CARNOTAURUS: FLESH-EATING BULL

Carnotaurus sastrei, a genus of large theropod dinosaurs that roamed the southern tip of Argentina, South America during the Late Cretaceous, 72 to 69.9 million years ago. His name means "flesh-eating bull,' and he lives up to it.

This fellow — or at least his robust skull with the short, knobby eyebrow horns and fierce-looking teeth — is on display at the Natural History Museum in Madrid, Spain. For now, he is the only known genus of this species of bipedal predator.

The first specimen of Carnotaurus sastrei was found in Chubut on vast plains between the Andes Mountains and the Atlantic Ocean. A physician, Dr. A'ngel Tailor noticed a large concretion showing some bone fragments. A team led by José F. Bonaparte excavated the find in 1984 as part of a paleontological expedition funded by the Argentine Museum of Natural Sciences.

Sadly, Bonaparte — the Maestro del Mesozoico — passed away the 18th February 20220 at the age of 91. He spent the majority of his career as head of the Vertebrate Palaeontology Division of the Museo Argentino de Ciencias Naturales “Bernardino Rivadavia,” in Buenos Aires. Bonaparte opened up the vertebrate finds of Argentina to the world. He was instrumental in the finding, excavating and naming many iconic dinosaurs — Carnotaurus, Amargasaurus, Abelisaurus, Argentinosaurus, Noasaurus along with the finding of the first fossilised remains of Mesozoic South American mammals. He mentored many palaeontologists who will miss his keen eye and tremendous work ethic — Luis Chiappe, Rodolfo Coria, Agustín Martinelli, Fernando Novas, Jaime Powell, Guillermo Rougier, Leonardo Salgado, Sebastián Apesteguía and many others.

His excavation of Carnotaurus was the first of its kind and he recognized that the skull is quite unusual. Initially, it has a very marine reptile feel — but make no mistake this guy is clearly a terrestrial theropod. He had smallish, underdeveloped arms — teeny by theropod standards. Once you look closer you see his bull-like horns from whence he gets his name — horns that imply battle between rivals for the best meal, sexual partner and to be the one who leads the herd. 

He was covered in leathery skin lined with rows of cone-shaped nodules or bumps. These get larger as they move towards his spine. He had forward-facing eyes, similar to tyrannosaurs like T-rex and smaller theropods like Velociraptor and Troodon — who had better vision even that T-rex — which would have given him the advantage of binocular vision and depth perception. Forward-facing eyes are also quite helpful with nocturnal hunting — think owls and cats — as they take in more light and help with nighttime predation. So perhaps this flesh-eating bull fancied a late-night snack on his menu from time to time.

Species like squirrels, pigeons and crocodiles have eyes on the sides of their heads. They lack the important competitive feature of well-developed depth perception — being able to easily and estimate distance — but perhaps make up for it with a panorama that offers a wider field of view.   

HO HO, OH! IS THAT A DINOSAUR TRACK?

This is a theropod track left by an Allosaurus walking through a muddy bank that turned to stone. These big beasties were large carnosaurian theropod dinosaurs that lived 155 to 145 million years ago.

This track and many others are at Dinosaur Ridge, a segment of the Dakota Hogback in the Morrison Fossil Area National Natural Landmark located in Jefferson County, Colorado, near the town of Morrison and just west of Denver.

The Dinosaur Ridge area is one of the world's most famous dinosaur fossil localities. Along with the dinosaur bones, you find plant fossils, crocodile bones and a variety of reptile tracks. 

In 1877, fossil excavation began at Dinosaur Ridge under the direction of palaeontologist Othniel Charles Marsh. Some of the best-known dinosaurs were found here, including Stegosaurus, Apatosaurus, Diplodocus, and Allosaurus. 

In 1973, the area was recognized for its uniqueness as well as its historical and scientific significance when it was designated the Morrison Fossil Area National Natural Landmark by the National Park Service. In 1989, the Friends of Dinosaur Ridge was formed to address increasing concerns regarding the preservation of the site and to offer educational programs on the area's resources.

Visit Dinosaur Ridge Morrison Fossil Area National Natural Landmark

Fancy a trip? You can visit these wonderful tracks, the Dinosaur Ridge Exhibit Hall and walk the trails through the tracks. They have put up helpful interpretive signs that explain the local geology, a volcanic ash bed, trace fossils, paleo-ecology, economic development of coal, oil and clay, and many other geologic and paleontological features.

There are two trails and a visitor centre at Dinosaur Ridge. The visitor centre features information on trails and a small gift shop. If you do not pack a lunch, you'll want to visit the Stegosaurus Snack Shack located outside the visitor centre offering coffee, cold drinks, burritos, pretzels and more. It is open from 10am-4pm Mon-Sat and 11am-4pm on Sun from June through August. 

Check updates on their website concerning Covid-19 restrictions. This is a vast site with easy trails so it makes for a great family trip when museums and other indoor facilities are closed.

Know Before You Go

Uncover Colorado: https://www.uncovercolorado.com/landmarks/morrison-golden-fossil-areas-dinosaur-ridge/

AWKWARD AND AWESOME: DIMORPHODON

This remarkable fellow is Dimorphodon — a genus of medium-sized pterosaur from the Early Jurassic. He is another favourite of mine for his charming awkwardness.

You can see this fellow's interesting teeth within his big, bulky skull. Dimorphodon had two distinct types of teeth in their jaws — an oddity amongst reptiles — and also proportionally short wings for their overall size. 

Just look at him. What an amazing beast. We understand their anatomy quite well today, but can you imagine being the first to study their fossils and try to make sense of them. 

The first fossil remains now attributed to Dimorphodon were found in England by fossil collector Mary Anning, at Lyme Regis in Dorset, the United Kingdom in December 1828. While she faced many challenges in her life, she was blessed to live in one of the richest areas in Britain for finding fossils. 

She walked the beaches way back in the early 1800s of what would become the Jurassic Coast UNESCO World Heritage Site. The Jurassic Coast holds some of the most interesting fossils ever found — particularly within the strata of the Blue Lias which date back to the Hettangian-Sinemurian. It is one of the world’s most famous fossil sites. Millions come to explore the eroding coastline looking for treasures that provide delight and inspiration to young and old.

These fossil treasures provide us with tremendous insights into our world 185 million years ago when amazing animals like Dimorphodon ruled the skies. 

Mary's specimen was acquired by William Buckland and reported in a meeting of the Geological Society on 5 February 1829. Six years later, in 1835, William Clift and William John Broderip built upon the work by Buckland to publish in the Transactions of the Geological Society, describing and naming the fossil as a new species. 

As was the case with most early pterosaur finds, Buckland classified the remains in the genus Pterodactylus, coining the new species Pterodactylus macronyx. The specific name is derived from Greek makros, "large" and onyx, "claw", in reference to the large claws of the hand. The specimen, presently NHMUK PV R 1034, consisted of a partial and disarticulated skeleton on a slab — notably lacking the skull. Buckland in 1835 also assigned a piece of the jaw from the collection of Elizabeth Philpot to P. macronyx. 

Later, the many putative species assigned to Pterodactylus had become so anatomically diverse that they began to be broken into separate genera.

In 1858, Richard Owen reported finding two new specimens, NHMUK PV OR 41212 and NHMUK PV R 1035, again partial skeletons but this time including the skulls. Having found the skull to be very different from that of Pterodactylus, Owen assigned Pterodactylus macronyx its own genus, which he named Dimorphodon. 

His first report contained no description and the name remained a nomen nudum. In 1859, however, a subsequent publication by Owen provided a description. After several studies highlighting aspects of Dimorphodon's anatomy, Owen finally made NHMUK PV R 1034 the holotype in 1874  — 185 million years after cruising our skies the Dimorphodon had finally fully arrived.

PROSAUROLOPHUS MAXIMUS

Reconstruction of Prosaurolophus maximus

Prosaurolophus maximus was a large-headed duckbill dinosaur, or hadrosaurid, in the ornithischian family Hadrosauridae.

The most complete Prosaurolophus maximus specimen had a massive skull an impressive 0.9 metres (3.0 ft) long that graced a skeleton about 8.5 metres (28 ft) long. 

He had a small, stout, triangular crest in front of his eyes. The sides of the crest are concave, forming depressions. 

The crest grew isometrically — without changing in proportion — throughout the lifetime of each individual, leading one to wonder if Prosaurolophus had had a soft tissue display structure such as inflatable nasal sacs. We see this feature in hooded seals, Cystophora cristata, who live in the central and western North Atlantic today. Prosaurolophus maximus may have used their inflatable nasal sac for a display to warn a predator or to entice the ladies, attracting the attention of a female.

When this good looking fellow was originally described by Brown, Prosaurolophus maximus was known only from a skull and jaw. Half of the skull was badly weathered at the time of examination, and the level of the parietal was distorted and crushed upwards to the side. You can imagine that these deformations in preservation created some grief in the final description.

The different bones of the skull are easily defined with the exception of the parietal and nasal bones. Brown found that the skull of the already described genus Saurolophus was very similar overall, just smaller than the skull of Prosaurolophus maximus. The unique feature of a shortened frontal in lambeosaurines is also found in Prosaurolophus maximus, and the other horned hadrosaurines Brachylophosaurus, Maiasaura, and Saurolophus. Although they lack a shorter frontal, the genera Edmontosaurus and Shantungosaurus share an elongated dentary structure.

Prosaurolophus maximus, Ottawa Museum of Nature

Patches of preserved skin are known from two juvenile specimens, TMP 1998.50.1 and TMP 2016.37.1; these pertain to the ventral extremity of the ninth through fourteenth dorsal ribs, the caudal margin of the scapular blade, and the pelvic region. 

Small basement scales (scales that make up the majority of the skin surface), 3–7 millimetres (0.12–0.28 in) in diameter, are preserved on these patches - this is similar to the condition seen in other saurolophine hadrosaurs.

More uniquely, feature scales (larger, less numerous scales which are interspersed within the basement scales) around 5 millimetres (0.20 in) wide and 29 millimetres (1.1 in) long are found interspersed in the smaller scales in the patches from the ribs and scapula (they are absent from the pelvic patches). 

Similar scales are known from the tail of the related Saurolophus angustirostris (on which they have been speculated to indicate pattern), and it is considered likely adult Prosaurolophus would've retained the feature scales on their flanks like the juveniles.

Image: Three-dimensional reconstruction of Prosaurolophus maximus. Created using the skull reconstructions in the original description as reference. (Fig. 1 and 3 in Brown 1916). According to Lull and Wright (1942), the muzzle was restored too long in its original description. The colours and/or patterns, as with nearly all reconstructions of prehistoric creatures, are speculative. Created & uploaded to Wikipedia by Steveoc 86.

APATOSAURUS: PLANT-EATING GIANTS

Apatosaurus, one of the largest animals to ever live

Apatosaurus was one of the largest animals to ever live on our planet. Picture this fellow at over 22 metres (75 feet) long and over 22,679 kg (50,000 lbs). 

That's about two-and-a-half times as long as a London bus and two-and-a-half times as heavy as a Tyrannosaurus rex. 

Given their large size, these big boys did not have to worry about most predators. It would take a posse of large apex predators like the large carnosaurian theropod dinosaur Allosaurus to take down a full-grown Apatosaurus. 

Their young would be vulnerable and their eggs even more so, but my money would be on the large Apatosaurus defending the smaller members of their family groups against these risks.

This sauropod had a really long neck and whip-like tail and a teeny, tiny head. These massive beasts liked to dine on vegetation and foliage at the tops of the tree canopies.

Othniel Charles Marsh (of Cope & Marsh fame) first described the Apatosaurus in 1877, giving it the name Apatosaurus ajax. There are two recognized species of Apatosaurus, the Apatosaurus ajax (the type species) and Apatosaurus louisae who lived in North America during the Late Jurassic — 152 million years ago.

TRACKING THEROPODS HIGH IN THE PERUVIAN ANDES

Left, right, one, two... Theropod Tracks

Left, right, one, two... the wonderfully preserved theropod trackway you see here was found by eagle-eyed construction workers blasting out a tunnel for a road near Yanashallash in the Chavin de Huantar region of Peru. 

You would be surprised how many fossils have been found this way!

The footprints are trace fossils from a big fellow who marched through here back in the Cretaceous. The inflated rust coloured prints were found alongside the fossil crocodile, pterosaurs, primitive tortoise and fish.

Antamina Mining and the Asociacion Ancash have provided funding to turn this remarkable find into an educational exhibit with a research team led by palaeontologist Carlos Vildoso. 

Vildoso along with palaeontologist Patricia Sciammaro (the two are married) founded the Instituto Peruano de Estudios en Paleovertebrados (IPEP) is a non-profit, non-government institution. Their centre focuses on vertebrate palaeontology. Over the years they have built an enviable database of significant Peruvian fossil sites and publish Contribuciones Paleontológicas, a quarterly journal devoted to vertebrate palaeontology. Chévere!

PISTA DE BAILE JURÁSICA

This busy slate grey dinosaur trackway from the Iberian Peninsula looks more like a dance floor than the thoroughfare it is. 

The numerous theropod dinosaur tracks — with a few enormous sauropod tracks thrown in for good measure — cover the entire surface. 

The local soil has a bit of rusty iron ore in it that highlights each print nicely when the soil is blown into the depressions the tracks left. 

The dinosaurs crossed this muddy area en masse sometime back in the Jurassic.

The Iberian Peninsula is the westernmost of the three major southern European peninsulas — the Iberian, Italian, and Balkan. It is bordered on the southeast and east by the Mediterranean Sea, and on the north, west, and southwest by the Atlantic Ocean. The Pyrenees mountains are situated along the northeast edge of the peninsula, where it adjoins the rest of Europe. Its southern tip is very close to the northwest coast of Africa, separated from it by the Strait of Gibraltar and the Mediterranean Sea.

The Iberian Peninsula contains rocks of every geological period from the Ediacaran to the recent, and almost every kind of rock is represented. To date, there are 127 localities of theropod fossil finds ranging from the Callovian-Oxfordian — Middle-Upper Jurassic — to the Maastrichtian (Upper Cretaceous), with most of the localities concentrated in the Kimmeridgian-Tithonian interval and the Barremian and Campanian stages. The stratigraphic distribution is interesting and suggests the existence of ecological and/or taphonomic biases and palaeogeographical events that warrant additional time and attention.

As well as theropods, we also find their plant-eating brethren. This was the part of the world where the last of the hadrosaurs, the duck-billed dinosaurs, lived then disappeared in the Latest Cretaceous K/T extinction event 65.5 million years ago.

The core of the Iberian Peninsula is made up of a Hercynian cratonic block known as the Iberian Massif. On the northeast, this is bounded by the Pyrenean fold belt, and on the southeast, it is bounded by the Baetic System. These twofold chains are part of the Alpine belt. To the west, the peninsula is delimited by the continental boundary formed by the magma-poor opening of the Atlantic Ocean. The Hercynian Foldbelt is mostly buried by Mesozoic and Tertiary cover rocks to the east but nevertheless outcrops through the Sistema Ibérico and the Catalan Mediterranean System. The photo you see here is care of the awesome Pedro Marrecas from Lisbon, Portugal. Hola, Pista de baile jurásica!

Pereda-Suberbiola, Xabier; Canudo, José Ignacio; Company, Julio; Cruzado-Caballero, Penélope; Ruiz-Omenaca, José Ignacio. "Hadrosauroid dinosaurs from the latest Cretaceous of the Iberian Peninsula" Journal of Vertebrate Paleontology 29(3): 946-951, 12 de septiembre de 2009.

Pereda-Suberbiola, Xabier; Canudo, José Ignacio; Cruzado-Caballero, Penélope; Barco, José Luis; López-Martínez, Nieves; Oms, Oriol; Ruíz-Omenaca, José Ignacio. Comptes Rendus Palevol 8(6): 559-572 septiembre de 2009.

FERRISAURUS SUSTUTENSIS: A NEW NON-AVIAN DINOSAUR IN BC

Say hello to Ferrisaurus sustutensis —  “A new leptoceratopsid dinosaur from Maastrichtian-aged deposits of the Sustut Basin, northern British Columbia, Canada."

You may recall Dr. Victoria Arbour, curator of palaeontology at the Royal BC Museum from her work on ankylosaurs & that interesting specimen from Hornby Island thought to be a pterosaur but further study revealed to be a saurodontid fish, an ambush predator with very sharp serrated teeth and elongate, torpedo-like body. Not a pterosaur but still a massively exciting find. Arbour was very gracious about the new interpretation, taking it in stride. She has since gone on to name this partial ornithischian dinosaur from Sustut Basin, as well as the ankylosaurs Zuul, Zaraapelta, Crichtonpelta, and Ziapelta. She's been a busy bee.

For this latest find, she’s partnered up & published her findings with David Evans from the Royal Ontario Museum in the peer-reviewed scientific journal PeerJ - the Journal of Life and Environmental Sciences last year. Their paper describes this partial dinosaur skeleton found amongst the inhospitable boreal forests and folded rock of the Canadian Cordillera near the Sustut Basin of northern British Columbia, Canada.

The first bones were collected by geologist Kenny F. Larsen who was surveying for uranium along the then in-construction BC Rail line along the Sustut River. The bones were later donated to Dalhousie University in Halifax, Nova Scotia then accessioned by the Royal British Columbia Museum in Victoria, BC. The skeleton includes parts of the pectoral girdles, left forelimb, left hindlimb, and right pes. Their rationale for a new species distinguished from other named leptoceratopsids is based on the proportions of the ulna and pedal phalanges.

This specimen was previously described in 2008 as an indeterminate small-bodied, bipedal neornithischian, possibly representing either a pachycephalosaur or a basal ornithopod similar to Thescelosaurus. With more material to work with, Arbour and Evans reinterpreted the remains as a leptoceratopsid ceratopsian, Ferrisaurus sustutensis, gen. et. sp. nov.

Figure 2: Preserved elements of RBCM P900

The news deserves some fanfare. While Alberta, our sister province to the east is practically littered with dinosaur remains, they are relatively rare in BC. This is the first unique non-avian dinosaur species reported from British Columbia.

It has been placed, within a reasonably resolved phylogenetic context, with Ferrisaurus recovered as more closely related to Leptoceratops than Montanoceratops. At 68.2–67.2 Ma in age, Ferrisaurus falls between, and slightly overlaps with, both Montanoceratops and Leptoceratops, and represents a western range extension for Laramidian leptoceratopsids. Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and Europe. They resembled and were closely related to, other neoceratopsians, such as Protoceratopsidae and Ceratopsidae, but they are more primitive and generally smaller.

Figure 3: Pectoral Elements of Laramidian leptoceratopsids

Back in 2017, Arbour led an expedition to the Sustut River in Northern British Columbia to relocate the site where Ferrisaurus was originally discovered forty-six years earlier in 1971 along the BC Rail line near the intersection of Birdflat Creek and the Sustut River. The expedition was a huge success as the team found the remains of this new species of dinosaur and also recovered several species of fossil plants.

The fossil plant finds may not seem that exciting in comparison to a dinosaur but Cretaceous plants in BC are also relatively rare. Most of our best fossil plant sites are Eocene, the ancient lakebed sites at McAbee and Princeton — so a good 15 million or so years earlier.

During that expedition, the team recovered a fragment of a large Cretaceous terrestrial trionychoid turtle Basilemys from the family Nanhsiungchelyidae near the confluence of Birdflat Creek and the Sustut River. This largely North American turtle along with the plants will allow us to make correlations with terrestrial finds from other sites including those from the Nanaimo group, the inland island construction sites and the Trent River on Vancouver Island and Horseshoe Canyon in southwestern Alberta. Jordan Mallon and Donald Brinkman have done some good work on the Basilemys morrinensis from the Upper Cretaceous Horseshoe Canyon Formation. The Sustut Basin turtle and plant remains have been accessioned into the Royal BC Museum’s collections in Victoria.

It wasn't until last summer that Arbour was able to extract more of this dinosaur and not all of it as their field season was shortened by a cold snap that brought snow and ice, freezing the ground they were working in the high alpine. Arbour plans to continue her work searching for dinosaur fossils in the high alpine plateaus of northern British Columbia. A fresh grant this year from the Natural Sciences and Engineering Research Council of Canada (NSERC) will help pave the way for both her and some summer students to continue their fieldwork.

Reference: Arbour VM, Evans DC. 2019. A new leptoceratopsid dinosaur from Maastrichtian-aged deposits of the Sustut Basin, northern British Columbia, Canada. PeerJ 7:e7926 https://doi.org/10.7717/peerj.7926. Here's a link to the paper: https://peerj.com/articles/7926/

Figure 1: RBCM P900, the holotype of Ferrisaurus sustutensis, was collected along the BC Rail line near the intersection of Birdflat Creek and the Sustut River in 1971, in the Sustut Basin of northern British Columbia, Canada. Map modified from Evenchick et al. (2003).

Figure 2: Preserved elements of RBCM P900, holotype of Ferrisaurus sustutensis, in white (gray represents missing parts of incomplete bones). RBCM P900 includes a partial right coracoid, partial left scapular blade, complete left radius, partial left ulna, partial left tibia, fibula, and coossified astragalus and ?calcaneum, partial left metatarsals I-IV, and digits III (phalanges 2–4) and IV (phalanges 2–5) of the right pes.

Figure 3: Pectoral elements of RBCM P900, holotype of Ferrisaurus sustutensis, compared to other Laramidian leptoceratopsids. (A) Fragmentary right coracoid of RBCM P900 in lateral view, compared to (B) complete right scapulocoracoid of CMN 8889, Leptoceratops gracilis, lateral view centered on coracoid with scapula in oblique view. Fragmentary left scapular blade of RBCM P900 in (C) lateral and (D) medial view, compared to (E) left scapula of MOR 300, Cerasinops hodgskissi in medial view, and (F) left scapula of TCM 2003.1.9, Prenoceratops pieganensis in lateral view. Abbreviations: sp, sternal process.

HADROSAUR TOOTH FROM ALBERTA

A rare and very beautifully preserved Cretaceous Hadrosaur Tooth. This lovely specimen is from one of our beloved herbivorous "Duck-Billed" dinosaurs from 68 million-year-old outcrops near Drumheller, Alberta, Canada, and is likely from an Edmontosaurus.

When you scour the badlands of southern Alberta, most of the dinosaur material you'll find are from hadrosaurs. These lovely tree-less valleys make for excellent-searching grounds and have led us to know more about hadrosaur anatomy, evolution, and paleobiology than for most other dinosaurs.

We have oodles of very tasty specimens and data to work with. We've got great skin impressions and scale patterns from at least ten species and interesting pathological specimens that provide valuable insights into hadrosaur behaviour. Locally, we have an excellent specimen you can visit in the Courtenay and District Museum on Vancouver Island, Canada. The first hadrosaur bones were found on Vancouver Island a few years back by Mike Trask, VIPS, on the Trent River near Courtenay.

The Courtenay hadrosaur is a first in British Columbia, but our sister province of Alberta has them en masse. Given the ideal collecting grounds, many of the papers on hadrosaurs focus on our Canadian finds. These herbivorous beauties are also found in Europe, South America, Mexico, Mongolia, China, and Russia. Hadrosaurs had teeth arranged in stacks designed for grinding and crushing, similar to how you might picture a cow munching away on the grass in a field. These complex rows of "dental batteries" contained up to 300 individual teeth in each jaw ramus. But even with this great number, we rarely see them as individual specimens.

They didn't appear to shed them all that often. Older teeth that are normally shed in our general understanding of vertebrate dentition, were resorped, meaning that their wee osteoclasts broke down the tooth tissue and reabsorbed the yummy minerals and calcium.

As the deeply awesome Mike Boyd notes, "this is an especially lucky find as hadrosaurs did not normally shed so much as a tooth, except as the result of an accident when feeding or after death. Typically, these fascinating dinosaurs ground away their teeth... almost to nothing."

In hadrosaurs, the root of the tooth formed part of the grinding surface as opposed to a crown covering over the core of the tooth. And curiously, they developed this dental arrangement from their embryonic state, through to hatchling then full adult.

There's some great research being done by Aaron LeBlanc, Robert R. Reisz, David C. Evans and Alida M. Bailleul. They published in BMC Evolutionary Biology on work that looks at the histology of hadrosaurid teeth analyzing them through cross-sections. Jon Tennant did a nice summary of their research. I've included both a link to the original journal article and Jon Tennant's blog below.

LeBlanc et al. are one of the first teams to look at the development of the tissues making up hadrosaur teeth, analyzing the tissue and growth series (like rings of a tree) to see just how these complex tooth batteries formed.

They undertook the first comprehensive, tissue-level study of dental ontogeny in hadrosaurids using several intact maxillary and dentary batteries and compared them to sections of other archosaurs and mammals. They used these comparisons to pinpoint shifts in the ancestral reptilian pattern of tooth ontogeny that allowed hadrosaurids to form complex dental batteries.

References:

LeBlanc et al. (2016) Ontogeny reveals function and evolution of the hadrosaurid dinosaur dental battery, BMC Evolutionary Biology. 16:152, DOI 10.1186/s12862-016-0721-1 (OA link)

To read more from Jon Tennant, visit: https://blogs.plos.org/paleocomm/2016/09/14/all-the-better-to-chew-you-with-my-dear/

Photo credit: Derrick Kersey. For more awesome fossil photos like this from Derrick, visit his page: https://www.facebook.com/prehistoricexpedition/

PLAZA DE ESPANA, SEVILLE

The Plaza de España is a plaza in the Parque de María Luisa, in Seville, Spain. It was built in 1928 for the Ibero-American Exposition of 1929. 

It is a landmark example of Regionalism Architecture, mixing elements of the Baroque Revival, Renaissance Revival and Moorish Revival styles of Spanish architecture. You can stroll through the grounds and explore each of the buildings. There is amazing tile work.

The Plaza de España, designed by Aníbal González, was a principal building built on the Maria Luisa Park's edge to showcase Spain's industry and technology exhibits. González combined a mix of 1920s Art Deco and Spanish Renaissance Revival, Spanish Baroque Revival and Neo-Mudéjar styles. The Plaza de España complex is a huge half-circle; the buildings are accessible by four bridges over the moat, which represent the ancient kingdoms of Spain. In the centre is the Vicente Traver fountain.

Many tiled alcoves were built around the plaza, each representing a different province of Spain. Each alcove is flanked by a pair of covered bookshelves, now used by visitors in the manner of a "Little Free Library". Each bookshelf often contains works with information about each province. Visitors have also donated favourite novels and other books for others to read.

Today the buildings of the Plaza de España have been renovated and adapted for use as offices for government agencies. The central government departments, with a sensitive adaptive redesign, are located within it. Toward the end of the park, the grandest mansions from the fair have been adapted as museums. The most distant museum contains the city's archaeology collections. The main exhibits are Roman mosaics and artefacts from nearby Italica.

The Plaza de España has been used as a filming location, including scenes for Lawrence of Arabia (1962). The building was used as a location in the Star Wars movie series Star Wars: Episode II – Attack of the Clones (2002) — in which it featured in shots of the City of Theed on the Planet Naboo. It also featured in the 2012 film The Dictator.

CERECINOS DE CAMPOS: DEINOTHERIUM

This partial specimen of Deinotherium giganteum hails from Middle-Upper Miocene, c. 15.97-5.33 Million Years outcrops near Cerecinos de Campos, Zamora Castile and León, northwestern Spain.

Deinotherium means "terrible beast," which feels a bit unkind to this vegetarian — though he was one of the largest elephants to walk this Earth. 

They are relatively recent in the evolutionary story of the Earth. They first appeared 17 million years ago, had a short run of it and became extinct relatively recently — just 1.6 million years ago. This fellow's cousin, Deinotherium bozasi would likely have interacted with some of our oldest relatives. 

One of the distinguishing features of Deinotherium is their curved tusks inserted only in the jaw. One of the tusks from this fellow, on display at the Museo Nacional De Ciencias Naturales in Madrid, Spain, while incomplete, was preserved rather nicely and shows the detail of where the tusk meets the jaw. Deinotherium could reach a height of over 3.5 meters. Its structure and size are similar to those of the present-day elephant. 

EL TORCAL DE ANTEQUERA

El Torcal de Antequera

El Torcal de Antequera is a nature reserve in the Sierra del Torcal mountain range south of the city of Antequera, in Andalusia, Spain. 

From the tops of the hillsides, you can see far into the fertile grazing lands of the province of Málaga. 

There are numerous hiking routes throughout the park, some for serious walkers and climbers, as well as for those who might prefer a more gentle meander. 

El Torcal is known for its unusual landforms and is regarded as one of the most impressive karst landscapes in Europe. Karst topography forms from the dissolution of soluble rocks like limestone, dolomite, and gypsum. It often has underground drainage systems with sinkholes and caves. 

Water loves to dissolve the softer rocks but it works its erosional magic on harder, more weathering-resistant quartzites given the right conditions. El Torcal has many wonderful caves and thousands of chasms for the small animals living in this area to call home. Some are quite small, while others are large enough to be explored. The rock we see at El Torcal formed over several hundred million years. 

About 200 million years ago, much of Europe and the Middle East were submerged under the Tethys Sea. 

This was a time of carbonate sedimentation as the skeletons, shells and shells of small marine animals lived and died, depositing their remains at the bottom of the sea. 

Over vast amounts of time, these wee bits of marine matter built up until 175 million years later, the sediments have built up and compacted to form strat thousands of metres deep. 

Towards the Middle Miocene, the Iberian plates to the north of the Tethys Sea and the African plates to the south, compressed, deformed and fractured those sediments. This process is slow and continuous and still continues today. Water, wind and ice continue to shape the landscape and present the continually eroding karst landscape you can hike through today at El Torcal de Antequera.

El Torcal Natural Park is a UNESCO site. Hiking through the hills, you can see the large mushroom-shaped folds, with a very wide upper part and horizontal layers, and short and abrupt flanks. Karst acts as a large sponge, storing rainwater and releasing it within the rock to encourage the limestone to dissolve. 

Gravity pulls the water down and it trickles out again as streams along the edge of the cliffs. One of the sites that the water gathers is in the Nacimiento de La Villa spring on El Torcal's north side.

El Torcal, Karst Topography

Along with its distinct hoodoos, sprinkled amongst the limestones, you will find a wealth of interesting plants and wildlife. Look for lilies, red peonies, wild rose trees and thirty varieties of orchid.  

The many species of reptiles include the Montpellier snake and ocellated lizard, both endemic to El Torcal. 

Other wildlife to look for are the resident Griffon vultures and Spanish Ibex, Andalusian mountain goats, voles, fox and rabbits. If you are here in the evening, look for some of the nocturnal mammals who call these hills home — badgers and weasels.

The park has an excellent Visitor Centre which makes a natural starting point for your exploration of the reserve. There you will find details about the park, parking and walking routes. Guided walks are available, including the popular ‘Route of the 5 Senses’, a night-time ‘El Torcal Under Moonlight’ walk and a fossil-hunting walk, Route of the Ammonites. The visitor centre includes a very reasonably priced restaurant which offers a good selection of traditional food, all made with locally sourced ingredients.

For those who might enjoy some sightseeing in the heavens, this area of Spain has extremely favourable conditions for stargazing and astronomy. The Astronomical Observation of El Torcal (OAT) is located within the park. They host regular observation evenings that take advantage of the lack of light pollution in this region.  

Places to Stay: Finca Gran Cerros Rural Retreat: The epitome of tranquil, rural Spain, Finca Gran Cerros nestles into the Andalusian hillside just a few minutes drive from the traditional white villages’ of Álora and Valle de Abdalajis. Visit them: https://www.fincagrancerros.com. Fina Gran Cerros is about 30 km south of El Torcal de Antequera nature reserve in the Sierra del Torcal mountains.

TORVOSAURUS: SAVAGE LIZARD

This toothy fellow is Torvosaurus tanneri and he hails from Late Jurassic outcrops in the Carnegie Quarry at Dinosaur National Monument, Morrison Formation, western United States — where we have found a single bone, his humerus telling us about his mighty size. 

The specimen you see here is currently on display at the Museo Nacional De Ciencias Naturales in Madrid, Spain.

Torvosaurus were one of the largest and most robust carnivores of the Jurassic. 

These "savage lizards," were true to their name. They were skilled bipedal hunters who weighed over two tons. They had powerful dentition, large, sharp teeth and strong claws on their forelegs — ferocious predators of the Upper Jurassic. He would have roamed alongside the mighty Camarasaurus, Diplodocus, Apatosaurus, Stegosaurus and Allosaurus.

Palaeontologist Earl Douglass, 1909

Fossil specimens of Torvosaurus have been found in the Lourinha Formation near Lisbon, Portugal. Here, he would have towered over the smaller Allosaurus of the region who were just over eight metres or 27 feet on average, while he towered at over ten metres or 35 feet. 

This was not the case for the Allosaurus — famed brontosaur hunters — who roamed the fern-covered floodplains of the Jurassic west and what would one day become the United States. Here they grew massive, passing twelve metres or 40 feet in length and towering over the local Tovosaurus. Allosaurus had a large bite, their jaws opening up very wide, making them capable of taking very big bites and positioning them as the top carnivores of the Late Jurassic.

Still, both of these hunters had to contend with Sauophaganax, the largest Jurassic theropod at a whopping twelve to thirteen metres — making it the largest Allosaurus and maybe even a wee bit larger than the mighty Tyrannosaurus rex roaming around western North America back when it was the island continent of Laramidia. This would have been fearsome land to roam as the juvenile of any species as all of these brutes would have the skill, speed and teeth to take you down. 

Photo One: Tovosaurus tannerion display at the Museo Nacional De Ciencias Naturales in Madrid, Spain.

Photo Two: Palaeontologist Earl Douglass digging up the remains of a Brontosaurus at the Carnegie Quarry, 1909. To learn more about this fossil site, visit: https://carnegiemnh.org/celebrated-fossil-quarry/

A PASSION FOR PALAEONTOLOGY

An old friend connected via social media to ask how he can best support his seven-year-old daughter's love of palaeontology. That is a question I love to hear! Now, my personal response is a bit of a tidal wave — buy her books, and rocks and a rock tumbler... take her out on fossil field trips, bring her to museums — fuel the flames of that passion for palaeo. Take no prisoners. Get her good and hooked! 

A love of palaeontology spills over to other areas of science and will help spark an interest in biology, ecology and natural history. In a perfect storm, the whole family catches the bug and summer field trips turn to trips from March to October or as soon as the snow clears.

If you are looking to purchase some fossils — be mindful not to purchase Canadian specimens — then Etsy is a good general source. Since we are living in the new normal of Covid, I would also turn to Amazon as a book source and take a boo at their starter rock collections and rock tumblers.

Local museums are a wonderful source of inspiration and tend to favour local fossil specimens. I particularly like the Royal Tyrrell Museum in Drumheller, Alberta and the Courtenay and District Museum on Vancouver Island, British Columbia. Seeing these is useful as it gives you the visual aid you'll need when collecting out in the field.  

Eyewitness Books: Fossils

If you are looking for resources and are readying this from a laptop, you'll see a column down the right-hand side of the page with a whole host of yummy options from books to gear. It is targeted at a slightly older audience, but I'll add some titles that might appeal to a younger audience. 

Ashley Hall did up quite a good children's book targeted for those aged 6-8 years old: Fossils for Kids: A Junior Scientist's Guide to Dinosaur Bones, Ancient Animals, and Prehistoric Life on Earth. It is available on Amazon and includes some wonderful images and covers all the introductory topics one would want to see in a first book on fossils. She also has a nice homage to her parents who inspired and encouraged her love of palaeontology. Dean Lomax and Darren Naish have published some worthy books that make a great addition to the family library.

Eye Witness has produced some wonderfully visual books on fossils. They are general, but that is the perfect place to start. Some folk love dinosaurs, others are into shark's teeth. Myself, I love all the wee invertebrates.  I love a good visual with a bite-size bit of information so you can digest it easily. I have sliced more than one Eyewitness book apart to laminate a section for use in kid's palaeontology courses. 

Some of these topics were touched upon in Season One of the Fossil Huntress Podcast. There is a wee cast on the legal side of palaeo that is worth a listen if you are planning to head out collecting or find yourself tempted to purchase Canadian specimens.

One of the best things about palaeontology is that it can be enjoyed at any age and everyone can contribute to science. Young, old, rich, poor, boy, girl, professional or vocational — fossils do not discriminate. You can be in elementary school and find a new dinosaur or marine reptile species. 

Some of the most significant finds in Canada and around the world are credited to youngsters — from the likes of Mary Anning to British Columbia's first marine reptile and dinosaur finds. The first elasmosaur in British Columbia was found by a young girl and her father. The dinosaurs up near Tumbler Ridge were found by two boys tubing along a river. 

Families and friends out for a stroll have found fossil bits and bones from many new species. The pterosaur Vectidraco was found by a four-year-old, who was honoured through the species name V. daisymorrisae. The Late Jurassic herbivorous dinosaur, Chilesaurus diegosuarezi, from Chile, was discovered by a seven-year-old while his parents briefly distracted — a lucky bit of timing for us all.  

So, if you're reading this, JD, I'm thrilled for you! Fuel the flames. Encourage her love of fossils, science and the natural world.

THERIZINOSAURUS: DINOSAUR EGGS

The brood of eggs you see here belong to the slow-moving but massive dinosaur Therizinosaurus. He belonged to a genus of sizable therizinosaurid that lived during the Late Cretaceous, 70 million years ago. 

Therizinosaurus was a colossal therizinosaur that could grow up to 9–10 m (30–33 ft) long and weigh possibly over 3 t (3,000 kg). Like other therizinosaurs, it would have been a bit of a slowpoke on the ground. These fellows had a rhamphotheca (horny beak) and a wide torso for food processing. 

The forelimbs were particularly robust and had three fingers that bore unguals which, unlike other relatives, were very stiffened, elongated, and only had significant curvatures at the tips. After years of taxonomic debate, nevertheless, they are now placed in one of the major dinosaur clades, Theropoda, specifically as maniraptorans. 

DINOSAUR GEORGE / EPISODE #143

 

PALAEONTOLOGIST EARL DOUGLASS: THE CARNEGIE QUARRY

Palaeontologist Earl Douglass, 1909

About 150 million years ago, a severe drought ravaged the western interior of North America. In eastern Utah, malnourished dinosaurs gathered near a dwindling river to live out their last days. 

Today, this site is known as the Carnegie Quarry at Dinosaur National Monument, and it is one of the most incredible fossil sites in the world.

The celebrated fossil quarry at what is now recognized as Dinosaur National Monument in Utah was discovered in 1909 by Carnegie Museum field collector Earl Douglass.

“I saw eight of the tail bones of a Brontosaurus in exact position. It was a beautiful sight.”  — Earl Douglass in his diary on August 17, 1909, recounting the moment he found the first dinosaur remains of a Brontosaurus at the Carnegie Quarry. Those vertebrae were part of a fully articulated skeleton that became the type for a new species, Apatosaurus louisae, (Gilmore, 1936), published a detailed quarry map showing the skeleton, with "outcrop" identifying the discovery bones. The specimen is now mounted in the Carnegie Museum and those eight tail bones, freed from their sandstone tomb. 

From 1909–1923, Douglass and his crews collected more than 350 tons (700,000 pounds) of fossils from that site alone. Several dinosaur skeletons discovered by Douglass at this quarry are featured in our core exhibition hall, Dinosaurs in Their Time.

Others grace the exhibit halls of other prominent North American museums, such as the American Museum of Natural History in New York, the Smithsonian Institution’s National Museum of Natural History in Washington, DC, the Denver Museum of Nature and Science, and the Royal Ontario Museum in Toronto.

If you would like to visit Dinosaur National Monument, you can explore extensive outcrops of the Morrison at the Dinosaur Quarry, on the Fossil Discovery Trail, the Sounds of Silence Trail, and other areas in the park.

To learn more about this fossil site, visit: https://carnegiemnh.org/celebrated-fossil-quarry/

CRETACEOUS HADROSAUR TOOTH

A rare and very beautifully preserved Cretaceous Hadrosaur Tooth. This lovely specimen is from one of our beloved herbivorous "Duck-Billed" dinosaurs from 68 million-year-old outcrops near Drumheller, Alberta, Canada, and is likely from an Edmontosaurus.

When you scour the badlands of southern Alberta, most of the dinosaur material you'll find are from hadrosaurs. These lovely tree-less valleys make for excellent-searching grounds and have led us to know more about hadrosaur anatomy, evolution, and paleobiology than for most other dinosaurs.

We have oodles of very tasty specimens and data to work with. We've got great skin impressions and scale patterns from at least ten species and interesting pathological specimens that provide valuable insights into hadrosaur behaviour. Locally, we have an excellent specimen you can visit in the Courtenay and District Museum on Vancouver Island, Canada. The first hadrosaur bones were found on Vancouver Island a few years back by Mike Trask, VIPS, on the Trent River near Courtenay.

The Courtenay hadrosaur is a first in British Columbia, but our sister province of Alberta has them en masse. Given the ideal collecting grounds, many of the papers on hadrosaurs focus on our Canadian finds. These herbivorous beauties are also found in Europe, South America, Mexico, Mongolia, China, and Russia. Hadrosaurs had teeth arranged in stacks designed for grinding and crushing, similar to how you might picture a cow munching away on the grass in a field. These complex rows of "dental batteries" contained up to 300 individual teeth in each jaw ramus. But even with this great number, we rarely see them as individual specimens.

They didn't appear to shed them all that often. Older teeth that are normally shed in our general understanding of vertebrate dentition, were resorped, meaning that their wee osteoclasts broke down the tooth tissue and reabsorbed the yummy minerals and calcium.

As the deeply awesome Mike Boyd notes, "this is an especially lucky find as hadrosaurs did not normally shed so much as a tooth, except as the result of an accident when feeding or after death. Typically, these fascinating dinosaurs ground away their teeth... almost to nothing."

In hadrosaurs, the root of the tooth formed part of the grinding surface as opposed to a crown covering over the core of the tooth. And curiously, they developed this dental arrangement from their embryonic state, through to hatchling then full adult.

There's some great research being done by Aaron LeBlanc, Robert R. Reisz, David C. Evans and Alida M. Bailleul. They published in BMC Evolutionary Biology on work that looks at the histology of hadrosaurid teeth analyzing them through cross-sections. Jon Tennant did a nice summary of their research. I've included both a link to the original journal article and Jon Tennant's blog below.

LeBlanc et al. are one of the first teams to look at the development of the tissues making up hadrosaur teeth, analyzing the tissue and growth series (like rings of a tree) to see just how these complex tooth batteries formed.

They undertook the first comprehensive, tissue-level study of dental ontogeny in hadrosaurids using several intact maxillary and dentary batteries and compared them to sections of other archosaurs and mammals. They used these comparisons to pinpoint shifts in the ancestral reptilian pattern of tooth ontogeny that allowed hadrosaurids to form complex dental batteries.

References:

LeBlanc et al. (2016) Ontogeny reveals function and evolution of the hadrosaurid dinosaur dental battery, BMC Evolutionary Biology. 16:152, DOI 10.1186/s12862-016-0721-1 (OA link)

To read more from Jon Tennant, visit: https://blogs.plos.org/paleocomm/2016/09/14/all-the-better-to-chew-you-with-my-dear/

Photo credit: Derrick Kersey. For more awesome fossil photos like this from Derrick, visit his page: https://www.facebook.com/prehistoricexpedition/

PERMIAN-TRIASSIC MASS EXTINCTION: EVOLUTIONARY ARMS RACE

Yesterday, on the Fossil Huntress Podcast, we wrestled with the question of whether dinosaurs were warm-blooded or cold-blooded. It is an excellent question and there is good evidence on both sides of that debate.

Many dinosaurs stood upright — a warm-blooded trait. They are also the ancestors of birds who are warm-blooded. Dinosaurs often began life with porous bones, moving to denser bones later in life. This is as much a mark of growth rate as it is for the warm-cold debate. 

And, dinosaurs had small brains relative to body size — a trait of our cold-blooded animals. So, which is it? Cold or warm? My money is on the latter, but we'll likely have some time to wait before we have enough evidence to say for sure one way or the other. One thing we do know to be true is that we see a trend of the Earth's animals moving from cold-bloodedness to warm-bloodedness over time. 

What was the driver for that adaptation? One of the drivers looks to be the Permian-Triassic mass extinction event some 250 million years ago. It was a catastrophic event that killed ninety-five percent of all life on Earth. The remaining species were left to fight for survival against an inhospitable planet and one another. The few surviving species found themselves in a turbulent world —repeatedly hit by ice ages, rapid warming and ocean acidification cycles.

Through all of that, two main groups of tetrapods survived; the synapsids and archosaurs, ancestors of mammals and birds. The ancestors of both mammals and birds became warm-blooded at the same time.

Warm-bloodedness, or endothermy, is the ability to regulate your body temperature using your metabolism rather than relying on the external environment. Humans are endothermic. We eat food and wear warm sweaters to guard against the cold. Warm-bloodedness is key for both survival and reproductive fitness.

There is evidence of warm-bloodedness, including a diaphragm and whiskers in the synapsids as far back as the Triassic. This is supported by a more porous bone structure in both synapsids and archosaurs. Warm-blooded animals tend to have highly vascularized bone tissue. Cold-blooded animals have a denser bone structure that even exhibits annual growth rings. 

Dinosaurs show both traits. They start off life with highly vascularized bone which becomes denser as they mature. This move from vascular to dense bone may have more to do with growth rates than to whether the animals were warm or cold-blooded. 

Another factor in warmth is hair. We know that mammal ancestors had hair from the beginning of the Triassic. More recently, we have learned that archosaurs had feathers from 250 million years ago. Archosaurs are a group of diapsids and are broadly classified as reptiles. The living representatives of this group are birds and crocodilians. It also includes all extinct dinosaurs, pterosaurs, and extinct close relatives of crocodilians. 

Medium-sized and large tetrapods switched from sprawling to erect posture right at the Permian-Triassic boundary. As you know, most warm-blooded animals have an erect or upright posture and our cold-blooded friends tend to walk on all fours. 

The mass posture change and early origin of hair and feathers all speak to the beginning of a species arms race. In ecological terms, an arms race occurs when predators and prey compete on an escalated scale for survival. This pressure caused a rapid change in their evolution as their adaptations escalate. 

When we look at our world today, warm-blooded animals populate all areas of the Earth. They have fewer offspring and show intense parental care, taking months or years to care for their young before they become independent. These adaptations give birds and mammals an edge over amphibians and reptiles and we see this in their domination of the ecosystems in our world.

This revolution in ecosystems was triggered by the independent origins of endothermy in birds and mammals. This particular adaptation lives on as these species survive and thrive in an Earth that can be fickle in terms of environmental conditions.

Reference: Benton, Michael J. The origin of endothermy in synapsids and archosaurs and arms races in 
the Triassic, Gondwana Research, School of Earth Sciences, Life Sciences Building, University of Bristol, Bristol BS8 1TH, UKThe evolution of main groups through the Triassic. Image: Nobu Tamura

FIRST BC DINOSAUR WEST OF THE ROCKIES

This dapper fellow is a pine needle and horsetail connoisseur. He's a hadrosaurus — also known as "duck-billed" dinosaurs. They were a very successful group of plant-eaters that thrived throughout western Canada during the late Cretaceous, some 70 to 84 million years ago.

This beautiful specimen graces the back galleries of the Courtenay and District Museum on Vancouver Island, British Columbia, Canada. I was very fortunate to have a tour this past summer with the deeply awesome Mike Trask joined by the lovely Lori Vesper. 

The museum houses an extensive collection of palaeontological and archaeological material found on Vancouver Island, many of which have been donated by the Vancouver Island Palaeontological Society.

Hadrosaurs lived as part of a herd, dining on pine needles, horsetails, twigs and flowering plants. They are ornithischians — an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. They are close relatives and possibly descendants of the earlier iguanodontid dinosaurs. They had slightly webbed, camel-like feet with pads on the bottom for cushioning and perhaps a bit of extra propulsion in water. They were primarily terrestrial but did enjoy feeding on plants near and in shallow water. There had a sturdy build with a stiff tail and robust bone structure. 

At their emergence in the fossil record, they were quite small, roughly three meters long. That's slightly smaller than an American bison. They evolved during the Cretaceous with some of their lineage reaching up to 20 meters or 65 feet.

Hadrosaurs are very rare in British Columbia but a common fossil in our provincial neighbour, Alberta, to the east. Here, along with the rest of the world, they were more abundant than sauropods and a relatively common fossil find. They were common in the Upper Cretaceous of Europe, Asia, and North America.

There are two main groups of Hadrosaurs, crested and non-crested. The bony crest on the top of the head of the hadrosaurs was hollow and attached to the nasal passages. It is thought that the hollow crest was used to make different sounds. These sounds may have signalled distress or been the hadrosaur equivalent of a wolf whistle used to attract mates. Given their size it would have made for quite the trumpeting sound.

Dan Bowen, Chair of the Vancouver Island Palaeontological Society, shared the photo you see here of the first partly articulated dinosaur from Vancouver Island ever found. The vertebrate photo and illustration are from a presentation by Dr. David Evans at the 2018 Paleontological Symposium in Courtenay.  

The research efforts of the VIPS run deep in British Columbia and this new very significant find is no exception. A Hadrosauroid dinosaur is a rare occurrence and further evidence of the terrestrial influence in the Upper Cretaceous, Nanaimo Group, Vancouver Island — outcrops that we traditionally thought of as marine from years of collecting well-preserved marine fossil fauna.

The fossil bone material was found years ago by Mike Trask of the Vancouver Island Palaeontological Society. You may recall that he was the same fellow who found the Courtenay Elasmosaur on the Puntledge River.

Mike was leading a fossil expedition on the Trent River. While searching through the Upper Cretaceous shales, the group found an articulated mass of bones that looked quite promising.

Given the history of the finds in the area, the bones were thought to be from a marine reptile.

Since that time, we've found a wonderful terrestrial helochelydrid turtle, Naomichelys speciosa, but up to this point, the Trent had been known for its fossil marine fauna, not terrestrial. Efforts were made to excavate more of the specimen, and in all more than 25 associated vertebrae were collected with the help of some 40+ volunteers. Identifying fossil bone is a tricky business. Encased in rock, the caudal vertebrae were thought to be marine reptile in origin. Some of these were put on display in the Courtenay Museum and mislabeled for years as an unidentified plesiosaur.

In 2016, after years collecting dust and praise in equal measure, the bones were reexamined. They didn't quite match what we'd expect from a marine reptile. Shino Sugimoto, Fossil Preparator, Vertebrate Palaeontology Technician at the Royal Ontario Museum was called in to work her magic — painstakingly prepping out each caudal vertebrae from the block.

Once fully prepped, seemingly unlikely, they turned out to be from a terrestrial hadrosauroid. This is the second confirmed dinosaur from the Upper Cretaceous Nanaimo Group. The first being a theropod from Sucia Island. The partial left thigh bones the first dinosaur fossil ever found in Washington state.

Dr. David Evans, Temerty Chair in Vertebrate Palaeontology, Department of Natural History, Palaeobiology from the Royal Ontario Museum, confirmed the ID and began working on the partial duck-billed dinosaur skeleton to publish on the find.

Now fully prepped, the details of this articulated Hadrosauriod caudal vertebrae come to light. We can see the prominent chevron facets indicative of caudal vertebrae with it's a nice hexagonal centrum shape on anterior view.

There are well-defined long, raked neural spines that expand distally — up and away from the acoelous centrum. 

Between the successive vertebrae, there would likely have been a fibrocartilaginous intervertebral body with a gel-like core —  the nucleus pulposus — which is derived from the embryonic notochord. This is a handy feature in a vertebrate built as sturdily as a hadrosaur. Acoelous vertebrae have evolved to be especially well-suited to receive and distribute compressive forces within the vertebral column.

This fellow has kissing cousins over in the state of New Jersey where this species is the official state fossil. The first of his kind was found by John Estaugh Hopkins in New Jersey back in 1838. Since that time, we've found many hadrosaurs in Alberta, particularly the Edmontosuaurs, another member of the subfamily Hadrosaurine.

In 1978, Princeton University found fifteen juvenile hadrosaurs, Maiasaura ("good mother lizard") on a paleontological expedition to the Upper Cretaceous, Two Medicine Formation of Teton County in western Montana. 

Their initial finds of several small skeletons had them on the hunt for potential nests — and they found them complete with wee baby hatchlings!

Photo One: Fossil Huntress / Heidi Henderson, VIPS

Photo Two / Sketch Three: Danielle Dufault, Palaeo-Scientific Ilustrator, Research Assistant at the Royal Ontario Museum, Host of Animalogic. 

The vertebrate photo and illustration were included in a presentation by Dr. David Evans at the 2018 BCPA Paleontological Symposium in Courtenay, British Columbia, Canada.

Photo Four: Illustration by the talented Greer Stothers, Illustrator & Natural Science-Enthusiast.