FROM FISH TO TETRAPODS

Elpistostege watsoni

In the late 1930s, our understanding of the transition of fish to tetrapods — and the eventual jump to modern vertebrates — took an unexpected leap forward. The evolutionary a'ha came from a single partial fossil skull found on the shores of a riverbank in Eastern Canada. 

Meet the Stegocephalian, Elpistostege watsoni, an extinct genus of finned tetrapodomorphs that lived during the Late Givetian to Early Frasnian of the Late Devonian — 382 million years ago. 

Elpistostege watsoni — perhaps the sister taxon of all other tetrapods — was first described in 1938 by British palaeontologist and elected Fellow of the Royal Society of London, Thomas Stanley Westoll. Westoll's research interests were wide-ranging. He was a vertebrate palaeontologist and geologist best known for his innovative work on Palaeozoic fishes and their relationships with tetrapods. 

As a specialist in early fish, Westoll was asked to interpret that single partial skull roof discovered at the Escuminac Formation in Quebec, Canada. His findings and subsequent publication named Elpistostege watsoni and helped us to better understand the evolution of fishes to tetrapods — four-limbed vertebrates — one of the most important transformations in vertebrate evolution. 

Hypotheses of tetrapod origins rely heavily on the anatomy of but a few tetrapod-like fish fossils from the Middle and Late Devonian, 393–359 million years ago. These taxa — known as elpistostegalians — include Panderichthys, Elpistostege and Tiktaalik — none of which had yet to reveal the complete skeletal anatomy of the pectoral fin. 

Elpistostege watsoni

None until 2010 that is, when a complete 1.57-metre-long articulated specimen was found and described by Richard Cloutier et al. in 2020. 

The specimen helped us to understand the origin of the vertebrate hand. Stripped from its encasing stone, it revealed a set of paired fins of Elpistostege containing bones homologous to the phalanges (finger bones) of modern tetrapods and is the most basal tetrapodomorph known to possess them. 

Once the phalanges were uncovered, prep work began on the fins. The fins were covered in wee scales and lepidotrichia (fin rays). The work was tiresome, taking more than 2,700 hours of preparation but the results were thrilling. 

Origin of the Vertebrate Hand

We could now clearly see that the skeleton of the pectoral fin has four proximodistal rows of radials — two of which include branched carpals — as well as two distal rows organized as digits and putative digits. 

Despite this skeletal pattern — which represents the most tetrapod-like arrangement of bones found in a pectoral fin to date blurring the line between fish and land vertebrates — the fin retained lepidotrichia (those wee fin rays) distal to the radials. 

This arrangement confirmed an age-old question — showing us for the first time that the origin of phalanges preceded the loss of fin rays, not the other way around.

E. watsoni is very closely related to Tiktaalik roseae found in 2004 in the Canadian Arctic — a tetrapodomorpha species also known as a Choanata. These were advanced forms transitional between fish and the early labyrinthodonts playfully referred to as fishapods — half-fish, half-tetrapod in appearance and limb morphology. 

Up to that point, the relationship of limbed vertebrates (tetrapods) to lobe-finned fish (sarcopterygians) was well known, but the origin of major tetrapod features remained obscure for lack of fossils that document the sequence of evolutionary changes — until Tiktaalik. While Tiktaalik is technically a fish, this fellow is as far from fish-like you can be and still be a card-carrying member of the group. 

Tiktaalik roseae

Complete with scales and gills, this proto-fish lacked the conical head we see in modern fish but had a rather flattened triangular head more like that of a crocodile. 

Tiktaalik had scales on its back and fins with fin webbing but like early land-living animals, it had a distinctive flat head and neck. He was a brawny brute. The shape of his skull and shoulder look part fish and part amphibian.

The watershed moment came as Tiktaalik was prepped. Inside Tiktaalik's fins, we find bones that correspond to the upper arm, forearm and even parts of the wrist — all inside a fin with webbing — remarkable! 

Its fins have thin ray bones for paddling like most fish, but with brawny interior bones that gave Tiktaalik the ability to prop itself up, using his limbs for support. I picture him propped up on one paddle saying, "how you doing?" 

Six years after Tiktaalik was discovered by Neil Shubin and team in the ice-covered tundra of the Canadian Arctic on southern Ellesmere Island, a team working the outcrops at Miguasha on the Gaspé Peninsula discovered the only fully specimen of E. watsoni found to date — greatly increasing our knowledge of this finned tantalizingly transitional tetrapodomorph. 

E. watsoni fossils are rare — this was the fourth specimen collected in over 130 years of hunting. Charmingly, the specimen was right on our doorstop — extracted but a few feet away from the main stairs descending onto the beach of Miguasha National Park. 

L'nu Mi’gmaq First Nations of the Gespe’gewa’gi Region

Miguasha is nestled in the Gaspésie or Gespe’gewa’gi region of Canada — home to the Mi’gmaq First Nations who self-refer as L’nu or Lnu. The word Mi’gmaq or Mi’kmaq means the family or my allies/friends in Mi'kmaw, their native tongue (and soon to be Nova Scotia's provincial first language). They are the people of the sea and the original inhabitants of Atlantic Canada having lived here for more than 10,000 years. 

The L'nu were the first First Nation people to establish contact and trade with European explorers in the 16th and 17th centuries — and perhaps the Norse as early as the turn of the Millenium. Sailing vessels filled with French, British, Scottish, Irish and others arrived one by one to lay claim to the region — settling and fighting over the land. As each group rolled out their machinations of discovery, tensions turned to an all-out war with the British and French going head to head. I'll spare you the sordid details but for everyone caught in the crossfire, it went poorly.

North America Map 1775 (Click to Enlarge)

Cut to 1760, the British tipped the balance with their win at the Battle of the Restigouche, the last naval battle between France and England for possession of the North American continent — Turtle Island. 

The bittersweet British victory sparked the American War of Independence. 

For the next twenty years, the L'nu would witness and become embroiled in yet another war for these lands, their lands — first as bystanders, then as American allies, then intimidated into submission by the British Royal Navy with a show of force by way of a thirty-four gun man-of-war, encouraging L'nu compliance — finally culminating in an end to the hostilities with the 1783 Treaty of Paris. 

The peace accord held no provisions for the L'nu, Métis and First Nations impacted. None of these newcomers was Mi'kmaq — neither friends nor allies.

It was to this area some sixty years later that the newly formed Geological Survey of Canada (GSC) began exploring and mapping the newly formed United Province of Canada. Geologists in the New Brunswick Geology Branch traipsed through the rugged countryside that would become a Canadian province in 1867. 

It was on one of these expeditions that the Miguasha fossil outcrops were discovered. They, too, would transform in time to become Miguasha National Park or Parc de Miguasha, but at first, they were simply the promising sedimentary exposures on the hillside across the water —  a treasure trove of  Late Devonian fauna waiting to be discovered.

In the summer of 1842, Abraham Gesner, New Brunswick’s first Provincial Geologist, crossed the northern part of the region exploring for coal. Well, mostly looking for coal. Gesner also had a keen eye for fossils and his trip to the Gaspé Peninsula came fast on the heels of a jaunt along the rocky beaches of Chignecto Bay at the head of the Bay of Fundy and home to the standing fossil trees of the Joggins Fossil Cliffs. 

Passionate about geology and chemistry, he is perhaps most famous for his invention of the process to distil the combustible hydrocarbon kerosene from coal oil — a subject on which his long walks exploring a budding Canada gave him a great deal of time to consider. We have Gesner to thank for the modern petroleum industry. He filed many patents for clever ways to distil the soft tar-like coal or bitumen still in use today.

He was skilled in a broad range of scientific disciplines — being a geologist, palaeontologist, physician, chemist, anatomist and naturalist — a brass tacks geek to his core. Gesner explored the coal exposures and fossil outcrops across the famed area that witnessed the region become part of England and not France — and no longer L'nu.

Following the Restigouche River in New Brunswick through the Dalhousie region, Gesner navigated through the estuary to reach the southern coast of the Gaspé Peninsula into what would become the southeastern coast of Quebec to get a better look at the cliffs across the water. He was the first geologist to lay eyes on the Escuminac Formation and its fossils.

In his 1843 report to the Geologic Survey, he wrote, “...I found the shore lined with a coarse conglomerate. Farther eastward the rocks are light blue sandstones and shales, containing the remains of vegetables. (...) In these sandstone and shales, I found the remains of fish and a small species of tortoise with fossil foot-marks.”

We now know that this little tortoise was the famous Bothriolepis, an antiarch placoderm fish. It was also the first formal mention of the Miguasha fauna in our scientific literature. Despite the circulation of his report, Gesner’s discovery was all but ignored — the cliffs and their fossil bounty abandoned for decades to come. Geologists like Ells, Foord and Weston, and the research of Whiteaves and Dawson, would eventually follow in Gesner's footsteps.

North America Map 1866 (Click to Enlarge)

Over the past 180 years, this Devonian site has yielded a wonderfully diverse aquatic assemblage from the Age of Fishes — five of the six fossil fish groups associated with the Devonian including exceptionally well-preserved fossil specimens of the lobe-finned fishes. 

This is exciting as it is the lobe-finned fishes — the sarcopterygians — that gave rise to the first four-legged, air-breathing terrestrial vertebrates – the tetrapods. 

Fossil specimens from Miguasha include twenty species of lower vertebrates — anaspids, osteostra-cans, placoderms, acanthodians, actinopterygians and sarcopterygians — plus a limited invertebrate assemblage, along with terrestrial plants, scorpions and millipedes.

Originally interpreted as a freshwater lacustrine environment, recent paleontological, taphonomic, sedimentological and geochemical evidence corroborates a brackish estuarine setting — and definitely not the deep waters of the sea. This is important because the species that gave rise to our land-living animals began life in shallow streams and lakes. It tells us a bit about how our dear Elpistostege watsoni liked to live — preferring to lollygag in cool river waters where seawater mixed with fresh. Not fully freshwater, but a wee bit of salinity to add flavour.  

• Photos: Elpistostege watsoni (Westoll, 1938 ), Upper Devonian (Frasnian), Escuminac formation, Parc de Miguasha, Baie des Chaleurs, Gaspé, Québec, Canada. John Fam, VanPS
• Origin of the Vertebrate Hand Illustration, https://www.nature.com/articles/s41586-020-2100-8
• Tiktaalik Illustration: By Obsidian Soul - Own work, CC BY 4.0, https://commons.wikimedia.org/w/index.php?curid=47401797

References & further reading:

• From Water to Land: https://www.miguasha.ca/mig-en/the_first_discoveries.php
• UNESCO Miguasha National Park: https://whc.unesco.org/en/list/686/
• Office of L'nu Affairs: https://novascotia.ca/abor/aboriginal-people/
• Cloutier, R., Clement, A.M., Lee, M.S.Y. et al. Elpistostege and the origin of the vertebrate hand. Nature 579, 549–554 (2020). https://doi.org/10.1038/s41586-020-2100-8
• Daeschler, E. B., Shubin, N. H. & Jenkins, F. A. Jr. A Devonian tetrapod-like fish and the evolution of the tetrapod body plan. Nature 440, 757–763 (2006).
• Shubin, Neil. Your Inner Fish: A Journey into the 3.5 Billion History of the Human Body.
• Evidence for European presence in the Americas in AD 1021: https://www.nature.com/articles/s41586-021-03972-8

AMMONITE FROM FOLKSTONE

Sheer beauty — a beautiful Euhoplites ammonite from Folkstone, UK. I've been really enjoying looking at all oil-in-water colouring and chunkiness of these ammonites.

Euhoplites is an extinct ammonoid cephalopod from the Lower Cretaceous, characterized by strongly ribbed, more or less evolute, compressed to inflated shells with flat or concave ribs, typically with a deep narrow groove running down the middle.

In some, ribs seem to zigzag between umbilical tubercles and parallel ventrolateral clavi. In others, the ribs are flexious and curve forward from the umbilical shoulder and lap onto either side of the venter.

Its shell is covered in the lovely lumps and bumps we associate with the genus. The function of these adornments are unknown. I wonder if they gave them greater strength to go deeper into the ocean to hunt for food. 

They look to have been a source of hydrodynamic drag, likely preventing Euhoplites from swimming at speed. Studying them may give some insight into the lifestyle of this ancient marine predator. Euhoplites had shells ranging in size up to a 5-6cm. 

We find them in Lower Cretaceous, middle to upper Albian age strata. Euhoplites has been found in Middle and Upper Albian beds in France where it is associated respectively with Hoplites and Anahoplites, and Pleurohoplites, Puzosia, and Desmoceras; in the Middle Albian of Brazil with Anahoplites and Turrilites; and in the Cenomanian of Texas.

This species is the most common ammonite from the Folkstone Fossil Beds in southeastern England where a variety of species are found, including this 37mm beauty from the collections of José Juárez Ruiz.

UNESCOCERATOPS BY JULIUS CSOTONYI

Unescoceratops koppelhusae, Julius Csotonyi

A very sweet small leptoceratopsid dinosaur, Unescoceratops koppelhusae — a new species in the collections of the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta.

The colourful and beautifully detailed painting you see here is by the very talented Julius Csotonyi who captured the magnificence of form, texture and palette to bring this small leptoceratopsid dinosaur to life.

The Royal Tyrrell Museum of Palaeontology, named in honour of Joseph Burr Tyrrell, is a palaeontology museum and research facility in Drumheller, Alberta, Canada. 

This jaw is the holotype specimen of this small leptoceratopsid dinosaur. Only a handful of isolated fossils have been found from this species, including a jaw that is the holotype specimen now in collections at the Royal Tyrell. 

The Royal Tyrrell Museum of Palaeontology, named in honour of Joseph Burr Tyrrell, is a palaeontology museum and research facility in Drumheller, Alberta, Canada. 

Unescoceratops koppelhusae, RTMP Collections

The rusty chocolate jaw bone you see here is the puzzle piece that helped all of the research come together and help us to better understand more about the diminutive leptoceratopsid dinosaurs from Alberta. 

The Cleveland Museum of Natural History's Michael Ryan and David Evans of the Royal Ontario Museum in Toronto recently determined that the specimen was a new genus and species. 

Unescoceratops is a genus of leptoceratopsid ceratopsian dinosaurs known from the Late Cretaceous (about 76.5-75 million years ago) of Alberta, Canada. Unescoceratops is thought to have been between one and two meters long and less than 91 kilograms. A plant-eater, its teeth were the roundest of all Leptocertopsids.

Dinosaur Provincial Park, Alberta, Canada

The genus name acknowledges the UNESCO  World Heritage Site, Dinosaur Provincial Park, where the fossil was found. 

In addition to its particularly beautiful scenery, Dinosaur Provincial Park – located at the heart of the province of Alberta's badlands – is unmatched in terms of the number and variety of high-quality specimens.

To date, they represent more than 44 species, 34 genera and 10 families of dinosaurs, dating back 75-77 million years. This provides us with remarkable insight into life millions of years ago.

The park contains exceptional riparian habitat features as well as badlands of outstanding aesthetic value.

The creamy honey, beige and rust coloured hills around the fossil locality are outstanding examples of major geological processes and fluvial erosion patterns in semi-arid steppes — think glorious! 

The scenic badlands stretch along 26 kilometres of high quality and virtually undisturbed riparian habitat, presenting a landscape of stark but exceptional natural beauty.

The species name honours Dr. Eva Koppelhus, who has made significant contributions to vertebrate palaeontology and palynology. 

The genus is named to honour the UNESCO World Heritage Site designation for the locality where the specimen was found and from the Greek “ceratops,” which means 'horned face'. 

Dr Michael Ryan explained that he meant to honour UNESCO's efforts to increase understanding of natural history sites around the world.

© Julius T. Csotonyi An illustration of Unescoceratops koppelhusae, a plant-eating dinosaur from the Late Cretaceous period that lived approximately 75 million years ago shared with his gracious permission. 

ABOUT THE ARTIST

Dr. Julius Csotonyi is a Vancouver-based scientific illustrator and natural history fine artist. He is a featured paleoartist on Season One and Season Two of BC's Fossil Bounty. Julius has a scientific background in ecology (MSc) and microbiology (PhD) which has taken him to study sensitive ecosystems, from sand dunes in the Rocky Mountain parks to hydrothermal vents at the bottom of the Pacific Ocean. 

These experiences have fuelled his strong resolve to work toward preserving our Earth’s biota. Painting biological subjects is one means that he uses to both enhance public awareness of biological diversity and to motivate concern for its welfare.   

He paints murals and panels that have appeared in numerous museums including the Smithsonian’s National Museum of Natural History, press release images for scientific publications, books, stamp sets — including the outstanding 2018 “Sharks of Canada” set for Canada Post — and coins for the Royal Canadian Mint. To view more of Julius Csotonyi's exquisite work visit: https://csotonyi.com/

NOOTKA FOSSILS AND FIRST NATIONS HISTORY

Nootka Fossil Field Trip. Photo: John Fam

The rugged west coast of Vancouver Island offers spectacular views of a wild British Columbia. Here the seas heave along the shores slowly eroding the magnificent deposits that often contain fossils. 

Just off the shores of Vancouver Island, east of Gold River and south of Tahsis is the picturesque and remote Nootka Island.

This is the land of the proud and thriving Nuu-chah-nulth First Nations who have lived here always. 

Always is a long time, but we know from oral history and archaeological evidence that the Mowachaht and Muchalaht peoples lived here, along with many others, for many thousands of years — a time span much like always. 

While we know this area as Nootka Sound and the land we explore for fossils as Nootka Island, these names stem from a wee misunderstanding. 

Just four years after the 1774 visit by Spanish explorer Juan Pérez — and only a year before the Spanish established a military and fur trading post on the site of Yuquot — the Nuu-chah-nulth met the Englishman, James Cook.  

Captain Cook sailed to the village of Yuquot just west of Vancouver Island to a very warm welcome. He and his crew stayed on for a month of storytelling, trading and ship repairs. Friendly, but not familiar with the local language, he misunderstood the name for both the people and land to be Nootka. In actual fact, Nootka means, go around, go around. 

Two hundred years later, in 1978, the Nuu-chah-nulth chose the collective term Nuu-chah-nulth — nuučaan̓uł, meaning all along the mountains and sea or along the outside (of Vancouver Island) — to describe themselves. 

It is a term now used to describe several First Nations people living along western Vancouver Island, British Columbia. 

It is similar in a way to the use of the United Kingdom to refer to the lands of England, Scotland and Wales — though using United Kingdom-ers would be odd. Bless the Nuu-chah-nulth for their grace in choosing this collective name.  

An older term for this group of peoples was Aht, which means people in their language and is a component in all the names of their subgroups, and of some locations — Yuquot, Mowachaht, Kyuquot, Opitsaht. While collectively, they are the Nuu-chah-nulth, be interested in their more regional name should you meet them. 

But why does it matter? If you have ever mistakenly referred to someone from New Zealand as an Aussie or someone from Scotland as English, you have likely been schooled by an immediate — sometimes forceful, sometimes gracious — correction of your ways. The best answer to why it matters is because it matters.

Each of the subgroups of the Nuu-chah-nulth viewed their lands and seasonal migration within them (though not outside of them) from a viewpoint of inside and outside. Kla'a or outside is the term for their coastal environment and hilstis for their inside or inland environment.

It is to their kla'a that I was most keen to explore. Here, the lovely Late Eocene and Early Miocene exposures offer up fossil crab, mostly the species Raninid, along with fossil gastropods, bivalves, pine cones and spectacularly — a singular seed pod. These wonderfully preserved specimens are found in concretion along the foreshore where time and tide erode them out each year.

Five years after Spanish explorer Juan Pérez's first visit, the Spanish built and maintained a military post at Yuquot where they tore down the local houses to build their own structures and set up what would become a significant fur trade port for the Northwest Coast — with the local Chief Maquinna's blessing and his warriors acting as middlemen to other First Nations. 

Following reports of Cook's exploration British traders began to use the harbour of Nootka (Friendly Cove) as a base for a promising trade with China in sea-otter pelts but became embroiled with the Spanish who claimed (albeit erroneously) sovereignty over the Pacific Ocean. 

Dan Bowen searching an outcrop. Photo: John Fam

The ensuing Nootka Incident of 1790 nearly led to war between Britain and Spain (over lands neither could actually claim) but talk of war settled and the dispute was settled diplomatically. 

George Vancouver on his subsequent exploration in 1792 circumnavigated the island and charted much of the coastline. His meeting with the Spanish captain Bodega y Quadra at Nootka was friendly but did not accomplish the expected formal ceding of land by the Spanish to the British. 

It resulted however in his vain naming the island "Vancouver and Quadra." The Spanish captain's name was later dropped and given to the island on the east side of Discovery Strait. Again, another vain and unearned title that persists to this day.

Early settlement of the island was carried out mainly under the sponsorship of the Hudson's Bay Company whose lease from the Crown amounted to 7 shillings per year — that's roughly equal to £100.00 or $174 CDN today. Victoria, the capital of British Columbia, was founded in 1843 as Fort Victoria on the southern end of Vancouver Island by the Hudson's Bay Company's Chief Factor, Sir James Douglas. 

With Douglas's help, the Hudson's Bay Company established Fort Rupert on the north end of Vancouver Island in 1849. Both became centres of fur trade and trade between First Nations and solidified the Hudson's Bay Company's trading monopoly in the Pacific Northwest.

The settlement of Fort Victoria on the southern tip of Vancouver Island — handily south of the 49th parallel — greatly aided British negotiators to retain all of the islands when a line was finally set to mark the northern boundary of the United States with the signing of the Oregon Boundary Treaty of 1846. Vancouver Island became a separate British colony in 1858. British Columbia, exclusive of the island, was made a colony in 1858 and in 1866 the two colonies were joined into one — becoming a province of Canada in 1871 with Victoria as the capital.

Dan Bowen, Chair of the Vancouver Island Palaeontological Society (VIPS) did a truly splendid talk on the Fossils of Nootka Sound. With his permission, I have uploaded the talk to the ARCHEA YouTube Channel for all to enjoy. Do take a boo, he is a great presenter. Dan also graciously provided the photos you see here. The last of the photos you see here is from the August 2021 Nootka Fossil Field Trip. Photo: John Fam, Vice-Chair, Vancouver Paleontological Society (VanPS).

Know Before You Go — Nootka Trail

The Nootka Trail passes through the traditional lands of the Mowachaht/Muchalat First Nations who have lived here since always. They share this area with humpback and Gray whales, orcas, seals, sea lions, black bears, wolves, cougars, eagles, ravens, sea birds, river otters, insects and the many colourful intertidal creatures that you'll want to photograph.

This is a remote West Coast wilderness experience. Getting to Nootka Island requires some planning as you'll need to take a seaplane or water taxi to reach the trailhead. The trail takes 4-8 days to cover the 37 km year-round hike. The peak season is July to September. Permits are not required for the hike. 

Access via: Air Nootka floatplane, water taxi, or MV Uchuck III

• Dan Bowen, VIPS on the Fossils of Nootka: https://youtu.be/rsewBFztxSY
• https://www.thecanadianencyclopedia.ca/en/article/sir-james-douglas
• file:///C:/Users/tosca/Downloads/186162-Article%20Text-199217-1-10-20151106.pdf
• Nootka Trip Planning: https://mbguiding.ca/nootka-trail-nootka-island/#overview

FOSSILS OF CANADA'S EASTERN SHORES

Hylonomus lyelli, Ancestor of all dinosaurs

The fossil cliffs at Joggins are one of Canada's gems, now a UNESCO World Heritage Site, you can visit to see our ancient world frozen in time. 

Preserved in situ is a snapshot of an entire food chain of a terrestrial Pennsylvanian Coal Age wetland.

The outcrop holds fossil plant life — including impressive standing lycopsid trees that formed the framework of these wetlands — decomposing detritivores in the invertebrates and tetrapods, the predatory carnivores of the day.

The Coal Age trees were fossilized where they stood 300-million-years ago with the remains of the earliest reptiles entombed within. The preservation is quite marvellous with the footprints of creatures who once lived in these wetlands are frozen where they once walked and the dens of amphibians are preserved with remnants of their last meal. 

Nowhere is a record of plant, invertebrate and vertebrate life within now fossilized forests rendered more evocatively. The fossil record at Joggins contains 195+ species of plants, invertebrates and vertebrates. The fossil plant life became the vast coal deposits for which this period of Earth's history is named. 

Recorded in the rock are vertebrate and invertebrate fauna both aquatic and terrestrial. This broad mix of specimens gives us a view into life back in the Pennsylvanian and sets us up to understand their ecological context.

Pennsylvanian Coal Age Ecosystem, 300-Million-Years-Old

The fossil record includes species first defined at Joggins, some of which are found nowhere else on Earth. 

It was here that Sir Charles Lyell, with Sir William Dawson, founder of modern geology, discovered tetrapods — amphibians and reptiles — entombed in the upright fossil trees. 

Later work by Dawson would reveal the first true reptile, Hylonomus lyelli, ancestor of all dinosaurs that would rule the Earth 100 million years later. 

This tiny reptile serves as the reference point where animals finally broke free of the water to live on land. This evolutionary milestone recorded at Joggins remains pivotal to understanding the origins of all vertebrate life on land, including our own species. 

Sir Charles Lyell, author of Principles of Geology, first noted the exceptional natural heritage value of the Joggins Fossil Cliffs, calling them “...the finest example in the world of a natural exposure in a continuous section ten miles long, occurs in the sea cliffs bordering a branch of the Bay of Fundy in Nova Scotia.” Indeed, the world-famous Bay of Fundy with its impressive tides, the highest in the world, and stormy nature exposed much of this outcrop. 

Geological accounts of the celebrated coastal section at Joggins first appear in the published literature in 1828–1829, by Americans C.T. Jackson and F. Alger, and by R. Brown and R. Smith, managers for the General Mining Association in the Sydney and Pictou coal fields. Brown and Smith’s account is the first to document the standing fossil trees.

Joggins Fossil Cliffs Map (Click to Enlarge)

Plan Your Joggins Fossil Cliffs Staycation

Joggins Fossil Cliffs is a Canadian gem — and they welcome visitors. They offer hands-on learning and discovery microscope activities in their Fossil Lab.

You can explore interpretive displays in the Joggins Fossil Centre before heading out to the beach and cliffs with an interpreter.

Their guided tours of the fossil site include an educational component that tells you about the geology, ecology, palaeontology and conservation of this very special site. 

Joggins / Chegoggin / Mi'kmaq L'nu

We know this area as Joggins today. In Mi'kmaw, the language spoken in Mi'kma'ki, the territory of the Mi'kmaq L'nu, the area bears another name, Chegoggin, place of fishing weirs.

Booking Your Class Field Trip

If you are a teacher and would like to book a class field trip, contact the Director of Operations via the contact information listed below. They will walk you through Covid safety and discuss how to make your visit educational, memorable and fun.

Know Before You Go

The Bay of Fundy has the highest tides in the world. Beach walks are scheduled according to the tides and run regardless of the weather. Good low tides but raining, the beach walk goes on. Lovely and sunny but with a high tide, the beach walk must wait. So, you will want to dress for it as they will not be cancelled in the event of rain. Should severe weather be a factor, bookings may need to be rescheduled at the discretion of the Joggins staff.

Any questions about booking your school field trip? Feel free to email:  operations@jogginsfossilcliffs.net or call: 1 (902) 251-2727 EXT 222.

References & further reading:

Joggins Fossil Cliffs: https://jogginsfossilcliffs.net/cliffs/history/

Image: Hylonomus lyelli, Una ricostruzione di ilonomo by Matteo De Stefano/MUSEThis file was uploaded by MUSE - Science Museum of Trento in cooperation with Wikimedia Italia., CC BY-SA 3.0, https://commons.wikimedia.org/w/index.php?curid=48143186

Image: Arthropleura: Par Tim Bertelink — Travail personnel, CC BY-SA 4.0, https://commons.wikimedia.org/w/index.php?curid=48915156

Joggins Map: Joggins Fossil Cliffs: https://jogginsfossilcliffs.net/cliffs/history/

FOSSIL AMPHIBIANS OF NOVA SCOTIA

Dendrerpeton acadianum, an extinct amphibian

One of the best Canadian fossil finds stems from a random boulder picked up on the beach near the town of Joggins, Nova Scotia. Inside were the bones of a fully articulated skeleton of Dendrerpeton acadianum, a Temnospondyli from the Lower Pennsylvanian. 

These little cuties belong to an extinct genus of amphibians who loved wet, swampy wetlands similar to those we find in the bayous of Mississippi today.   

Dendrerpeton is the primitive sister-group to a clade of Temnospondyls that includes Trimerorhachoids, the Eryopoids — Ervops, Parioxys, & Sclerocephalus — Zatracheids & Dissorophoids. 

This little guy along with finding the first true reptile, Hylonomus lyelli, ancestor of all dinosaurs that would rule the Earth 100 million years later serve as the reference point where animals finally broke free of the water to live on land. This evolutionary milestone recorded at Joggins remains pivotal to understanding the origins of all vertebrate life on land, including our own species. 

Joggins records life in a once a wet, swampy wetland

Sir Charles Lyell, the author of Principles of Geology, first noted the exceptional natural heritage value of the Joggins Fossil Cliffs. He described them as: 

“...the finest example in the world of a natural exposure in a continuous section ten miles long, occurs in the sea cliffs bordering a branch of the Bay of Fundy in Nova Scotia.” 

Indeed, the world-famous Bay of Fundy with its impressive tides, the highest in the world, and stormy nature exposed much of this outcrop. 

FOSSIL FROM THE GAULT

The chunky ammonite Proeuhoplites subtuberculatus, bed II (iv), Folkstone Gault Clay, county of Kent, southeast England.

This matrix you see here is the Gault Clay, known locally as the Blue Slipper. This fine muddy clay was deposited 105-110 million years ago during the Lower Cretaceous (Upper and Middle Albian) in a calm, fairly deep-water continental shelf that covered what is now southern England and northern France.

Lack of brackish or freshwater fossils indicates that the gault was laid down in open marine environments away from estuaries. The maximum depth of the Gault is estimated 40-60m a figure which has been reached by the presence of Borings made by specialist Algal-grazing gastropods and supported by a study made by Khan in 1950 using Foraminifera. Estimates of the surface water temperatures in the Gault are between 20-22°c and 17-19°c on the seafloor. These estimates have been reached by bulk analysis of sediments which probably register the sea surface temperature for calcareous nanofossils.

It is responsible for many of the major landslides around Ventnor and Blackgang the Gault is famous for its diverse fossils, mainly from mainland sites such as Folkestone in Kent.

Folkestone, Kent is the type locality for the Gault clay yielding an abundance of ammonites, the same cannot be said for the Isle of Wight Gault, however, the south-east coast of the island has proved to be fossiliferous in a variety of ammonites, in particular, the Genus Hoplites, Paranahoplites and Beudanticeras.

While the Gault is less fossiliferous here on the island it can still produce lovely marine fossils, mainly ammonites and fish remains from these muddy mid-Cretaceous seas. The Gault clay marine fossils include the ammonites (such as Hoplites, Hamites, Euhoplites, Anahoplites, and Dimorphoplites), belemnites (such as Neohibolites), bivalves (notably Birostrina and Pectinucula), gastropods (including the lovely Anchura), solitary corals, fish remains (including shark teeth), scattered crinoid remains, and crustaceans (look for the crab Notopocorystes).

Occasional fragments of fossil wood may also be found. The lovely ammonite you see here is from the Gault Clays of Folkstone. Not all who name her would split the genus Euhoplites. There’s a reasonable argument for viewing this beauty as a very thick form of E. loricatus with Proeuhoplites being a synonym of Euhoplites. Collected, photographed and prepped by Thomas Miller. Approx 35mm across.

Jack Wonfor shared a wealth of information on the Gault and has many lovely examples of the ammonites found here in his collections. If you wish to know more about the Gault clay a publication by the Palaeontological Association called 'Fossils of the Gault clay' by Andrew S. Gale is available in Dinosaur Isle's gift shop.

There is a very good website maintained by Fred Clouter you can look at for reference. It also contains many handy links to some of the best fossil books on the Gault Clay and Folkstone Fossil Beds. Check it out here: http://www.gaultammonite.co.uk/