CTENOPHORES: COMB JELLIES

Cannibalistic Comb Jellies

This festive lantern looking lovely belongs to a group of invertebrates known as comb jellies.

Comb jellies are named for their unique plates of giant fused cilia, or combs, which run in eight rows up and down the length of their bodies. 

They are armed with sticky cells or colloblasts, that do not sting but display wonderful bioluminescent colouring as they move through the sea.

Ctenophores or comb jellies are one of the phylogenetically most important and controversial metazoan groups. 

Looks can be deceiving. At first glance you might think you are looking at a jellyfish but this is not the case. Surprisingly, they are not jellyfish and are not closely related, though they do share some characteristics with the gelatinous members of the subphylum Medusozoa. 

Comb jellies are not picky eaters. Their tastes range to what is at hand, including cannibalizing other comb jellies. They will feast on their kin along with tasty plankton, zooplankton, crustaceans and wee fish.

Interest in their fossil record has been catalysed by spectacularly preserved soft-bodied specimens from Cambrian Lagerstätten of the 518-million-years-old Chengjiang Biota, the 505-million-years-old Burgess Shale and other Burgess Shale-like deposits. 

We find them in the Late Devonian Escuminac Formation at Miguasha National Park, Quebec, Canada — a UNESCO world heritage site famous for its abundance of well-preserved vertebrate fossils including most major evolutionary groups of Devonian lower vertebrates from jawless fish to stem-tetrapods.

Based on morphological similarities of this Canadian fossil with stem-ctenophore fossils from the Cambrian Lagerstätte of the Chinese locality Chengjiang, they have been assessed for their affinity to stem-group ctenophores (dinomischids, Siphusauctum, scleroctenophorans) and early crown-group ctenophores. Modern ctenophores and many fossil forms lack mineralized hard parts, which renders the rare fossils that have been extracted from several Lagerstätten quite remarkable. 

Like the soft bodies of jellyfish and the polyps of hydrozoans and anthozoans, the probability for such soft bodies (or body regions) to become fossilized is extremely low. In spite of this low preservation potential, remains of stem-ctenophores have become known from several Cambrian and younger conservation deposits, and with even older candidate ctenophores in the Ediacaran. 

While Cambrian Lagerstätten have yielded several genera, ctenophore remains are much rarer in the Devonian; in particular, two studies, describing material from the German Hunsrück Slate. 

Bioluminescent Comb Jellies

This Early Devonian material, however, appears to belong to crown ctenophores morphologically similar to living forms such as Pleurobrachia, unlike the stem Cambrian taxa and the new Devonian stem taxon described here.

The most basal stem ctenophores are the dinomischids: sessile benthic petaloid invertebrates, many of which are equipped with a stalk. This group first was described from the Middle Cambrian Burgess Shale. Based on the genus Dinomischus, these early stalked forms were commonly called ‘dinomischids’. 

Zhao et al. shared that dinomischids "form a grade in the lower part of the ctenophore stem group” and include taxa such as Xianguangia, Daihua, and Dinomischus that have hexaradiate-based symmetry (e.g., sixfold, 18-fold). 

Some later, skeletonised stem-ctenophores were termed ‘Scleroctenophora’; ‘scleroctenophorans’ have a shorter stalk, lack the ‘petals’ and have no bracts and might be monophyletic. 

To date, all known dinomischids and scleroctenophorans are Cambrian. Remarkably, analysis of the material described here suggests it is a very late-surviving member of this part of the ctenophore tree, occurring in strata over a hundred million years younger with no intervening known record, thus making it a Lazarus taxon with an extensive ghost lineage. 

Palaeozoic sediments yield a growing number of fossil invertebrates with radial symmetries, some being quite enigmatic with body plans differing radically from those of extant organisms.

The morphological similarities to Cambrian forms and the mix of characters regarding overall shape and symmetries render this discovery important. The aims of this study are to describe the only known specimen of this Devonian ctenophore, discuss its phylogenetic and systematic position, and the impact of fossil data for ctenophore affinities, and assess its palaeoecological role.

SMILODON NORTH OF THE 49TH PARALLEL

This fierce predator with the luxurious coat is Smilodon fatalis — a compact but robust killer that weighed in around 160 to 280 kg and was 1.5 - 2.2 metres long.

Smilodon is a genus of the extinct machairodont subfamily of the felids. It is one of the most famous prehistoric mammals and the best known saber-toothed cat. Although commonly known as the saber-toothed tiger, it was not closely related to the tiger or other modern cats.

Up until a few years ago, all the great fossil specimens of this apex predator were found south of us in the United States. That was until some interesting bones from Medicine Hat, Alberta got a second look.

A few years ago, a fossil specimen caught the eye of researcher Ashley Reynolds as she was rummaging through the collections at the Royal Ontario Museum in Toronto. 

Back in the 1960s,  University of Toronto palaeontologist C.S. Churcher and his team had collected and donated more than 1,200 specimens from their many field seasons scouring the bluffs of the South Saskatchewan River near Medicine Hat, Alberta.

Churcher is a delightful storyteller and a palaeontologist with a keen eye. I had the very great pleasure of listening to many of his talks out at the University of British Columbia and a few Vancouver Paleontological Society meetings in the mid-2000s. 

"Rufus" was a thoroughly charming storyteller and shared many of his adventures from the field. 

He moved out to the West Coast for his retirement, first to Gabriola Island then to Victoria, but his keen love of the science kept him giving talks to enthralled listeners keen to hear about his survey of the Dakhleh Oasis in the Western Desert of Egypt, geomorphology, stratigraphy, recent biology, Pleistocene and Holocene lithic cultures, insights learned from Neolithic Islamic pottery to Roman settlements.

The specimens he had collected had been roughly sorted but never examined in detail. Reynolds, who was researching the growth patterns and life histories of extinct cats saw a familiar-looking bone from an ancient cat's right front paw. That tiny paw bone had reached through time and was positively identified as Canada's first Smilodon.

These Apex Predators used their exceptionally long upper canine teeth to hunt large mammals. 

Isotopes preserved in the bones of S. fatalis in the La Brea Tar Pits in California tell us that they liked to dine on bison (Bison antiquus) and camels (Camelops) along with deer and tapirs. Smilodon is thought to have killed its prey by holding it still with its forelimbs and biting it. And that was quite the bite!

Their razor-sharp incisors were arranged in an arch. Once they bit down, the teeth would hold their prey still and stabilize it while the canine bite was delivered — and what a bite that was. They could open their mouths a full 120 degrees.

Smilodon died out at the same time that most North and South American megafauna disappeared, about 10,000 years ago. Its reliance on large animals has been proposed as the cause of its extinction, along with climate change and competition with other species. 

APEX HUNTER OF ITS TIME: ANKYLORHIZA

Back in the 1880s, from fragments of bone weathered by time and tide, a most curious creature emerged into scientific view — an ancient toothed dolphin later named Ankylorhiza tiedemani. 

Its name, drawn from the Greek ankylo — bound or fused — and rhiza — root — hints at one of its more unusual traits: teeth with mostly single, fused roots. 

A formidable grin, and not at all what we might expect from the dolphins we know today.

We often think of dolphins as gentle, clever denizens of the sea. 

But cast your mind back to the Oligocene, and a rather different picture takes shape. Here was a hunter — swift, powerful, and armed with a mouthful of sharp teeth. Ankylorhiza tiedemani stood as the largest member of the Odontoceti — the great lineage of toothed whales that includes dolphins, porpoises, sperm whales, beaked whales, river dolphins, pilot whales, and their kin — all hunters of prey larger than plankton, all bearing teeth instead of baleen.

More clues surfaced in the decades that followed. Fragments in the 1970s and 1990s, and then something far more revealing — a nearly complete skeleton, now resting at the Mace Brown Museum of Natural History. A beautifully preserved skull, ribcage, much of the vertebral column, and even a solitary flipper. 

Rare treasures, these, for creatures of the sea. 

Together, they whisper a clearer story: a 4.8-metre predator, tracing its lineage back some 35–36 million years, diverging from baleen whales yet evolving strikingly similar features through convergence.

This was no languid swimmer. Some 24 million years ago, Ankylorhiza coursed through ancient seas with speed and purpose. 

Its body tells the tale — a narrow tailstock, additional tail vertebrae, and a shortened humerus in its flippers. Like modern dolphins, it likely powered itself with strong, rhythmic thrusts of its flukes, adjusting its course with hydrofoil-like flippers. 

Beneath the skin, robust muscles anchored to a relatively rigid torso — a design honed for movement, for pursuit, for the hunt.

The fossil record, however, does not always give up its secrets easily. Eocene whale skeletons show us the early transition from land to sea — limbs shrinking, bodies streamlining. 

But Oligocene specimens are rare, and with them, much of the story of how whales mastered fluke-powered swimming has remained elusive. 

Did these early dolphins possess the same refinements for speed? For a long time, we could only speculate.

Then came the work of Robert Boessenecker and colleagues. Their study of this remarkable skeleton reveals an animal poised between worlds — its forelimb structure bridging stem cetaceans and modern whales, its spine showing the beginnings of rigidity at the tail while retaining flexibility through the lower back. 

A body in transition, yet already capable.

And what a role it played. Its skull, teeth, vertebrae, and size all point to a macrophagous predator — one that hunted large prey and moved with relative speed. 

In life, Ankylorhiza may well have filled a niche much like that of today’s killer whales — an apex hunter of its time, commanding the ancient seas with quiet authority.

A fossil, yes — but also a story. One of innovation, convergence, and the relentless shaping of life in motion.

HOLLARDOPS: LE MAÎTRE

Hollardops sp. Devonian Trilobite

Hollardops is a genus of trilobite in the order Phacopida that lived during the Eifelian of the Middle Devonian. It was described by Le Maître in 1952 under type species Metacanthina mesocristata. 

The genus underwent reclassification in 1997 and emerged as Hollardops. We find this extinct arthropod in present-day Morocco. They share similarities with Greenops of New York and Canada but are generally larger than most Greenops species.

Hollardops have schizochroal eyes and a glabella that is slightly raised on the surface of the cephalon. Genal spines extend from the cephalon and extend to approximately the 6th thoracic segment.

Hollardops has eleven thoracic segments and also has five pairs of spines extending from the segments of the pygidium. Length ranges from approximately 3 to 9 cm.

Palaeo Coordinates — If you are a keen bean to head out in search of this lovely yourself, head to the Tazoulait Formation at Jbel (Jebel) Oufatène 30.8374368°N 4.9018067°W and Issimour 30.9669834°N 5.0373266°W SE of Alnif, western of Oued Alnif, Ma'ider region, Morocco.

CRETACEOUS PREDATORS: KOURISODON PUNTLEDGENSIS

Kourisodon puntledgensis

You would be a brave soul to be swimming in the warm, shallow seas of the Late Cretaceous—and braver still to linger near the surface. 

These waters belonged to the mosasaurs—sleek, powerful marine reptiles that ruled the global oceans with quiet authority. 

Since their first discovery in 1766, their bones have surfaced from nearly every corner of the world—New Zealand to Antarctica, Africa to Europe, the Americas to Japan—whispers of a dynasty that once circled the globe.

And yet, some of their most intriguing stories are written close to home.

Along the banks of the Puntledge River on Vancouver Island, a remarkable assemblage has come to light. Here, tucked into ancient marine sediments, we find the remains of both elasmosaurids and mosasaurs—echoes of a coastal ecosystem long vanished beneath forest and freshwater flow. 

As Dan Bowen of the Vancouver Island Paleontological Society notes, this stretch of river has yielded not one, but multiple marine reptiles from a time when this land lay beneath a teeming inland sea.

The first mosasaur material recovered here—around ten vertebrae belonging to Platecarpus—was discovered by Tim O’Bear and carefully excavated by a dedicated team led by Dr. Rolf Ludvigsen. Later prepared by Bowen and Joe Morin, these bones offer a tantalising glimpse of fast-moving predators that once patrolled these waters.

But it is a second discovery that truly sharpens the tale.

Kourisodon photo by Roland Tanglao

In 1993, upstream from the elasmosaur site, Joe Zembiliwich uncovered something altogether special on a field excursion led by the legendary Mike Trask. 

What emerged was Kourisodon puntledgensis—a name drawn from the Greek kourís and odon, meaning “razor tooth,” and an apt one at that.

Small by mosasaur standards—roughly 3.75 to 4.5 metres in length—Kourisodon was nonetheless a nimble and capable predator. 

First described within the “Leiodontini” and now placed among the clidastines, it hunted with precision in Pacific waters rich with life. Alongside it swam long-necked elasmosaurids, turtles, and other mosasaurs, though notably absent were the polycotylids so common elsewhere.

What makes this animal especially intriguing is its story of place—and of connection.

The type specimen of Kourisodon puntledgensis comes from a single locality within the Santonian-aged Pender Formation along the Puntledge River—the very place that lends the species its name. And yet, across the vast Pacific, its relatives appear again in the Upper Cretaceous rocks of Japan’s Izumi Group. 

There, fragmentary remains—including those of juveniles—hint at at least one additional, as-yet-unnamed species, distinguished by features such as longer maxillary teeth (Tanimoto, 2005; Caldwell & Konishi, 2007).

We see a similar trans-Pacific kinship echoed in the ammonites of these regions—shared lineages linking distant shores. But curiously, this connection does not extend inland. The marine reptiles of the Western Interior Seaway tell a different story entirely.

This is provinciality in action—ancient ecosystems shaped by geography, currents, and isolation. As detailed by Nicholls and Meckert (2002), the Pacific faunas of British Columbia evolved along their own path, distinct from their contemporaries to the east.

Today, a full-scale replica of Kourisodon puntledgensis—a sleek, 12-foot echo of those razor-toothed hunters—resides at the Canadian Fossil Discovery Centre in Morden, Manitoba. A long way from the Puntledge, perhaps, but still tethered to that riverbank story.

And the Puntledge continues to give.

It is also the home of a newly described elasmosaur—Traskasaura—named in honour of Mike, Pat, and Heather Trask. Discovered in 1988 and formally described in 2025, it adds yet another layer to this rich and ever-unfolding story of Vancouver Island’s ancient seas.

Stand along the Puntledge now, with the river slipping quietly past your boots, and you can almost feel it—the weight of deep time, the flicker of ancient oceans, the swift shadow of something moving just beneath the surface.

And, if you’ve spent enough time in the field, you’ll feel something else too.

The memory of those who walked these shores with you—who swung their last hammer, shared their last laugh, and left their stories folded gently into the stones we still turn over today.

References

Nicholls, E. L. and Meckert, D. (2002). Marine reptiles from the Nanaimo Group (Upper Cretaceous) of Vancouver Island. Canadian Journal of Earth Sciences 39(11):1591–1603.

Tanimoto, M. (2005). Mosasaur remains from the Upper Cretaceous Izumi Group of Southwest Japan. Netherlands Journal of Geosciences 84(3):373–378.

Caldwell, M., & Konishi, T. (2007). Taxonomic re-assignment of the first-known mosasaur specimen from Japan, and a discussion of circum-Pacific mosasaur paleobiogeography. Journal of Vertebrate Paleontology 27(2):517–520.

CBC News (2018). “Ferocious” new mosasaur skeleton coming to Morden.

CJOB (2018). “Ferocious, razor-like teeth”: new mosasaur comes to Morden's fossil centre.

Winnipeg News (2018). Morden museum's collection of mosasaur skeletons grows with new addition.

Image Two: By Roland Tanglao from Vancouver, Canada - Dinos at Courtenay Museum -20090628-7Uploaded by FunkMonk, CC BY 2.0, https://commons.wikimedia.org/w/index.php?curid=10364342

SAKARA MADAGASGAR: OXFORDIAN OUTCROPS

Lobolytoceras costellatum

This big beastie is a superb specimen of the ammonite Lobolytoceras costellatum showing the intricate fractal pattern of its septa. 

This lovely measures to a whopping 230 mm and hails from Oxfordian outcrops near Sakara, Madagascar. Lovingly prepped by the supremely talented José Juárez Ruiz.

Ammonites were predatory, squidlike creatures that lived inside coil-shaped shells. 

Like other cephalopods, ammonites had sharp, beak-like jaws inside a ring of squid-like tentacles that extended from their shells. They used these tentacles to snare prey — plankton, vegetation, fish and crustaceans — similar to the way a squid or octopus hunt today.

Catching a fish with your hands is no easy feat, as I'm sure you know. Ammonites did the equivalent, catching prey in their tentacles. They were skilled and successful hunters. They caught their prey while swimming and floating in the water column. 

Within their shells, they had a number of chambers, called septa, filled with gas or fluid that were interconnected by a wee air tube. By pushing air in or out, they were able to control their buoyancy in the water column.

They lived in the last chamber of their shells, continuously building new shell material as they grew. As each new chamber was added, the squid-like body of the ammonite would move down to occupy the final outside chamber.

They were a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids — octopuses, squid, and cuttlefish) then they are to shelled nautiloids such as the living Nautilus species.

Lobolytoceras costellatum

Ammonites have intricate and complex patterns on their shells called sutures. The suture patterns differ across species and tell us what time period the ammonite is from. 

If they are geometric with numerous undivided lobes and saddles and eight lobes around the conch, we refer to their pattern as goniatitic, a characteristic of Paleozoic ammonites.

Ammonites first appeared about 240 million years ago, though they descended from straight-shelled cephalopods called bacrites that date back to the Devonian, about 415 million years ago, and the last species vanished in the Cretaceous–Paleogene extinction event.

They were prolific breeders that evolved rapidly. If you could cast a fishing line into our ancient seas, it is likely that you would hook an ammonite, not a fish. 

They were prolific back in the day, living (and sometimes dying) in schools in oceans around the globe. We find ammonite fossils (and plenty of them) in sedimentary rock from all over the world.

In some cases, we find rock beds where we can see evidence of a new species that evolved, lived and died out in such a short time span that we can walk through time, following the course of evolution using ammonites as a window into the past.

For this reason, they make excellent index fossils. An index fossil is a species that allows us to link a particular rock formation, layered in time with a particular species or genus found there. 

Generally, deeper is older, so we use the sedimentary layers rock to match up to specific geologic time periods, rather the way we use tree-rings to date trees. A handy way to compare fossils and date strata across the globe.

BARNACLES: CUVIER TO DARWIN

Barnacles All Closed Up

One of the most interesting and enigmatic little critters we find at the seashore are barnacles. 

They cling to rocks at the waters' edge, closed to our curiosity, their domed mounds like little closed beaks shut to the water and the world.

They choose their permanent homes as larvae, sticking to hard substrates that will become their permanent homes for the rest of their lives. 

It has taken us a long time to find how they actually stick or what kind of "glue" they were using.

A clever fellow from Duke University's Marine Laboratory in Durhan, North Carolina finally cracked that puzzle. 

Instead of chopping up barnacles to see what makes them stick, he observed and collected the oozing glue from some Amphibalanus amphitrite as they secreted it.

Remarkably, the barnacle glue sticks to rocks in a similar way to how red cells bind together. Red blood cells bind and clot with a little help from some enzymes. These work to create long protein fibres that first blind, clot then form a scab. The mechanism barnacles use, right down to the enzyme, is very similar. That's especially interesting as about a billion years separate our evolutionary path from theirs.

So, with the help of their clever enzymes, they can affix to most anything – ship hulls, rocks, and even the skin of whales. If you find them in tidepools, you begin to see their true nature as they open up, their delicate feathery finger-like projections flowing back and forth in the surf.

Barnacle Cirri Seeking Tasty Plankton

Those wee feather-like bits you see are called cirri. Eight pairs of these thoracic limbs help barnacles to filter tasty bits of plankton from the surrounding water into their mouths.

Barnacles are cirripedes, a kind of crustacean that is covered with hard plates of calcium carbonate. Named for their cirri, they live stuck to hard surfaces in and around our world's oceans. While they do not look like crustaceans, they are definitely part of this taxonomic grouping that includes crab, lobster, crayfish, prawn, krill, and woodlice.

BARNACLES IN KWAK'WALA

In the Kwak̓wala language of the Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, barnacles are known as k̕wit̕a̱'a and broken barnacle shells are known as t̕sut̕su'ma.

BARNACLES IN THE FOSSIL RECORD

They have an old history. Their ancestors can be traced back to animals such as Priscansermarinus that lived during the Middle Cambrian – some 510 to 500 million years ago. 

I found my first barnacle fossil at a fossil site called Muir Creek on the south end of Vancouver Island. The fossil exposures at Muir are Oligocene, 20-25 million years old. This is about the time that barnacles can be found more readily as skeletal remains.

One of the reasons for the limited number of barnacle remains in the fossil record is their preferred habitat – high energy, shallow ocean environments. These tend to see a lot of tidal action that leads to erosion and barnacles being broken apart, slowly eroded down to bits too small to recognize for what they are.

One of the fossil remains we do find are not the barnacles themselves, but trace fossils of acrothoracican barnacle borings from Rogerella. These are commonly found in the fossil record beginning in the Devonian right up to today. Rogerella is a small pouch-shaped boring (a type of trace fossil) with a slit-like aperture currently produced by acrothoracican barnacles. 

These crustaceans extrude their legs upwards through the opening for filter-feeding (Seilacher, 1969; Lambers and Boekschoten, 1986). They are known in the fossil record as borings in carbonate substrates (shells and hardgrounds) from the Devonian to the Recent (Taylor and Wilson, 2003).

Barnacle Ancestry Goes Back to the Middle Cambrian

FROM MOLLUSCA TO ARTICULATA

Barnacles were originally classified by Linnaeus and Cuvier as Mollusca, but in 1830 John Vaughan Thompson published observations showing the metamorphosis of the nauplius and cypris larvae into adult barnacles. 

He noted how these larvae were similar to those of crustaceans.

In 1834 Hermann Burmeister published further information, reinterpreting these findings. The effect was to move barnacles from the phylum of Mollusca to Articulata, showing naturalists that detailed study was needed to reevaluate their taxonomy.

Charles Darwin took up this challenge in 1846 and developed his initial interest in a major study published as a series of monographs in 1851 and 1854. 

Darwin undertook this study, at the suggestion of his friend Joseph Dalton Hooker, to thoroughly understand at least one species before making the generalizations needed for his theory of evolution by natural selection.

BARNACLES IN A NUT SHELL

Barnacles are suspension feeders, sweeping small food into their mouth with their curved 'feet'. They are cemented to rock (usually), and covered with hard calcareous plates, which they shut firmly when the tide goes out. 

Barnacles reproduce sexually and produce little nauplius larvae that disperse in the plankton. Eventually, the larvae change into cypris form and attach on other hard surfaces to form new barnacles.

CERVUS CANADENSIS: MAGNIFICENT ELK

Nature awes me everyday. Quiet moments often shared solo or if lucky, with a good friend or one of the amazing animals that walk this Earth.

I was especially lucky to have many of them while staying in Banff, Alberta. 

A morning stroll became an epic moment shared with a herd of wild but nonplussed elk enjoying their breakfast.

There is something quietly magnificent about an elk moving through fresh snow — head lowered, breath curling into the cold air, long legs parting the white silence of a winter morning in Banff. It feels timeless. And in a way, it is.

The elk you see here, Cervus canadensis, belongs to a lineage that stretches deep into the Pleistocene — a time when ice sheets advanced and retreated across much of North America, reshaping landscapes and the lives within them. 

Elk are members of the family Cervidae, a group that first appears in the fossil record during the Early Miocene, roughly 20 million years ago. These early deer were small, forest-dwelling creatures, lacking the impressive antlers we associate with their modern kin.

By the Late Miocene and into the Pliocene, cervids began to diversify in both form and habitat. Antlers — those seasonal crowns of bone — became more elaborate, evolving as tools of display and combat. 

The genus Cervus, which includes modern elk, appears later, with fossils known from Eurasia before spreading into North America via the Bering Land Bridge during the Pleistocene, likely within the last 2 million years.

Once here, elk flourished.

Pleistocene deposits across North America — from tar seeps like Rancho La Brea in California to river gravels and cave assemblages further north — preserve their bones alongside an Ice Age cast of giants: mammoths, mastodons, dire wolves and short-faced bears. 

Elk held their own in this formidable company, adaptable grazers and browsers able to navigate shifting climates and changing ecosystems.

In Canada, elk fossils are known from a number of Quaternary sites, including Alberta and the Yukon, where their remains speak to a long history on these lands. 

As the glaciers withdrew at the end of the last Ice Age, elk expanded into newly opened habitats, tracking the spread of grasslands and open forests.

What you are seeing in Banff today is the continuation of that story — a survivor of ice and upheaval, still moving with quiet purpose through a landscape shaped by deep time.

I've been lucky enough to get to spend some time in Banff, looking for fossils, as an artist and exploring nature in all its glory.  It was heartwarming to see Elk most every day there and snow multiple times a week—and all this in April and May!

CAMBRIAN CROWN: THE SPINED ELEGANCE OF ORGMASPIS

This calcified beauty is Orygmaspis (Parabolinoides) spinula (Westrop, 1986), an Upper Cambrian trilobite recovered from the McKay Group near Tanglefoot Mountain in the Kootenay Rockies—one of those quietly extraordinary places where deep time peeks through in layered stone.

A member of the Order Asaphida, Orygmaspis carries the elegant geometry so characteristic of its kin: an inverted, egg-shaped outline, a broad and gently arched cephalon, modestly sized eyes, and a thorax adorned with a procession of finely spined segments. 

Twelve thoracic segments form its articulated middle, each bearing spines that lengthen progressively toward the ninth before tapering again—a subtle rhythm of form that feels almost architectural in its precision.

Asaphids themselves tell a longer, more dramatic story. Emerging in the Cambrian and flourishing into the Ordovician, they diversified into six superfamilies—Anomocaroidea, Asaphoidea, Cyclopygoidea, Dikelocephaloidea, Remopleuridoidea and Trinucleioidea—each experimenting with variations on a successful marine design. 

Some evolved remarkable visual adaptations, including the long-stalked eyes of Asaphus kowalewskii, which would have lifted their gaze above the seafloor haze, scanning for both prey and peril in the shifting Ordovician seas.

By the close of the Ordovician, a great extinction event swept away five of these six lineages, claiming roughly 60% of marine life. Only the resilient Trinucleioidea persisted, carrying the torch a little further into the Silurian before another global upheaval drew the final curtain on the Asaphida (Fortey & Chatterton, 1988).

Returning to our Kootenay traveller, the cephalon of Orygmaspis is parabolic, less than twice as wide as long, with a well-defined glabella—the central raised axis—measuring roughly three-quarters as wide as it is long. Its surface is modestly convex, tapering forward with faint lateral furrows and a clearly expressed occipital ring marking the posterior boundary. The preglabellar field is short, about a quarter the length of the glabella, giving the headshield a compact, purposeful look.

The eyes, small but well placed, sit between the anterior and mid-length of the glabella, positioned about one-third of the way out from the axis. Surrounding cheeks—the fixigenae and librigenae—are relatively flat, divided by facial sutures that trace an elegant path: diverging just before the eyes, running parallel near the border, then sweeping inward again in a graceful convergence. 

Behind the eyes, these sutures arc outward and back at roughly 45°, cutting the posterior margin in classic opisthoparian fashion.

At the rear, a diminutive pygidium—just a third the width of the cephalon—completes the form. It is twice as wide as long, with a central axis composed of up to four rings that nearly reach the margin. The pleural fields are gently expressed, their segmentation subdued, while the posterior edge carries three to four pairs of spines, each diminishing toward the rear like the final notes of a fading refrain.

Altogether, Orygmaspis spinula is a study in balance—armoured, yes, but refined. A small, spined voyager from Cambrian seas, preserved in stone and beautifully calcified yet still whispering of movement, adaptation, and survival in a world more than half a billion years removed from our own.

The fingers you see holding this specimen are those of the deeply awesome Chris Jenkins. If you're reading this, Chris, I owe you a visit!

SQUAKING BY THE SEA: SEAGULLS: T'SIK'WI

A gull cries in protest at not getting his share of a meal

Many of us have the good fortune to live near the sea. It is one of the places I seek out to reset my energy and soak up the atmosphere.

I love the feeling of the wind on my face as I take my best-loved path down towards the water —the sand and shells under my feet.

In those moments, the foreshore is alive with the harsh, laughing cries of seagulls, their calls slicing through the steady hush of the tide. 

Wings flash white in the sunlight as they wheel and dive, squabbling over scraps, webbed feet slapping wet sand with a slap-slap before they lift again. The air is thick with the briny tang of seaweed and salt, mingled with the faint sourness of rotting kelp and shells cracked open by the tide. 

Each wave leaves behind a shining film on the rocks, and the gulls pick and probe at it with sharp yellow beaks, clattering and clucking in between their shrieks. The smell of the ocean mixes with the dry, feathery musk of the birds themselves, grounding the scene in a rhythm as ancient as the sea. This is the domain of the seagulls who call these shores home. 

Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. The Laridae are known from not-yet-published fossil evidence from the Early Oligocene — 30–33 million years ago. 

Three gull-like species were described by Alphonse Milne-Edwards from the early Miocene of Saint-Gérand-le-Puy, France. 

Another fossil gull from the Middle to Late Miocene of Cherry County, Nebraska, USA, has been placed in the prehistoric genus Gaviota. 

These fossil gulls, along with undescribed Early Oligocene fossils are all tentatively assigned to the modern genus Larus. Among those of them that have been confirmed as gulls, Milne-Edwards' "Larus" elegans and "L." totanoides from the Late Oligocene/Early Miocene of southeast France have since been separated in Laricola.

Gulls are most closely related to the terns in the family Sternidae and only distantly related to auks, skimmers and distantly to waders. 

A historical name for gulls is mews, which is cognate with the German möwe, Danish måge, Swedish mås, Dutch meeuw, Norwegian måke/måse and French mouette. We still see mews blended into the lexicon of some regional dialects.

In the Kwak̓wala language of the Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, gulls are known as t̕sik̕wi. Most folk refer to gulls from any number of species as seagulls. This name is a local custom and does not exist in the scientific literature for their official naming. Even so, it is highly probable that it was the name you learned for them growing up.

If you have been to a coastal area nearly everywhere on the planet, you have likely encountered gulls. They are the elegantly plumed but rather noisy bunch on any beach. You will recognize them both by their size and colouring. 

Gulls are typically medium to large birds, usually grey or white, often with black markings on the head or wings. 

They typically have harsh shrill cries and long, yellow, curved bills. Their webbed feet are perfect for navigating the uneven landscape of the foreshore when they take most of their meals. 

Most gulls are ground-nesting carnivores that take live food or scavenge opportunistically, particularly the Larus species. 

Food often includes crab, clams (which they pick up, fly high and drop to crack open), fish and small birds. Gulls have unhinging jaws which allow them to consume large prey which they do with gusto. 

Their preference is to generally live along the bountiful coastal regions where they can find food with relative ease. Some prefer to live more inland and all rarely venture far out to sea, except for the kittiwakes. 

The larger species take up to four years to attain full adult plumage, but two years is typical for small gulls. Large white-headed gulls are typically long-lived birds, with a maximum age of 49 years recorded for the herring gull.

Gulls nest in large, densely packed, noisy colonies. They lay two or three speckled eggs in nests composed of vegetation. The young are precocial, born with dark mottled down and mobile upon hatching. Gulls are resourceful, inquisitive, and intelligent, the larger species in particular, demonstrating complex methods of communication and a highly developed social structure. Many gull colonies display mobbing behaviour, attacking and harassing predators and other intruders. 

Certain species have exhibited tool-use behaviour, such as the herring gull, using pieces of bread as bait with which to catch goldfish. Many species of gulls have learned to coexist successfully with humans and have thrived in human habitats. 

Others rely on kleptoparasitism to get their food. Gulls have been observed preying on live whales, landing on the whale as it surfaces to peck out pieces of flesh. They are keen, clever and always hungry. Near where I live along the west coast, I hear their calls and they always bring a smile to my day.

ANCIENT AMBUSH KILLER: MACHAIRODUS

Saber-Toothed Cat, Machairodus aphanistus

The skull before you lies cradled in a glass case at the Museo Nacional de Ciencias Naturales in Madrid, Spain.

This museum—one of my most cherished anywhere in the world—houses extraordinary treasures in the heart of a city I adore.

Even at a distance, the skull seems almost unreal, its sweeping lines and lethal symmetry more like an artifact of myth than a product of natural selection.

The upper canines of Machairodus aphanistus sweep downward in a deadly curve, their bases thick and reinforced, their blades tapering into elegant, murderous crescents. 

Grooves along their sides lighten the teeth without robbing them of strength, an evolutionary compromise that allowed this ancient predator to deliver precise, slicing blows. The zygomatic arches flare outward with commanding confidence, a testament to the enormous jaw muscles that once powered the bite. Even the wide nasal opening hints at a creature ruled by scent, finely attuned to the faintest whispers of prey on a warm Miocene wind.

This skull—stripped of flesh, muscle, and fur—remains a vivid record of a predator that walked the Earth between nine and five million years ago, long before the saber-toothed icons of the Americas made their mark. 

Machairodus aphanistus lived in the shifting landscapes of the Late Miocene, a time when Europe and western Asia were giving way to broader grasslands and open woodlands. Forest canopies receded. Herds grew larger and faster. Predators had to adapt or perish, and Machairodus responded with a design both beautiful and deadly.

Unlike its more famous descendant, Smilodon, with its compact body and powerful forelimbs, Machairodus moved with the grace of a panther. It was long-limbed and athletic, relying on bursts of speed and stealth to launch an ambush. But its skull tells a more nuanced story—one of tension between speed and specialization. The tall sagittal crest reveals a powerhouse of jaw muscles anchoring deep into the bone. 

The forward-facing orbits provide the stereoscopic vision needed to track prey with extraordinary accuracy. The sheer length of the canines required a jaw capable of opening nearly ninety degrees, a gape far wider than that of any modern cat, allowing those great blades to descend unobstructed into vulnerable regions like the throat.

You will be relieved to hear that our ancestors did not hunt and were not hunted by this impressive predator. Machairodus aphanistus went extinct in the Late Miocene, roughly 5–9 million years ago.

The earliest members of the human lineage (Homo) did not appear until about 2.8 million years ago, in the early Pleistocene. Even our more ancient relatives—Australopithecines—don’t show up until 4–4.5 million years ago.

So there is a gap of millions of years between the disappearance of Machairodus and the emergence of anything that could be considered human or human-adjacent. For that, I think we can all breath a collective sigh.

Still, others were alive on the plains that were their hunting grounds. Both hunters and prey.

In the warm, open savannas of the Miocene, the world of Machairodus was alive with competition. Packs of early hyenas honed their bone-crushing skills. Bear-dogs patrolled the river valleys. Other machairodonts—kin, rivals, or both—shared the same hunting grounds. 

The herbivores were just as diverse: early horses galloped across the plains in tight herds, while rhinocerotids, camelids, and horned antelope moved in cautious groups, ever aware of shadows that shifted in the tall grass. To survive in this dynamic ecosystem, Machairodus embraced an ambush strategy refined over countless generations. It would stalk silently, using shrubs, boulders, or dim forest edges for cover. 

When the distance closed, it lunged with explosive force, using its muscular forelimbs to pin or destabilize its prey before delivering a swift, slicing bite to the neck. Death came quickly—less by crushing force and more by catastrophic blood loss.

There is a sense, looking at its skull that you are seeing an evolutionary idea mid-transformation.

Machairodus aphanistus stands at a pivotal moment in the story of the saber-toothed cats. Its body remained agile and panther-like, but its cranial features were edging ever closer to the extreme adaptations that would define later giants like Homotherium and Smilodon. It represents a crucial chapter in which nature was experimenting, refining, and pushing the boundaries of what a predator could become.

The skull contains all of this history within its bone: the open grasslands, the pounding hooves of prey, the quiet tension of ambush, and the relentless arms race that shaped predator and prey alike. 

In its silence, it speaks. It tells of a world both familiar and wild, a world where the line between beauty and brutality was sharpened to a sabre’s edge.

A MASSIVE AMMONITE THE SIZE OF A CAR: THE FERNIE AMMONITE

Titanites occidentalis, Fernie Ammonite

The Fernie ammonite—long known as Titanites occidentalis—has officially been given a new name: Corbinites occidentalis, a fresh genus erected after a meticulous re-evaluation of this Western Giant’s anatomy and lineage. 

What hasn’t changed is its breathtaking presence high on Coal Mountain near Fernie, British Columbia, where this colossal cephalopod has rested for roughly 150 million years.

This extraordinary fossil belongs to the family Lithacoceratinae within the ataxioceratid ammonites. 

Once thought to be a close cousin of the great Titanites of Dorset, new material—including two additional large specimens discovered at nearby mine sites—reveals ribbing patterns and growth-stage features that simply didn’t match Titanites. 

With these multiple overlapping growth stages finally available, paleontologists had the missing pieces needed to correct its identity.

So, Titanites occidentalis no more—meet Corbinites occidentalis, a giant ammonite likely endemic to the relatively isolated early Alberta foreland basin of the Late Jurassic.

Fernie, British Columbia, Canada

The Fernie ammonite is a carnivorous cephalopod from the latest Jurassic (Tithonian). 

The spectacular individual on Coal Mountain measures 1.4 metres across—about the size of a small car tire and absolutely staggering when you first see it hugged by the mountainside.

The first specimen, discovered in 1947 by a British Columbia Geophysical Society mapping team at Coal Creek, was initially mistaken for a “fossil truck tire.” 

Fair enough—if a truck tire had been forged in the Jurassic and left on a mountaintop. It was later described by GSC paleontologist Hans Frebold, who gave it the name Titanites occidentalis, inspired by the giant ammonites of Dorset. 

For decades, that name stuck, even though paleontologists suspected the attribution was shaky due to poor preservation of the holotype’s inner whorls.

Recent discoveries of two additional specimens at Teck Resources’ Coal Mountain Mine finally provided the evidence needed for reassessment. 

With intact inner whorls and beautifully preserved ribbing—including hallmark variocostate and ataxioceratoid ornamentation—researchers Terence P. Poulton and colleagues demonstrated that the Canadian ammonite does not belong in Titanites. 

Their work (Volumina Jurassica, 2023) established Corbinites as a brand-new genus, with C. occidentalis as its type and only known species.

These specimens—one exceeding a metre, another about 64 cm—confirm a resident ammonite population within this basin. And as of now, these giants are unique to Western Canada.

A Journey Up Coal Mountain

If you’re keen to meet Corbinites occidentalis in the wild, you’ll want to head to Fernie, in southeastern British Columbia, close to the Alberta border. 

As your feet move up the hillside, you can imagine this land 10,000 years ago, rising above great glaciers. Where footfalls trace the steps of those that came before you. This land has been home to the Yaq̓it ʔa·knuqⱡi ‘it First Nation and Ktunaxa or Kukin ʔamakis First Nations whose oral history have them living here since time immemorial. Like them, take only what you need and no more than the land offers — packing out anything that you packed in. 

Active logging in the area since 2021 means that older directions are now unreliable—trailheads have shifted, and a fair bit of bushwhacking is the price of admission. Though clear-cutting reshaped the slope, loggers at CanWel showed admirable restraint: they worked around the fossil, leaving it untouched.

The non-profit Wildsight has been championing efforts to protect the ammonite, hoping to establish an educational trail with provincial support and possible inclusion under the Heritage Conservation Act—where the fossil’s stewardship could be formally recognised.

HIKING TO THE FERNIE AMMONITE (IMPORTANT UPDATE: TRAIL CLOSED)

From the town of Fernie, British Columbia, you would traditionally head east along Coal Creek Road toward Coal Creek, with the ammonite site sitting 3.81 km from the road’s base as the crow flies. 

The classic approach begins at a roadside exposure of dark grey to black Cretaceous plant fossils, followed by a creek crossing and a steep, bushwhacking ascent.

However — and this is critical — the trail is currently closed.

The entire access route runs straight through an area of active logging, and conditions on the slope are extremely dangerous. Between heavy equipment, unstable cutblocks, and altered drainages, this is not a safe place for hikers right now.

Conservation groups, including Wildsight, continue working toward restoring safe public access and formalising the site under the Heritage Conservation Act. 

Their long-term goal is to reopen the trail as a designated educational hike with proper signage, but at present, the route should not be attempted. 

Once logging operations move out of the area and safety assessments are done, the possibility of reopening may return.

For now, the safest—and strongly recommended—way to view this iconic fossil is via the excellent cast on display at the Courtenay & District Museum on Vancouver Island, or at the Visitor Information Centre in Sparwood.

Photo credit: Vince Mo Media. Vince is an awesome photographer and drone operator based in Fernie, BC. Check out his work (and hire him!) by visiting his website at vmmedia.ca.

THIRST OF THE LOST CONTINENT: DODOS AT THE RIVER OF MAURITIA

Dodo Birds by Daniel Eskridge

Two dodo birds—one warm brown like sun-baked coconut husk, the other a pale, ghostly white with hints of grey—stand beak-deep in the shallows of a river that winds like a silver serpent through the tropical jungles of ancient Mauritia. 

Their feet sink into cool silt and damp leaves at a rivers edge. 

The air is thick with the scent of pandanus and damp leaves, heavy enough to taste. Dragonflies hover in lazy spirals above them, iridescent flashes stitching over the water’s skin.

The brown male dodo dips first, scooping up a beakful of water with a gentle glop, while the white female one pauses, head cocked, watching a fruit drift downstream. For a moment the world feels impossibly quiet—no humans, no predators bold enough to trouble them, only the chorus of the forest and the steady rhythm of their drinking.

These feathered oddities belong to an island that itself has slipped through time. Mauritia, a now-lost microcontinent once nestled between Madagascar and India, cracked and sank more than 60 million years ago as the Indian Ocean spread and rearranged the world’s geography. All that remains today are a few scattered fragments—Mauritius, Réunion, Rodrigues—emerald crumbs left atop an ancient submerged landmass.

Dodo Birds by Daniel Eskridge

It is on one of these volcanic islands, long after Mauritia’s foundering, that the dodo evolved into its peculiar glory. 

Descended from flighted pigeons that likely swept in on storm winds from Southeast Asia, the dodo abandoned the sky entirely. 

With no natural predators and an island full of fruits, nuts, and fallen seeds, wings became more decorative than practical. Their legs grew stout. Their bodies rounded. 

Their beaks curved into the iconic hooked silhouette now etched into the imagination of every natural historian.

The brown dodo nudges the white one aside, perhaps a sign of affection, perhaps mild irritation—dodos, after all, were social birds, not the clumsy caricatures drawn centuries later. 

They waddled in flocks, nested on the ground, and lived comfortably beneath the canopy of ebony forests. Their feathers, described by early visitors as soft and hair-like, varied from gray-brown to white depending on age, sex, and perhaps even seasonal cycles.

But their peace was fragile, vulnerable to change they could not see coming.

When humans finally set foot on Mauritius in the late 1500s, they brought ships that carried pigs, rats, goats, and monkeys, all eager for eggs, seedlings, and anything edible. 

Forests were cut, nests trampled, and the trusting dodos, unaccustomed to fear, walked directly into the hands of sailors who considered them a convenient, if not particularly tasty, meal. Within roughly a century, they were gone.

But in this imagined moment—two birds drinking from a clear jungle river on an island born from a drowned continent—they live again. 

The sun breaks through a gap in the canopy, scattering gold across their backs. The white dodo lifts its head, droplets falling like tiny jewels, and lets out a soft, throaty grunt.

Here, in the cool breath of Mauritia’s shadowed past, the dodos are a symbol of loss—curious, gentle, utterly at home.

And for a heartbeat, we remember them.

Illustration Credit: This image was created by the supremely talented Daniel Eskridge, Paleo Illustrator from Atlanta, Georgia, USA. I share it here with permission as I have licensed the use of many of his images over the years, including this one. 

To enjoy his works (and purchase them!) to adorn your walls, visit his website at www.danieleskridge.com

SHAGGY TITANS OF THE GRASSLANDS: BISON

Bison move across the prairie like living storms, vast and steady, with the weight of centuries in their stride. 

Their dark eyes hold a quiet, unwavering depth—as if they’ve looked into the heart of time itself and carry its secrets in silence. Look into the eyes of this fellow and tell me you do not see his deep intelligence as he gives the camera a knowing look.

Shaggy fur ripples in the wind, rich and earthy, brushed by sun and shadow, a cloak woven from wilderness. When they breathe, clouds rise in the cold air, soft and ephemeral, like whispered promises that vanish but leave warmth behind.

There is something profoundly romantic in their presence: strength wrapped in gentleness, endurance softened by grace.  To watch them is to feel the wild itself lean closer, reminding us of a love as vast as the horizon, as eternal as the ground beneath our feet.

When we think of bison today, images of great herds roaming the North American plains come to mind—dark, shaggy shapes against sweeping prairies. But the story of bison goes back far deeper in time. 

These massive grazers are part of a lineage that stretches millions of years into the past, their fossil record preserving the tale of their rise, spread, and survival.

Bison belong to the genus Bison, within the cattle family (Bovidae). Their story begins in Eurasia during the late Pliocene, around 2.6 million years ago, when the first true bison evolved from earlier wild cattle (Bos-like ancestors). 

Fossils suggest they descended from large bovids that roamed open grasslands of Eurasia as forests retreated and cooler, drier climates expanded.

The earliest known species, Bison priscus, or the Steppe Bison, was a giant compared to modern bison, sporting long horns that could span over six feet tip to tip. These animals thrived across Europe, Asia, and eventually crossed into North America via the Bering Land Bridge during the Pleistocene Ice Age.

The fossil record of bison stretches back about 2 million years in Eurasia and at least 200,000 years in North America, where they became one of the most successful large herbivores of the Ice Age. Fossil evidence shows that at least seven different species of bison once lived in North America, including the iconic Bison latifrons with its massive horns, and Bison antiquus, which is considered the direct ancestor of the modern American bison (Bison bison).

Some of the richest fossil bison deposits come from Siberia and Eastern Europe – home to abundant Bison priscus fossils, often preserved in permafrost with soft tissues intact. They are also found in Alaska, USA and in Canada's Yukon region – where Ice Age bison fossils are found alongside mammoth, horse, and muskox remains.

The Great Plains of the United States and Canada are rich in Bison antiquus and later species, often in mass bone beds where entire herds perished. We also find their remains in California and the American Southwest at sites like the La Brea Tar Pits. La Brea preserves bison remains from the Late Pleistocene and their museum of the same name has a truly wonderful display of Pleistocene wolves. Definitely worthy of a trip!

One particularly famous fossil site is the Hudson-Meng Bison Kill Site in Nebraska, where remains of over 600 Bison antiquus dating to about 10,000 years ago provide a window into Ice Age hunting practices and herd behavior.

By the end of the Ice Age, many megafauna species disappeared, but bison endured. Bison antiquus gradually gave rise to the modern American bison (Bison bison), which still carries echoes of its Ice Age ancestors. Though smaller than their Pleistocene relatives, today’s bison remain the largest land mammals in North America.

BACK IN THE USSR: KEPPLERITES

This glorious chocolate block contains the creamy grey ammonite Kepplerites gowerianus (Sowerby 1827) with a few invertebrate friends, including two brachiopods: Ivanoviella sp., Zeilleria sp. and the deep brown gastropod Bathrotomaria sp. 

There is also a wee bit of petrified wood on the backside.

These beauties hail from Jurassic, Lower Callovian outcrops in the Quarry of Kursk Magnetic Anomaly (51.25361,37.66944), Kursk region, Russia. Diameter ammonite 70мм. 

In the mid-1980s, during the expansion and development of one of the quarries, an unusual geological formation was found. This area had been part of the seafloor around an ancient island surrounded by Jurassic Seas. 

The outcrops of this geological formation turned out to be very rich in marine fossil fauna. This ammonite block was found there years ago by the deeply awesome Emil Black. 

In more recent years, the site has been closed to fossil collecting and is in use solely for the processing and extraction of iron ore deposits. Kursk Oblast is one of Russia's major producers of iron ore. The area of the Kursk Magnetic Anomaly has one of the richest iron-ore deposits in the world. Rare Earth minerals and base metals also occur in commercial quantities in several locations. Refractory loam, mineral sands, and chalk are quarried and processed in the region. 

The Kursk Magnetic Anomaly Quarry is not far from the Sekmenevsk Formation or Sekmenevska Svita in Russian, a Cretaceous (Albian to Cenomanian) terrestrial geologic formation where Pterosaur fossils have been found in the sandstones. 

OIL IN WATER BEAUTY: FOSSILS OF FOLKSTONE

Sheer beauty — a beautiful Euhoplites ammonite from Folkstone, UK. I've been really enjoying looking at all oil-in-water colouring and chunkiness of these ammonites.

Euhoplites is an extinct ammonoid cephalopod from the Lower Cretaceous, characterized by strongly ribbed, more or less evolute, compressed to inflated shells with flat or concave ribs, typically with a deep narrow groove running down the middle.

In some, ribs seem to zigzag between umbilical tubercles and parallel ventrolateral clavi. In others, the ribs are flexious and curve forward from the umbilical shoulder and lap onto either side of the venter.

Its shell is covered in the lovely lumps and bumps we associate with the genus. The function of these adornments are unknown. I wonder if they gave them greater strength to go deeper into the ocean to hunt for food. 

They look to have been a source of hydrodynamic drag, likely preventing Euhoplites from swimming at speed. Studying them may give some insight into the lifestyle of this ancient marine predator. Euhoplites had shells ranging in size up to a 5-6cm. 

We find them in Lower Cretaceous, middle to upper Albian age strata. Euhoplites has been found in Middle and Upper Albian beds in France where it is associated respectively with Hoplites and Anahoplites, and Pleurohoplites, Puzosia, and Desmoceras; in the Middle Albian of Brazil with Anahoplites and Turrilites; and in the Cenomanian of Texas.

This species is the most common ammonite from the Folkstone Fossil Beds in southeastern England where a variety of species are found, including this 37mm beauty from the collections of José Juárez Ruiz.

QUENSTEDTOCERAS WITH PATHOLOGY

What you are seeing here is a protuberance extruding from the venter of Quenstedtoceras cf. leachi (Sowerby). It is a pathology in the shell from hosting immature bivalves that shared the seas with these Middle Jurassic, Upper Callovian, Lamberti zone fauna from the Volga River basin. 

The collecting site is the now inactive Dubki commercial clay quarry and brickyard near Saratov, Russia. 

The site has produced thousands of ammonite specimens. A good 1,100 of those ended up at the Black Hills Institute of Geological Research in Hill City, South Dakota. 

Roughly 1,000 of those are Quenstedtoceras (Lamberticeras) lamberti and the other 100 are a mix of other species found in the same zone. These included Eboraciceras, Peltoceras, Kosmoceras, Grossouvria, Proriceras, Cadoceras and Rursiceras. 

What is especially interesting is the volume of specimens — 167 Quenstedtoceras (Lamberticeras) lamberti and 89 other species in the Black Hills collection — with healed predation injuries. It seems Quenstedtoceras (Lamberticeras) lamberti are the most common specimens found here and so not surprisingly the most common species found injured. 

Of the 1,000, 655 of the Quenstedtoceras (Lamberticeras) lamberti displayed some sort of deformation or growth on the shell or had grown in a tilted manner. 

Again, some of the Q. lamberti had small depressions in the centre likely due to a healed bite and hosting infestations of the immature bivalve Placunopsis and some Ostrea. 

The bivalves thrived on their accommodating hosts and the ammonites carried on, growing their shells right up and over their bivalve guests. 

This relationship led to some weird and deformities of their shells. They grow in, around, up and over nearly every surface of the shell and seem to have lived out their lives there. It must have gotten a bit unworkable for the ammonites, their shells becoming warped and unevenly weighted. 

Over time, both the flourishing bivalves and the ammonite shells growing up and over them produced some of the most interesting pathology specimens I have ever seen.    

In the photo here from Emil Black, you can see some of the distorted shapes of Quenstedtoceras sp. 

Look closely and you see a trochospiral or flattened appearance on one side while they are rounded on the other. 

All of these beauties hail from the Dubki Quarry near Saratov, Russia. The ammonites were collected in marl or clay used in brick making. The clay particles suggest a calm, deep marine environment. 

One of the lovely features of the preservation here is the amount of pyrite filling and replacement. It looks like these ammonites were buried in an oxygen-deficient environment. 

The ammonites were likely living higher in the water column, well above the oxygen-poor bottom. An isotopic study would be interesting to prove this hypothesis. 

There's certainly enough of these ammonites that have been recovered to make that possible. It's estimated that over a thousand specimens have been recovered from the site but that number is likely much higher. But these are not complete specimens. We mostly find the phragmocones and partial body chambers. Given the numbers, this may be a site documenting a mass spawning death over several years or generations.

If you fancy a read on all things cephie, consider picking up a copy of Cephalopods Present and Past: New Insights and Fresh Perspectives edited by Neil Landman and Richard Davis. Figure 16.2 is from page 348 of that publication and shows the hosting predation quite well. 

Photos: Courtesy of the deeply awesome Emil Black. These are in his personal collection that I hope to see in person one day. 

It was his sharing of the top photo and the strange anomaly that had me explore more about the fossils from Dubki and the weird and wonderful hosting relationship between ammonites and bivalves. Thank you, my friend!

ZENASPIS: DEVONIAN FISH MORTALITY PLATE

A Devonian fish mortality plate showing all lower shields of Zenaspis podolica (Lankester, 1869) and Stensiopelta pustulata (or Victoraspis longicornualis) from Lower Devonian deposits of Podolia, Ukraine.

Zenaspis is an extinct genus of jawless fish which existed during the early Devonian period. Due to it being jawless, Zenaspis was probably a bottom feeder.

The lovely 420 million-year-old plate you see here is from Podolia or Podilia, a historic region in Eastern Europe, located in the west-central and south-western parts of Ukraine, in northeastern Moldova. 

Podolia is the only region in Ukraine where Lower Devonian remains of ichthyofauna can be found near the surface.

For the past 150 years, vertebrate fossils have been found in more than 90 localities situated in outcrops along banks of the Dniester River and its northern tributaries, and in sandstone quarries. 

At present faunal list of Early Devonian agnathans and fishes from Podolia number 72 species, including 8 Thelodonti, 39 Heterostraci, 19 Osteostraci, 4 Placodermi, 1 Acanthodii, and 1 Holocephali (Voichyshyn 2001a, modified).

In Podolia, Lower Devonian redbeds strata (the Old Red Formation or Dniester Series) can be found up to 1800 m thick and range from Lochkovian to Eifelian in age (Narbutas 1984; Drygant 2000, 2003). 

In the lower part (Ustechko and Khmeleva members of the Dniester Series) they consist of multicoloured, mainly red, fine-grained cross-bedded massive quartz sandstones and siltstones with seams of argillites (Drygant 2000).

We see fossils beds of Zenaspis in the early Devonian of Western Europe. Both Zenaspis pagei and Zenaspis poweri can be found up to 25 centimetres long in Devonian outcrops of Scotland.

Reference: Voichyshyn, V. 2006. New osteostracans from the Lower Devonian terrigenous deposits of Podolia, Ukraine. Acta Palaeontologica Polonica 51 (1): 131–142. Photo care of Fossilero Fisherman.

BACK IN THE USSR: BEADANTICERAS OF THE NORTHERN CAUCASUS

This lovely oil in water coloured ammonite is the beauty Beudanticeras sp. from the Lower Cretaceous (Upper Aptian), Krasnodar region, Northern Caucasus, southern Russia. 

This area of the world has beautiful fossil specimens with their distinct colouring. The geology and paleontological history of the region are fascinating as is its more recent history. 

The territory of present Krasnodar Krai was inhabited as early as the Paleolithic, about 2 million years ago. It was inhabited by various tribes and peoples since ancient times. 

There were several Greek colonies on the Black Sea coast, which later became part of the Kingdom of the Bosporus. In 631, the Great Bulgaria state was founded in the Kuban. In the 8th-10th centuries, the territory was part of Khazaria.

In 965, the Kievan Prince Svyatoslav defeated the Khazar Khanate and this region came under the power of Kievan Rus, Tmutarakan principality was formed. At the end of the 11th century, in connection with the strengthening of the Polovtsy and claims of Byzantium, Tmutarakan principality came under the authority of the Byzantine emperors (until 1204).

In 1243-1438, this land was part of the Golden Horde. After its collapse, Kuban was divided between the Crimean Khanate, Circassia, and the Ottoman Empire, which dominated in the region. Russia began to challenge the protectorate over the territory during the Russian-Turkish wars.

In 1783, by decree of Catherine II, the right-bank Kuban and Taman Peninsula became part of the Russian Empire after the liquidation of the Crimean Khanate. 

In 1792-1793, Zaporozhye (Black Sea) Cossacks resettled here to protect new borders of the country along the Kuban River. 

During the military campaign to establish control over the North Caucasus (Caucasian War of 1763-1864), in the 1830s, the Ottoman Empire for forced out of the region and Russia gained access to the Black Sea coast.

Prior to the revolutionary events of 1917, most of the territory of present Krasnodar Krai was occupied by the Kuban region, founded in 1860. In 1900, the population of the region was about 2 million people. In 1913, it ranked 2nd by the gross harvest of grain, 1st place for the production of bread in the Russian Empire.

The Kuban was one of the centres of resistance after the Bolshevik revolution of 1917. In 1918-1920, there was a non-Bolshevik Kuban People’s Republic. In 1924, North-Caucasian krai was founded with the centre in Rostov-on-Don. In 1934, it was divided into Azov-Black Sea krai (Rostov-on-Don) and North Caucasus krai (Stavropol).

September 13, 1937, the Azov-Black Sea region was divided into the Rostov region and Krasnodar Krai that included Adygei autonomous oblast. During the Second World War, the region was captured by the Germans. After the battle for the Caucasus, it was liberated. There are about 1,500 monuments and memorials commemorating heroes of the war on the territory of Krasnodar Krai.

The lovely block you see here is in the collections of the awesome John Fam, Vice-Chair of the Vancouver Paleontological Society in British Columbia, Canada.