Puffbird similar to Fossil Birds found at Driftwood Canyon |
Metasequoia, the Dawn Redwood |
A Tapir showing off his prehensile nose trunk |
Puffbird similar to Fossil Birds found at Driftwood Canyon |
Metasequoia, the Dawn Redwood |
A Tapir showing off his prehensile nose trunk |
This material is very important as it greatly expands our understanding of the fauna and ranges of ammonites currently included in the North American regional ammonite zonation.
I had the very great honour of having the fellow below, Fergusonites hendersonae, a new species of nektonic carnivorous ammonite, named after me by palaeontologist Louse Longridge from the University of British Columbia.
I'd met Louise as an undergrad and was pleased as punch to hear that she would be continuing the research by Dr. Howard Tipper, the authority on this area of the Chilcotins and Haida Gwaii — which he dearly loved.
"Tip" was a renowned Jurassic ammonite palaeontologist and an excellent regional mapper who mapped large areas of the Cordillera. He made significant contributions to Jurassic paleobiogeography and taxonomy in collaboration with Dr. Paul Smith, Head of Earth and Ocean Science at the University of British Columbia.
Tip’s regional mapping within BC has withstood the test of time and for many areas became the regions' base maps for future studies. The scope of Tip’s understanding of Cordilleran geology and Jurassic palaeontology will likely never be matched. He passed away on April 21, 2005. His humour, knowledge and leadership will be sorely missed.
Fergusonites hendersonae |
Both Dan Bowen and John Fam were instrumental in planning those expeditions and each of them benefited greatly from the knowledge of Dr. Howard Tipper.
If not for Tipper's early work in the region, our shared understanding and much of what was accomplished in his last years and after his passing would not have been possible.
Over the course of three field seasons, we endured elevation sickness, rain, snow, grizzly bears and very chilly nights — we were sleeping right next to a glacier at one point — but were rewarded by the enthusiastic crew, helicopter rides — which really cut down the hiking time — excellent specimens including three new species of ammonites, along with a high-spired gastropod and lobster claw that have yet to be written up. This area of the world is wonderful to hike and explore — stunningly beautiful country. We were also blessed with access as the area is closed to all fossil collecting except with a permit.
This fauna understanding helps us to understand the correlations between different areas: (1) the Mineralense and Rursicostatum zones are present in Taseko Lakes and can be readily correlated with contemporaneous strata elsewhere in North America; (2) the Mineralense and Rursicostatum zones of North America are broadly equivalent to the Canadensis Zone and probably the Arcuatum horizon of the South American succession; (3) broad correlations are possible with middle–late Hettangian and earliest Sinemurian taxa in New Zealand; (4) the Mineralense and Rursicostatum zones are broadly equivalent to the circum‐Mediterranean Marmoreum Zone; (5) the Mineralense Zone and the lower to middle portion of the Rursicostatum Zone are probably equivalent to the Complanata Subzone whereas the upper portion of the Rursicostatum Zone may equate to the Depressa Subzone of the north‐west European succession.
Taseko Lake Area, BC |
Since then, knowledge of eastern Pacific Hettangian ammonite faunas has improved considerably.
Detailed systematic studies have been completed on faunas from localities in other areas of BC, Alberta, Alaska, Oregon, Nevada, Mexico and South America (e.g. Guex 1980, 1995; Imlay 1981; Hillebrandt 1981, 1988, 1990, 1994, 2000a–d; Smith and Tipper 1986; Riccardi et al. 1991; Jakobs and Pálfy 1994; Pálfy et al. 1994, 1999; Taylor 1998; Hall et al. 2000; Taylor and Guex 2002; Hall and Pitaru 2004).
These studies have demonstrated that Early Jurassic eastern Pacific ammonites had strong Tethyan affinities as well as a high degree of endemism (Guex 1980, 1995; Taylor et al. 1984; Smith et al. 1988; Jakobs et al. 1994; Pálfy et al. 1994). Frebold’s early studies were also hampered because they were based on small collections, which limited understanding of the diversity of the fauna and variation within populations. However, recent mapping has greatly improved our understanding of the geology of Taseko Lakes (Schiarizza et al. 1997; Smith et al. 1998; Umhoefer and Tipper 1998) and encouraged further collecting that has dramatically increased the size of the sample.
A study of the ammonite fauna from Taseko Lakes is of interest for several reasons. The data are important for increasing the precision of the late Hettangian portion of the North American Zonation.
Owing to the principally Tethyan or endemic nature of Early Jurassic ammonites in the eastern Pacific, a separate zonation for the Hettangian and Sinemurian of the Western Cordillera of North America has been established by Taylor et al. (2001). Except for information available from the early studies by Frebold (1951, 1967), the only Taseko Lakes taxa included in the North American Zonation of Taylor et al. (2001) were species of Angulaticeras studied by Smith and Tipper (2000).
Since then, Longridge et al. (2006) made significant changes to the zonation of the late Hettangian and early Sinemurian based on a detailed study of the Badouxia fauna from Taseko Lakes (Text‐fig. 2). An additional taxonomic study was recently completed on the late Hettangian ammonite Sunrisites (Longridge et al. 2008) and this information has not yet been included within the zonation.
Hettangian Zonation |
The Taseko Lakes fauna can improve Hettangian correlations within North America as well as between North America and the rest of the world.
North‐west European ammonite successions (e.g. Dean et al. 1961; Mouterde and Corna 1997; Page 2003) are considered the primary standard for Early Jurassic biochronology (Callomon 1984).
In north‐west Europe, the turnover from schlotheimiid dominated faunas in the late Hettangian to arietitid dominated faunas in the early Sinemurian was sharp (e.g. Dean et al. 1961; Bloos 1994; Bloos and Page 2002). In other areas, by contrast, these faunas were not so mutually exclusive and the transition was much more gradual.
This makes correlations between north‐west Europe and other areas difficult (e.g. Bloos 1994; Bloos and Page 2000, 2002). Correlations are further impeded by endemism and provincialism.
The Taseko Lakes fauna addresses these problems because it contains many taxa that are common throughout the eastern Pacific as well as several cosmopolitan taxa that make intercontinental correlation possible. Correlation between North America and other areas is of particular significance in that the interbedded volcanic and fossiliferous marine rocks in North America permit the calibration of geochronological and biochronological time scales (Pálfy et al. 1999, 2000).
This correlation between the late Hettangian fauna in the Taseko Lakes area and contemporaneous faunas in other areas of North America, South America, New Zealand, western and eastern Tethys, and north‐west Europe is of particular interest to me — especially the correlation of the faunal sequences of Nevada, USA.
Reference: PaleoDB 157367 M. Clapham GSC C-208992, Section A 09, Castle Pass Angulata - Jurassic 1 - Canada, Longridge et al. (2008)
L. M. Longridge, P. L. Smith, and H. W. Tipper. 2008. Late Hettangian (Early Jurassic) ammonites from Taseko Lakes, British Columbia, Canada. Palaeontology 51:367-404
PaleoDB taxon number: 297415; Cephalopoda - Ammonoidea - Juraphyllitidae; Fergusonites hendersonae Longridge et al. 2008 (ammonite); Average measurements (in mm): shell width 9.88, shell diameter 28.2; Age range: 201.6 to 196.5 Ma. Locality info: British Columbia, Canada (51.1° N, 123.0° W: paleo coordinates 22.1° N, 66.1° W)
Hallstatt Salt Mines, Austria / Permian Salt Diapir |
Hallstatt and the Hallstatt Sea, Austria |
Hallstatt Beinhaus Ossuary, Hallstatt, Austria |
Hallstatt, Salzkammergut region, Austria |
Bronze Age vessel with cow and calf |
Anahoplites is now included in the subfamily Anahoplitinae and separated from the Hoplitinae where it was placed in the older in the 1957 edition of the Treatise on Invertebrate Paleontology, Part L (Ammonoidea). Genera of the Hoplitinae tend to be more robust, with broader whorls and stronger ribs.
Anahoplites is found in Cretaceous (Middle to the Late Albian) deposits from England, through Europe, all the way to the Transcaspian Oblast region in Russia to the east of the Caspian Sea. The Aube department, named after the local river, is the type locality of the Albian stage (d'ORBIGNY, 1842).
A. planus from the French Coast |
This involute (113 mm) specimen shows evidence of cohabitation by some of his marine peers.
We see two different bryozoa, an oyster and some serpulids making a living and leaving trace fossils on her flat sides. The top specimen was prepared with potase by José Juárez Ruiz of Spain.
The lovely Anahoplites planus you see here to the lower right was found by Bertus op den Dries on the French coast in Albian deposits near Wissant, P5 and measures in at 8 cm. This on edge view gives you a very good sense of the keel.
Dan Bowen, Chair, VIPS, Trent River |
The Pacific Plate is an oceanic tectonic plate that lies beneath the Pacific Ocean. And it is massive. At 103 million km2 (40 million sq mi), it is the largest tectonic plate and continues to grow fed by volcanic eruptions that piggyback onto its trailing edge.
This relentless expansion pushes the Pacific Plate into the North American Plate. The pressure subducts it beneath our continent where it then melts back into the earth. Plate tectonics are slow but powerful forces.
The island chains that rode the plates across the Pacific smashed into our coastline and slowly built the province of British Columbia. And because each of those islands had a different origin, they create pockets of interesting and diverse geology.
It is these islands that make up the Insular Belt — a physio-geological region on the northwestern North American coast. It consists of three major island groups — and many smaller islands — that stretches from southern British Columbia up into Alaska and the Yukon. These bits of islands on the move arrived from the Late Cretaceous through the Eocene — and continues to this day.
The rocks that form the Insular Superterrane are allochthonous, meaning they are not related to the rest of the North American continent. The rocks we walk over along the Trent River are distinct from those we find throughout the rest of Vancouver Island, Haida Gwaii, the rest of the province of British Columbia and completely foreign to those we find next door in Alberta.
To discover what we do find on the Trent takes only a wee stroll, a bit of digging and time to put all the pieces of the puzzle together. The first geological forays to Vancouver Island were to look for coal deposits, the profitable remains of ancient forests that could be burned to the power industry.
Jim Monger and Charlie Ross of the Geological Survey of Canada both worked to further our knowledge of the complex geology of the Comox Basin. They were at the cutting edge of west coast geology in the 1970s. It was their work that helped tease out how and where the rocks we see along the Trent today were formed and made their way north.We know from their work that by 85 million years ago, the Insular Superterrane had made its way to what is now British Columbia.
The lands were forested much as they are now but by extinct genera and families. The fossil remains of trees similar to oak, poplar, maple and ash can be found along the Trent and Vancouver Island. We also see the lovely remains of flowering plants such as Cupanities crenularis, figs and breadfruit.
Heading up the river, you come to a delineation zone that clearly marks the contact between the dark grey marine shales and mudstones of the Haslam Formation where they meet the sandstones of the Comox Formation. Fossilized material is less abundant in the Comox sandstones but still contains some interesting specimens. Here you begin to see fossilized wood and identifiable fossil plant material.
Further upstream, there is a small tributary, Idle Creek, where you can find more of this terrestrial material in the sandy shales. As you walk up, you see identifiable fossil plants beneath your feet and jungle-like, overgrown moss-covered, snarly trees all around you.
Walking west from the Trent River Falls at the bottom, you pass the infamous Ammonite Alley, where you can find Mesopuzosia sp. and Kitchinites sp. of the Upper Cretaceous (Santonian), Haslam Formation. Minding the slippery green algae covering some of the river rocks, you can see the first of the Polytychoceras vancouverense zone.
Continuing west, you reach the first of two fossil turtle sites on the river — amazingly, one terrestrial and one marine. If you continue, you come to the Inland Island Highway.
The Trent River has yielded some very interesting marine specimens, and significant terrestrial finds. We have found a wonderful terrestrial helochelydrid turtle, Naomichelys speciosa, and the caudal vertebrae of a Hadrosauroid dinosaur. Walking down from the Hadrosaur site you come to the site of the fossil ratfish find — one of the ocean's oddest fish.
Ratfish, Hydrolagus Collie, are chimaera found in the north-eastern Pacific Ocean today. The fossil specimen from the Trent would be considered large by modern standards as it is a bruiser in comparison to his modern counterparts.This robust fellow had exceptionally large eyes and sex organs that dangled enticingly between them. You mock, but there are many ratfish who would differ. While inherently sexy by ratfish standards, this fellow was not particularly tasty to their ancient marine brethren (or humans today) — so not hugely sought after as a food source or prey.
A little further again from the ratfish site we reach the contact of the two Formations. The rocks here have travelled a long way to their current location. With them, we peel away the layers of the geologic history of both the Comox Valley and the province of British Columbia.
The Trent River is not far from the Puntledge, a river whose banks have also revealed many wonderful fossil specimens. The Puntledge is also the name used by the K'ómoks First Nation to describe themselves. They have lived here since time immemorial. Along with Puntledge, they refer to themselves as Sahtloot, Sasitla and Ieeksun.
References: Note on the occurrence of the marine turtle Desmatochelys (Reptilia: Chelonioidea) from the Upper Cretaceous of Vancouver Island Elizabeth L. Nicholls Canadian Journal of Earth Sciences (1992) 29 (2): 377–380. https://doi.org/10.1139/e92-033; References: Chimaeras - The Neglected Chondrichthyans". Elasmo-research.org. Retrieved 2017-07-01.
Directions: If you're keen to explore the area, park on the side of Highway 19 about three kilometres south of Courtenay and hike up to the Trent River. Begin to look for parking about three kilometres south of the Cumberland Interchange. There is a trail that leads from the highway down beneath the bridge which will bring you to the Trent River's north side.
Carnotaurus sastrei, a genus of large theropod dinosaurs that roamed the southern tip of Argentina, South America during the Late Cretaceous, 72 to 69.9 million years ago. His name means "flesh-eating bull,' and he lives up to it.
This fellow — or at least his robust skull with the short, knobby eyebrow horns and fierce-looking teeth — is on display at the Natural History Museum in Madrid, Spain. For now, he is the only known genus of this species of bipedal predator.
The first specimen of Carnotaurus sastrei was found in Chubut on vast plains between the Andes Mountains and the Atlantic Ocean. A physician, Dr. A'ngel Tailor noticed a large concretion showing some bone fragments. A team led by José F. Bonaparte excavated the find in 1984 as part of a paleontological expedition funded by the Argentine Museum of Natural Sciences.
Sadly, Bonaparte — the Maestro del Mesozoico — passed away the 18th February 20220 at the age of 91. He spent the majority of his career as head of the Vertebrate Palaeontology Division of the Museo Argentino de Ciencias Naturales “Bernardino Rivadavia,” in Buenos Aires. Bonaparte opened up the vertebrate finds of Argentina to the world. He was instrumental in the finding, excavating and naming many iconic dinosaurs — Carnotaurus, Amargasaurus, Abelisaurus, Argentinosaurus, Noasaurus along with the finding of the first fossilised remains of Mesozoic South American mammals. He mentored many palaeontologists who will miss his keen eye and tremendous work ethic — Luis Chiappe, Rodolfo Coria, Agustín Martinelli, Fernando Novas, Jaime Powell, Guillermo Rougier, Leonardo Salgado, Sebastián Apesteguía and many others.
His excavation of Carnotaurus was the first of its kind and he recognized that the skull is quite unusual. Initially, it has a very marine reptile feel — but make no mistake this guy is clearly a terrestrial theropod. He had smallish, underdeveloped arms — teeny by theropod standards. Once you look closer you see his bull-like horns from whence he gets his name — horns that imply battle between rivals for the best meal, sexual partner and to be the one who leads the herd.
He was covered in leathery skin lined with rows of cone-shaped nodules or bumps. These get larger as they move towards his spine. He had forward-facing eyes, similar to tyrannosaurs like T-rex and smaller theropods like Velociraptor and Troodon — who had better vision even that T-rex — which would have given him the advantage of binocular vision and depth perception. Forward-facing eyes are also quite helpful with nocturnal hunting — think owls and cats — as they take in more light and help with nighttime predation. So perhaps this flesh-eating bull fancied a late-night snack on his menu from time to time.
Species like squirrels, pigeons and crocodiles have eyes on the sides of their heads. They lack the important competitive feature of well-developed depth perception — being able to easily and estimate distance — but perhaps make up for it with a panorama that offers a wider field of view.
Ophthalmoplax brasiliana / Photo: José F. Ventura |
Rene was a mountain goat in the field, stalking the hills in his signature red t-shirt. He was tremendously knowledgeable about the natural world and delighted in it. For many years, he was Chair of the White Rock and Surrey Naturalists, while I was Chair of the Vancouver Paleontological Society. Together, we would plan and often co-lead field trips to many of the wonderful fossil outcrops in British Columbia and Washington state.
In 2002, we were planning a very exciting round of field trips. I was offered a fully paid trip to India with Karen Lund to hike to the headwaters of the Ganges, a trip which I was to forgo in favour of a hike up to the outcrops of the Cathedral Escarpment and Burgess Shale and then to yummy Lower Jurassic and Lower Cretaceous, Albian, outcrops accessed only by boat in Haida Gwaii.
Rene and I had talked about "walking in the shoes" of Joseph Whiteaves, the GSC's chief palaeontologist in Ottawa. He published a paper in 1876 describing the Jurassic and Cretaceous faunas of Skidegate Inlet and spent a significant portion of his career working out the fossil fauna of the Burgess Shale. Combining these two sites within the same field season was a fitting homage.
John Fam, Vancouver Paleontological Society (VanPS) and Dan Bowen, Vancouver Island Palaeontological Society (VIPS), did much of the planning for that Haida Gwaii trip, they too being inspired by Whiteaves papers and the work of James Richardson and George Dawson — as a whole, we were giddy with the prospect of the year ahead.
Rene and I had planned to do both, but in the end, I had to give up the hike to Burgess that year and Rene never made it back to join me in Haida Gwaii.
Rene Savenye |
I take heart that he lived and died doing what he loved most. I got the news a few days later and cried for the loss of a great friend. I am sharing my memory of him with you so that you can remember him, too, and share in the delight and loss of one of the loveliest men to ever walk our planet. His years of teaching, mentoring, encouragement and generosity have helped shape natural science and those who have gone on to make it their passion or career — or happily, both.
Rene's name will not be forgotten to science. His namesake, H. Savenyei, is a lovely fossil halictine bee from Early Eocene deposits near Quilchena, British Columbia — and the first bee body-fossil known from the Okanagan Highlands — and indeed from Canada.
As a school teacher, Rene once taught the, then student, now SFU biology instructor, Rolf Mathewes. Rene passed his scientifically valuable specimen to Mathews, knowing it was important to science. Mathewes brought it to the attention of Bruce Archibald and Michael Engel, who described Rene's bee in the Canadian Journal of Zoology. Their work is a lovely legacy to a wonderful man and a specimen from one of his favourite collecting sites — Quilchena — a small road-cut exposure of the Coldwater beds of the Princeton Group, one of several depositional basins in the Merritt region of south-central British Columbia.
Rene is also remembered in spirit by the British Columbia Paleontological Alliance (BCPA) Rene Savenye Award. It was established in 2003 to honour those who have demonstrated outstanding service to the science of palaeontology or to palaeontological education in British Columbia.
Notable past recipients are a veritable who's who from the Pacific Northwest — Graham Beard of Qualicum in 2005, Charles Helm of Tumbler Ridge in 2011, Pat Trask of Courtenay in 2014, Rod Bartlett in 2016, and Joseph "Joe" Haegert in 2018. I'll share a link to the award below so you can read more at your leisure about Rene and those who bear the award with his name.
About H. Savenyei, (Engel & Archibald, 2003): The type specimen is a fairly well preserved complete adult female preserved with portions of the fore-wings and hind-wings. The specimen is 7.04 millimetres (0.277 in) long with the possibility of alteration in length during fossilization. The sections of the forewing which are preserved are approximately 4.8 millimetres (0.19 in) long and show dark brown to black colouration. The presence of a pygidial plate bordered by setae on the fifth metasomal tergum supports the placement into the Halictidae subfamily Halictinae. Placement into the tribe Halictini is based on the lack of a medial cleft in the fifth tergum.
References:
Archibald, B. & R. W. Mathewes. 2000. “Early Eocene Insects from Quilchena, BC, and their Paleoclimatic Implications.” Canadian Journal of Zoology, Volume 78, Number 6: pp 1441-1462.
Grimaldi, D. 1999. “The Co-radiations of Pollinating Insects and Angiosperms in the Cretaceous.” Annals of the Missouri Botanical Garden. 86: 373-406.
Photo: Halictidae sp.; Archibald and Mathewes 2000: 1453.
Rene Savenye Award: https://bcfossils.ca/rene-savenye-award
My Norwegian cousins on my mother's side call them humle. Norway is a wonderful place to be something wild as the wild places have not been disturbed by our hands.
There are an impressive thirty-five species of bumblebee species that call Norway hjem (home), and one, Bombus consobrinus, boasts the longest tongue that they use to feast solely on Monkshood, genus Aconitum, you may know by the name Wolf's-bane.
In the Kwak̓wala language of the Kwakwaka'wakw, speakers of Kwak'wala, and my family in the Pacific Northwest, bumblebees are known as ha̱mdzalat̕si — though I wonder if this is actually the word for a honey bee, Apis mellifera, as ha̱mdzat̕si is the word for a beehive.
I have a special fondness for all bees and look for them both in the garden and in First Nation art.
Bumblebees' habit of rolling around in flowers gives us a sense that these industrious insects are also playful. In First Nation art they provide levity — comic relief along with their cousins the mosquitoes and wasps — as First Nation dancers wear masks made to mimic their round faces, big round eyes and pointy stingers. A bit of artistic license is taken with their forms as each mask may have up to six stingers. The dancers weave amongst the watchful audience and swoop down to playfully give many of the guests a good, albeit gentle, poke.
Honey bees actually do a little dance when they get back to the nest with news of an exciting new place to forage — truly they do. Bumblebees do not do a wee bee dance when they come home pleased with themselves from a successful foraging mission, but they do rush around excitedly, running to and fro to share their excitement. They are social learners, so this behaviour can signal those heading out to join them as they return to the perfect patch of wildflowers.
Bumblebees are quite passive and usually sting in defence of their nest or if they feel threatened. Female bumblebees can sting several times and live on afterwards — unlike honeybees who hold back on their single sting as its barbs hook in once used and their exit shears it off, marking their demise.They are important buzz pollinators both for our food crops and our wildflowers. Their wings beat at 130 times or more per second, literally shaking the pollen off the flowers with their vibration.
And they truly are busy bees, spending their days fully focused on their work. Bumblebees collect and carry pollen and nectar back to the nest which may be as much as 25% to 75% of their body weight.
And they are courteous — as they harvest each flower, they mark them with a particular scent to help others in their group know that the nectar is gone.
The food they bring back to the nest is eaten to keep the hive healthy but is not used to make honey as each new season's queen bees hibernate over the winter and emerge reinvigorated to seek a new hive each Spring. She will choose a new site, primarily underground depending on the bumblebee species, and then set to work building wax cells for each of her fertilised eggs.
Bumblebees are quite hardy. The plentiful hairs on their bodies are coated in oils that provide them with natural waterproofing. They can also generate more heat than their smaller, slender honey bee cousins, so they remain productive workers in cooler weather.
We see the first bumblebees arise in the fossil record 100 million years ago and diversify alongside the earliest flowering plants. Their evolution is an entangled dance with the pollen and varied array of flowers that colour our world.We have found many wonderful examples within the fossil record, including a rather famous Eocene fossil bee found by a dear friend and naturalist who has left this Earth, Rene Savenye.
His namesake, H. Savenyei, is a lovely fossil halictine bee from Early Eocene deposits near Quilchena, British Columbia — and the first bee body-fossil known from the Okanagan Highlands — and indeed from Canada.
It is a fitting homage, as bees symbolize honesty, playfulness and willingness to serve the community in our local First Nation lore and around the world — something Rene did his whole life.
Aioloceras besairiei (Collingnon, 1949) |
Red-Tailed Lemurs, Waiwai & Hedgehog |
Euhoplites is an extinct ammonoid cephalopod from the Lower Cretaceous, characterized by strongly ribbed, more or less evolute, compressed to inflated shells with flat or concave ribs, typically with a deep narrow groove running down the middle.
In some, ribs seem to zigzag between umbilical tubercles and parallel ventrolateral clavi. In others, the ribs are flexious and curve forward from the umbilical shoulder and lap onto either side of the venter.
Its shell is covered in the lovely lumps and bumps we associate with the genus. The function of these adornments are unknown. I wonder if they gave them greater strength to go deeper into the ocean to hunt for food.
They look to have been a source of hydrodynamic drag, likely preventing Euhoplites from swimming at speed. Studying them may give some insight into the lifestyle of this ancient marine predator. Euhoplites had shells ranging in size up to a 5-6cm.
We find them in Lower Cretaceous, middle to upper Albian age strata. Euhoplites has been found in Middle and Upper Albian beds in France where it is associated respectively with Hoplites and Anahoplites, and Pleurohoplites, Puzosia, and Desmoceras; in the Middle Albian of Brazil with Anahoplites and Turrilites; and in the Cenomanian of Texas.
This species is the most common ammonite from the Folkstone Fossil Beds in southeastern England where a variety of species are found, including this 37mm beauty from the collections of José Juárez Ruiz.
Humpback Whale, Megaptera novaeangliae |