LOWER LIAS LYTOCERAS

A superbly prepped and extremely rare Lytoceras (Suess, 1865) ammonite found as a green ammonite nodule by Matt Cape in the Lower Lias of Dorset. 

Lytoceras are rare in the Lower Lias of Dorset — apart from the Belemnite Stone horizon — so much so that Paul Davis, whose skilled prep work you see here, initially thought it might be a Becheiceras hidden within the large, lumpy nodule. 

One of the reasons these lovelies are rarely found from here is that they are a Mediterranean Tethyian genus. The fossil fauna we find in the United Kingdom are dominated by Boreal Tethyian genera. 

We do find Lytoceras sp. in the Luridum subzone of the Pliensbachian showing that there was an influx of species from the Mediterranean realm during this time. This is the first occurrence of a Lytoceras that he has ever seen in a green nodule and Paul's seen quite a few. 

This absolutely cracking specimen was found and is in the collections of the awesome Matt Cape. Matt recognized that whatever was hidden in the nodule would take skilled and careful preparation using air scribes. Indeed it did. It took more than five hours of time and skill to unveil the lovely museum-worthy specimen you see here. 

We find Lytoceras in more than 1,000 outcrops around the globe ranging from the Jurassic through to the Cretaceous, some 189.6 to 109.00 million years ago. Once this specimen is fully prepped with the nodule material cut or scraped away, you can see the detailed crinkly growth lines or riblets on the shell and none of the expected coarse ribbing. 

Lytoceras sp. Photo: Craig Chivers

If you imagine running your finger along these, you would be tracing the work of decades of growth of these cephalopods. 

While we cannot know their actual lifespans, but we can make a healthy guess. 

The nautilus, their closest living cousins live upwards of 20 years — gods be good — and less than three years if conditions are poor.

The flanges, projecting flat ribs or collars, develop at the edge of the mouth border on the animal's mantle as they grow each new chamber. 

Each delicate flange grows over the course of the ammonites life, marking various points in time and life stages as the ammonite grew. There is a large variation within Lytoceras with regards to flanges. They provide both ornamentation and strength to the shell to protect it from water pressure as they moved into deeper seas.

The concretion prior to prep

This distinctive genus with its evolute shells are found in the Cretaceous marine deposits of: 

Antarctica (5 collections), Austria (19), Colombia (1), the Czech Republic (3), Egypt (2), France (194), Greenland (16), Hungary (25), Italy (11), Madagascar (2), Mexico (1), Morocco (4), Mozambique (1), Poland (2), Portugal (1), Romania (1), the Russian Federation (2), Slovakia (3), South Africa (1), Spain (24), Tanzania (1), Trinidad and Tobago (1), Tunisia (25); and the United States of America (17: Alaska, California, North Carolina, Oregon).

We also find them in Jurassic marine outcrops in:

Austria (15), Canada (9: British Columbia), Chile (6), France (181), Germany (11), Greenland (1), Hungary (189), India (1), Indonesia (1), Iran (1), Italy (50), Japan (14), Kenya (2), Luxembourg (4), Madagascar (2), Mexico (1), Morocco (43), New Zealand (15), Portugal (1), Romania (5), the Russian Federation (1), Slovakia (1), Spain (6), Switzerland (2), Tunisia (11), Turkey (12), Turkmenistan (1), Ukraine (5), the United Kingdom (12), United States (11: Alaska, California) — in at least 977 known collections. 

References:

Sepkoski, Jack (2002). "A compendium of fossil marine animal genera (Cephalopoda entry)". Bulletins of American Paleontology. 363: 1–560. Archived from the original on 2008-05-07. Retrieved 2017-10-18.

Paleobiology Database - Lytoceras. 2017-10-19.

Systematic descriptions, Mesozoic Ammonoidea, by W.J Arkell, Bernhard Kummel, and C.W. Wright. 1957. Treatise on Invertebrate Paleontology, Part L. Geological Society of America and University of Kansas press.

UPPER CRETACEOUS MOTORCROSS SITE: VANCOUVER ISLAND

Steller's Jay, Cyanocitta stelleri

One of the classic Vancouver Island fossil localities is the Santonian-Maastrichtian, Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake, Nanaimo, British Columbia, Canada.

The quarry is no longer active as such though there is a busy little gravel quarry a little way down the road closer to Ammonite falls near Benson Creek Falls.

Today it is an active motocross site and remains one of the classic localities of the Nanaimo Group. We find well-preserved nautiloids and ammonites — Canadoceras, Pseudoschloenbachia, Epigoniceras — the bivalves — Inoceramus, Sphenoceramus— gastropods, and classic Nanaimo Group decapods — Hoploparia, Linuparus. We also find fossil fruit and seeds which tell the story of the terrestrial history of Vancouver Island.

Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake

It was John Fam, Vice-Chair, Vancouver Island Paleontological Society (VanPS), who originally told me about the locality. John is one of the most delightful and knowledgeable people you'd be well-blessed to meet.

While he lived on Vancouver Island, he was an active member of the VanPS back when I was Chair. Several of the best joint VIPS/VanPS paleontological expeditions were planned with or instigated by his passion for fossils. I tip my hat to him for his passion and shared love of all things paleo.

John grew up 15 minutes from the motocross locality and used to collect there a few times a week with his father. John has wonderful parents and since marrying his childhood sweetheart, the amazing Grace, those excellent genetics, curiosity and love of fossils are now being passed to a new generation. It's lovely to see John and Grace continuing tradition with two boys of their own.

I met John way back then and did an overnight at his parent's house the Friday before a weekend field trip to Jurassic Point. It was a joy to have him walk me through his collections and tell his stories from earlier years. After learning about the site from John, I headed up to the Motocross Pit with my Uncle Doug. He was a delightful man who grew up on the coast and had explored much of it but not the fossil site just 10-minutes from his home. It was wonderful to walk through time with him so many years ago and then again solo this past year with sadness in my belly that one of the best I've ever known has left this Earth.

Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake

There were some no trespassing signs up but no people around, so I walked the periphery looking for the bedrock of the Haslam.

The rocks we find here were laid down south of the equator as small, tropical islands. They rode across the Pacific heading north and slightly east over the past 80 million years to where we find them today.

Jim Haggart and Peter Ward have done much to increase our understanding of the molluscan fauna of the Nanaimo Group. Personally, both personify the charming Indiana Jones school of rugged manly palaeontologists you picture in popular film. Professionally, their singular contributions and collaborative efforts have helped shape our understanding of the correlation of Nanaimo Group fauna to those we find in the Gulf Islands of British Columbia and down in the San Juan Islands of Washington State.

Their work builds on the work of Usher (1952), Matsumoto (1959a, 1959b) and Mallory (1977). A healthy nod goes out to the work of Muller and Jeletzky (1970) for untangling the lithostratigraphic and biostratigraphic foundation for our knowledge of the Nanaimo Group.

Candoceras yokoyama, Photo: John Fam, VanPS

As I walked along the bedrock of the Haslam, a Steller's Jay, Cyanocitta stelleri, followed me from tree to tree making his guttural shook, shook, shook call. Instructive, he seemed to be encouraging me, timing his hoots to the beat of my hammer. Vancouver Island truly has glorious flora and fauna.

Fancy some additional reading? Check out a paper published in the Journal of Paleontology back in 1989 by Haggard and Ward on new Nanaimo Group Ammonites from British Columbia and Washington State.

In it, they look at the ammonite species Puzosia (Mesopuzosia) densicostata Matsumoto, Kitchinites (Neopuzosia) japonicus Spath, Anapachydiscus cf. A. nelchinensis Jones, Menuites cf. M. menu (Forbes), Submortoniceras chicoense (Trask), and Baculites cf. B. boulei Collignon are described from Santonian--Campanian strata of western Canada and northwestern United States.

Stratigraphic occurrences and ranges of the species are summarized and those taxa important for correlation with other areas in the north Pacific region and Late Cretaceous ammonite fauna of the Indo-Pacific region. Here's the link: https://www.jstor.org/stable/1305358?seq=1

Peter Ward is a prolific author, both of scientific papers and more popularized works. I highly recommend his book Gorgon: Paleontology, Obsession, and the Greatest Catastrophe in Earth's History. It is an engaging romp through a decade's research in South Africa's Karoo Desert.

Photo: Candoceras yokoyamai from Upper Cretaceous Haslam formation (Lower Campanian) near Nanaimo, British Columbia. One of the earliest fossils collected by John Fam (1993). Prepared using only a cold chisel and hammer. Photo & collection of John Fam, VIPS.

SAKARA MADAGASGAR: OXFORDIAN OUTCROPS

This big beastie is a superb specimen of the ammonite Lobolytoceras costellatum showing the intricate fractal pattern of its septa. 

This lovely measures to a whopping 230 mm and hails from Oxfordian outcrops near Sakara, Madagascar. Lovingly prepped by the supremely talented José Juárez Ruiz.

Ammonites were predatory, squidlike creatures that lived inside coil-shaped shells. Like other cephalopods, ammonites had sharp, beak-like jaws inside a ring of squid-like tentacles that extended from their shells. They used these tentacles to snare prey — plankton, vegetation, fish and crustaceans — similar to the way a squid or octopus hunt today.

Catching a fish with your hands is no easy feat, as I'm sure you know. Ammonites did the equivalent, catching prey in their tentacles. They were skilled and successful hunters. They caught their prey while swimming and floating in the water column. 

Within their shells, they had a number of chambers, called septa, filled with gas or fluid that were interconnected by a wee air tube. By pushing air in or out, they were able to control their buoyancy in the water column.

They lived in the last chamber of their shells, continuously building new shell material as they grew. As each new chamber was added, the squid-like body of the ammonite would move down to occupy the final outside chamber.

They were a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids — octopuses, squid, and cuttlefish) then they are to shelled nautiloids such as the living Nautilus species.

Ammonites have intricate and complex patterns on their shells called sutures. The suture patterns differ across species and tell us what time period the ammonite is from. If they are geometric with numerous undivided lobes and saddles and eight lobes around the conch, we refer to their pattern as goniatitic, a characteristic of Paleozoic ammonites.

Ammonites first appeared about 240 million years ago, though they descended from straight-shelled cephalopods called bacrites that date back to the Devonian, about 415 million years ago, and the last species vanished in the Cretaceous–Paleogene extinction event.

They were prolific breeders that evolved rapidly. If you could cast a fishing line into our ancient seas, it is likely that you would hook an ammonite, not a fish. They were prolific back in the day, living (and sometimes dying) in schools in oceans around the globe. We find ammonite fossils (and plenty of them) in sedimentary rock from all over the world.

In some cases, we find rock beds where we can see evidence of a new species that evolved, lived and died out in such a short time span that we can walk through time, following the course of evolution using ammonites as a window into the past.

For this reason, they make excellent index fossils. An index fossil is a species that allows us to link a particular rock formation, layered in time with a particular species or genus found there. Generally, deeper is older, so we use the sedimentary layers rock to match up to specific geologic time periods, rather the way we use tree-rings to date trees. A handy way to compare fossils and date strata across the globe.

ANAHOPLITES PLANUS OF VILLEMOYENNE QUARRY

A beautiful specimen of the ammonite, Anahoplites planus (Mantell, 1822) from Albian deposits in Villemoyenne Quarry, Courcelles, Aube, north-central France.

Anahoplites (Hyatt, 1900) is a genus of compressed hoplitid ammonites with flat sides, narrow, flat or grooved venters, and flexious ribs or striae arising from weak umbilical tubercles that end in fine dense ventrolateral nodes.

This lovely has attracted some roommates — an oyster, some bryozoans and worm tubes are attached to her shell.

Anahoplites is now included in the subfamily Anahoplitinae and separated from the Hoplitinae where it was placed in the older in the 1957 edition of the Treatise on Invertebrate Paleontology, Part L (Ammonoidea). Genera of the Hoplitinae tend to be more robust, with broader whorls and stronger ribs.

Anahoplites is found in Cretaceous (Middle to the Late Albian) deposits from England, through Europe, all the way to the Transcaspian Oblast region in Russia to the east of the Caspian Sea. The Aube department, named after the local river, is the type locality of the Albian stage (d'ORBIGNY, 1842). 

A. planus from the French Coast

Two formations are recognized in the clay facies (the "Gault" auct.) of the stratotype, the Argiles tégulines de Courcelles (82 m), overlain by the Marnes de Brienne (43 m). The boundary between the two formations is well-defined at the top of an indurated bed and readily identifiable in the field.

This involute (113 mm) specimen shows evidence of cohabitation by some of his marine peers. 

We see two different bryozoa, an oyster and some serpulids making a living and leaving trace fossils on her flat sides. The top specimen was prepared with potase by José Juárez Ruiz of Spain. 

The lovely Anahoplites planus you see here to the lower right was found by Bertus op den Dries on the French coast in Albian deposits near Wissant, P5 and measures in at 8 cm. This on edge view gives you a very good sense of the keel.

AMMONITE FROM FOLKSTONE

Sheer beauty — a beautiful Euhoplites ammonite from Folkstone, UK. I've been really enjoying looking at all oil-in-water colouring and chunkiness of these ammonites.

Euhoplites is an extinct ammonoid cephalopod from the Lower Cretaceous, characterized by strongly ribbed, more or less evolute, compressed to inflated shells with flat or concave ribs, typically with a deep narrow groove running down the middle.

In some, ribs seem to zigzag between umbilical tubercles and parallel ventrolateral clavi. In others, the ribs are flexious and curve forward from the umbilical shoulder and lap onto either side of the venter.

Its shell is covered in the lovely lumps and bumps we associate with the genus. The function of these adornments are unknown. I wonder if they gave them greater strength to go deeper into the ocean to hunt for food. 

They look to have been a source of hydrodynamic drag, likely preventing Euhoplites from swimming at speed. Studying them may give some insight into the lifestyle of this ancient marine predator. Euhoplites had shells ranging in size up to a 5-6cm. 

We find them in Lower Cretaceous, middle to upper Albian age strata. Euhoplites has been found in Middle and Upper Albian beds in France where it is associated respectively with Hoplites and Anahoplites, and Pleurohoplites, Puzosia, and Desmoceras; in the Middle Albian of Brazil with Anahoplites and Turrilites; and in the Cenomanian of Texas.

This species is the most common ammonite from the Folkstone Fossil Beds in southeastern England where a variety of species are found, including this 37mm beauty from the collections of José Juárez Ruiz.

GIGANTIC FOSSIL AMMONITE OF FERNIE

Titanites occidentalis, Fernie Ammonite

The Fernie ammonite, Titanites occidentalis, from outcrops on Coal Mountain near Fernie, British Columbia, Canada. 

This beauty is the remains of a carnivorous cephalopod within the family Dorsoplanitidae that lived and died in a shallow sea some 150 million years ago.

If you would like to get off the beaten track and hike up to see this ancient beauty, you will want to head to the town of Fernie in British Columbia close to the Alberta border. 

This is the traditional territory of the the Yaq̓it ʔa·knuqⱡi ‘it First Nation who have lived here since time immemorial. There was some active logging along the hillside in 2021, so if you are looking at older directions on how to get to the site be mindful that many of the trailheads have been altered and a fair bit of bushwhacking will be necessary to get to the fossil site proper. That being said, the loggers from CanWel may have clear-cut large sections of the hillside but they did give the ammonite a wide berth and have left it intact.

Wildsight, a non-profit environmental group out of the Kimberly Cranbrook area has been trying to gain grant funding to open up the site as an educational hike with educational signage for folks visiting the Fernie area. It is likely the province of British Columbia would top up those funds if they are able to place the ammonite under the Heritage Conservation Act. CanWel would remain the owners of the land but the province could assume the liability for those visiting this iconic piece of British Columbia's palaeontological history. 

Driving to the trail base is along an easy access road just east of town along Fernie Coal Road. There are some nice exposures of Cretaceous plant material on the north side (left-hand side) of the road as you head from Fernie towards Coal Creek. I recently drove up to Fernie to look at Cretaceous plant material and locate the access point to the now infamous Late Jurassic (Tithonian) Titanites (S.S. Buckman, 1921) site. While the drive out of town is on an easy, well-maintained road, the slog up to the ammonite site is often a wet, steep push.

Fernie, British Columbia, Canada

The first Titanites occidentalis was about one-third the size and was incorrectly identified as Lytoceras, a fast-moving nektonic carnivore. The specimen you see here is significantly larger at 1.4 metres (about four and a half feet) and rare in North America. 

Titanites occidentalis, the Western Giant, is the second known specimen of this extinct fossil species. 

The first was discovered in 1947 in nearby Coal Creek by a British Columbia Geophysical Society mapping team. When they first discovered this marine fossil high up on the hillside, they could not believe their eyes — both because it is clearly marine at the top of a mountain and the sheer size of this ancient beauty.

In the summer of 1947, a field crew was mapping coal outcrops for the BC Geological Survey east of Fernie. One of the students reported finding “a fossil truck tire.” Fair enough. The similarity of size and optics are pretty close to your average Goodridge. 

A few years later, GSC Paleontologist Hans Frebold described and named the fossil Titanites occidentalis after the large Jurassic ammonites from Dorset, England. The name comes from Greek mythology. Tithonus, as you may recall, was the Prince of Troy. He fell in love with Eos, the Greek Goddess of the Dawn. Eos begged Zeus to make her mortal lover immortal. Zeus granted her wish but did not grant Tithonus eternal youth. He did indeed live forever — ageing hideously. Ah, Zeus, you old trickster. It is a clever play on time placement. Dawn is the beginning of the day and the Tithonian being the latest age of the Late Jurassic. Clever Hans!

HIKING TO THE FERNIE AMMONITE

From the town of Fernie, British Columbia, head east along Coal Creek Road towards Coal Creek. The site is 3.81 km from the base of Coal Creek Road to the trailhead as the crow flies. I have mapped it here for you in yellow and added the wee purple GPS marker for the ammonite site proper. There is a nice, dark grey to black roadcut exposure of Cretaceous plants on the north side of the dirt road that is your cue to pull over and park.  

You access what is left of the trailhead on the south side of the road. You will need to cross the creek to begin your ascent. There is no easy way across the creek and you'll want to tackle this one with a friend when the water level is low. 

The beginning of the trail is not clear but a bit of searching will reveal the trailhead with its telltale signs of previous hikers. This is a moderate 6.3-kilometre hike up & back bushwhacking through scrub and fallen trees. Heading up, you will make about a 246-metre elevation gain. You will likely not have a cellular signal up here but if you download the Google Map to your mobile, you will have GPS to guide you. The area has been recently logged so much of the original trail has been destroyed. There may now be easier vehicle access up the logging roads but I have not driven them since the logging and new road construction.

If you are coming in from out of town, the closest airport is Cranbrook. Then it is about an hour and change to Fernie and another 15-minutes or so to park near the site.

You will want to leave your hammers with your vehicle (no need to carry the weight and this lovely should never be struck with anything more than a raindrop) as this site is best enjoyed with a camera. 

This is a site you will want to wear hiking boots to access. Know that these will get wet as you cross the creek. 

If you would like to see the ammonite but are not keen on the hike, a cast has been made by fossil preparator Rod Bartlett is on display at the Courtenay Museum in Courtenay, Vancouver Island, Canada. 

Respect for the Land / Leave No Trace

As your feet move up the hillside, you can imagine this land 10,000 years ago, rising above great glaciers. Where footfalls trace the steps of those that came before you. This land has been home to the Yaq̓it ʔa·knuqⱡi ‘it First Nation and Ktunaxa or Kukin ʔamakis First Nations whose oral history have them living here since time immemorial. Like them, take only what you need and no more than the land offers — packing out anything that you packed in. 

Fernie Ammonite Palaeo Coordinates: 49°29'04"N 115°00'49"W

AMMONITE TIME KEEPERS

Argonauticeras besairei, José Juárez Ruiz

An exceptional example of the fractal building of an ammonite septum, in this clytoceratid Argonauticeras besairei from the awesome José Juárez Ruiz.

Ammonites were predatory, squidlike creatures that lived inside coil-shaped shells.

Like other cephalopods, ammonites had sharp, beak-like jaws inside a ring of squid-like tentacles that extended from their shells. 

They used these tentacles to snare prey, — plankton, vegetation, fish and crustaceans — similar to the way a squid or octopus hunt today.

Catching a fish with your hands is no easy feat, as I am sure you know. But the Ammonites were skilled and successful hunters. They caught their prey while swimming and floating in the water column. 

Within their shells, they had a number of chambers, called septa, filled with gas or fluid that were interconnected by a wee air tube. By pushing air in or out, they were able to control their buoyancy in the water column.

They lived in the last chamber of their shells, continuously building new shell material as they grew. As each new chamber was added, the squid-like body of the ammonite would move down to occupy the final outside chamber.

They were a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. 

These molluscs, commonly referred to as ammonites, are more closely related to living coleoids — octopuses, squid, and cuttlefish) than they are to shelled nautiloids such as the living Nautilus species.

The Ammonoidea can be divided into six orders:

• Agoniatitida, Lower Devonian - Middle Devonian
• Clymeniida, Upper Devonian
• Goniatitida, Middle Devonian - Upper Permian
• Prolecanitida, Upper Devonian - Upper Triassic
• Ceratitida, Upper Permian - Upper Triassic
• Ammonitida, Lower Jurassic - Upper Cretaceous

Ammonites have intricate and complex patterns on their shells called sutures. The suture patterns differ across species and tell us what time period the ammonite is from. If they are geometric with numerous undivided lobes and saddles and eight lobes around the conch, we refer to their pattern as goniatitic, a characteristic of Paleozoic ammonites.

If they are ceratitic with lobes that have subdivided tips; giving them a saw-toothed appearance and rounded undivided saddles, they are likely Triassic. For some lovely Triassic ammonites, take a look at the specimens that come out of Hallstatt, Austria and from the outcrops in the Humboldt Mountains of Nevada.

Hoplites bennettiana (Sowby, 1826) Christophe Marot

If they have lobes and saddles that are fluted, with rounded subdivisions instead of saw-toothed, they are likely Jurassic or Cretaceous. If you'd like to see a particularly beautiful Lower Jurassic ammonite, take a peek at Apodoceras. Wonderful ridging in that species.

One of my favourite Cretaceous ammonites is the ammonite, Hoplites bennettiana (Sowby, 1826). This beauty is from Albian deposits near Carrière de Courcelles, Villemoyenne, near la région de Troyes (Aube) Champagne in northeastern France.

At the time that this fellow was swimming in our oceans, ankylosaurs were strolling about Mongolia and stomping through the foliage in Utah, Kansas and Texas. Bony fish were swimming over what would become the strata making up Canada, the Czech Republic and Australia. Cartilaginous fish were prowling the western interior seaway of North America and a strange extinct herbivorous mammal, Eobaatar, was snuffling through Mongolia, Spain and England.

In some classifications, these are left as suborders, included in only three orders: Goniatitida, Ceratitida, and Ammonitida. Once you get to know them, ammonites in their various shapes and suturing patterns make it much easier to date an ammonite and the rock formation where it is found.

Ammonites first appeared about 240 million years ago, though they descended from straight-shelled cephalopods called bacrites that date back to the Devonian, about 415 million years ago, and the last species vanished in the Cretaceous–Paleogene extinction event.

They were prolific breeders that evolved rapidly. If you could cast a fishing line into our ancient seas, it is likely that you would hook an ammonite, not a fish. They were prolific back in the day, living (and sometimes dying) in schools in oceans around the globe. We find ammonite fossils (and plenty of them) in sedimentary rock from all over the world.

In some cases, we find rock beds where we can see evidence of a new species that evolved, lived and died out in such a short time span that we can walk through time, following the course of evolution using ammonites as a window into the past.

For this reason, they make excellent index fossils. An index fossil is a species that allows us to link a particular rock formation, layered in time with a particular species or genus found there. 

Generally, deeper is older, so we use the sedimentary layers of rock to match up to specific geologic time periods, rather like the way we use tree rings to date trees. A handy way to compare fossils and date strata across the globe.

References: Inoue, S., Kondo, S. Suture pattern formation in ammonites and the unknown rear mantle structure. Sci Rep 6, 33689 (2016). https://doi.org/10.1038/srep33689

https://www.nature.com/articles/srep33689?fbclid=IwAR1BhBrDqhv8LDjqF60EXdfLR7wPE4zDivwGORTUEgCd2GghD5W7KOfg6Co#citeas

Photos: Argonauticeras besairei from the awesome José Juárez Ruiz.

Photo: Hoplites bennettiana from near Troyes, France. Collection de Christophe Marot

THE GIANT FOSSIL AMMONITE OF FERNIE, BRITISH COLUMBIA

Titanites occidentalis, Fernie Ammonite

The Fernie ammonite, Titanites occidentalis, from outcrops on Coal Mountain near Fernie, British Columbia, Canada. 

This beauty is the remains of a carnivorous cephalopod within the family Dorsoplanitidae that lived and died in a shallow sea some 150 million years ago.

If you would like to get off the beaten track and hike up to see this ancient beauty, you will want to head to the town of Fernie in British Columbia close to the Alberta border. 

This is the traditional territory of the the Yaq̓it ʔa·knuqⱡi ‘it First Nation who have lived here since time immemorial. There was some active logging along the hillside in 2021, so if you are looking at older directions on how to get to the site be mindful that many of the trailheads have been altered and a fair bit of bushwhacking will be necessary to get to the fossil site proper. That being said, the loggers from CanWel may have clear-cut large sections of the hillside but they did give the ammonite a wide berth and have left it intact.

Wildsight, a non-profit environmental group out of the Kimberly Cranbrook area has been trying to gain grant funding to open up the site as an educational hike with educational signage for folks visiting the Fernie area. It is likely the province of British Columbia would top up those funds if they are able to place the ammonite under the Heritage Conservation Act. CanWel would remain the owners of the land but the province could assume the liability for those visiting this iconic piece of British Columbia's palaeontological history. 

Driving to the trail base is along an easy access road just east of town along Fernie Coal Road. There are some nice exposures of Cretaceous plant material on the north side (left-hand side) of the road as you head from Fernie towards Coal Creek. I recently drove up to Fernie to look at Cretaceous plant material and locate the access point to the now infamous Late Jurassic (Tithonian) Titanites (S.S. Buckman, 1921) site. While the drive out of town is on an easy, well-maintained road, the slog up to the ammonite site is often a wet, steep push.

Fernie, British Columbia, Canada

The first Titanites occidentalis was about one-third the size and was incorrectly identified as Lytoceras, a fast-moving nektonic carnivore. The specimen you see here is significantly larger at 1.4 metres (about four and a half feet) and rare in North America. 

Titanites occidentalis, the Western Giant, is the second known specimen of this extinct fossil species. 

The first was discovered in 1947 in nearby Coal Creek by a British Columbia Geophysical Society mapping team. When they first discovered this marine fossil high up on the hillside, they could not believe their eyes — both because it is clearly marine at the top of a mountain and the sheer size of this ancient beauty.

In the summer of 1947, a field crew was mapping coal outcrops for the BC Geological Survey east of Fernie. One of the students reported finding “a fossil truck tire.” Fair enough. The similarity of size and optics are pretty close to your average Goodridge. 

A few years later, GSC Paleontologist Hans Frebold described and named the fossil Titanites occidentalis after the large Jurassic ammonites from Dorset, England. The name comes from Greek mythology. Tithonus, as you may recall, was the Prince of Troy. He fell in love with Eos, the Greek Goddess of the Dawn. Eos begged Zeus to make her mortal lover immortal. Zeus granted her wish but did not grant Tithonus eternal youth. He did indeed live forever — ageing hideously. Ah, Zeus, you old trickster. It is a clever play on time placement. Dawn is the beginning of the day and the Tithonian being the latest age of the Late Jurassic. Clever Hans!

HIKING TO THE FERNIE AMMONITE

From the town of Fernie, British Columbia, head east along Coal Creek Road towards Coal Creek. The site is 3.81 km from the base of Coal Creek Road to the trailhead as the crow flies. I have mapped it here for you in yellow and added the wee purple GPS marker for the ammonite site proper. There is a nice, dark grey to black roadcut exposure of Cretaceous plants on the north side of the dirt road that is your cue to pull over and park.  

You access what is left of the trailhead on the south side of the road. You will need to cross the creek to begin your ascent. There is no easy way across the creek and you'll want to tackle this one with a friend when the water level is low. 

The beginning of the trail is not clear but a bit of searching will reveal the trailhead with its telltale signs of previous hikers. This is a moderate 6.3-kilometre hike up & back bushwhacking through scrub and fallen trees. Heading up, you will make about a 246-metre elevation gain. You will likely not have a cellular signal up here but if you download the Google Map to your mobile, you will have GPS to guide you. The area has been recently logged so much of the original trail has been destroyed. There may now be easier vehicle access up the logging roads but I have not driven them since the logging and new road construction.

If you are coming in from out of town, the closest airport is Cranbrook. Then it is about an hour and change to Fernie and another 15-minutes or so to park near the site.

You will want to leave your hammers with your vehicle (no need to carry the weight and this lovely should never be struck with anything more than a raindrop) as this site is best enjoyed with a camera. 

This is a site you will want to wear hiking boots to access. Know that these will get wet as you cross the creek. 

If you would like to see the ammonite but are not keen on the hike, a cast has been made by fossil preparator Rod Bartlett is on display at the Courtenay Museum in Courtenay, Vancouver Island, Canada. 

Respect for the Land / Leave No Trace

As your feet move up the hillside, you can imagine this land 10,000 years ago, rising above great glaciers. Where footfalls trace the steps of those that came before you. This land has been home to the Yaq̓it ʔa·knuqⱡi ‘it First Nation and Ktunaxa or Kukin ʔamakis First Nations whose oral history have them living here since time immemorial. Like them, take only what you need and no more than the land offers — packing out anything that you packed in. 

Fernie Ammonite Palaeo Coordinates: 49°29'04"N 115°00'49"W

QUENSTEDTOCERAS WITH PATHOLOGY

What you are seeing here is a protuberance extruding from the venter of Quenstedtoceras cf. leachi (Sowerby). It is a pathology in the shell from hosting immature bivalves that shared the seas with these Middle Jurassic, Upper Callovian, Lamberti zone fauna from the Volga River basin. The collecting site is the now inactive Dubki commercial clay quarry and brickyard near Saratov, Russia. 

The site has produced thousands of ammonite specimens. A good 1,100 of those ended up at the Black Hills Institute of Geological Research in Hill City, South Dakota. 

Roughly 1,000 of those are Quenstedtoceras (Lamberticeras) lamberti and the other 100 are a mix of other species found in the same zone. These included Eboraciceras, Peltoceras, Kosmoceras, Grossouvria, Proriceras, Cadoceras and Rursiceras. 

What is especially interesting is the volume of specimens — 167 Quenstedtoceras (Lamberticeras) lamberti and 89 other species in the Black Hills collection — with healed predation injuries. It seems Quenstedtoceras (Lamberticeras) lamberti are the most common specimens found here and so not surprisingly the most common species found injured. Of the 1,000, 655 of the Quenstedtoceras (Lamberticeras) lamberti displayed some sort of deformation or growth on the shell or had grown in a tilted manner. 

Again, some of the Q. lamberti had small depressions in the centre likely due to a healed bite and hosting infestations of the immature bivalve Placunopsis and some Ostrea. 

The bivalves thrived on their accommodating hosts and the ammonites carried on, growing their shells right up and over their bivalve guests. This relationship led to some weird and deformities of their shells. They grow in, around, up and over nearly every surface of the shell and seem to have lived out their lives there. It must have gotten a bit unworkable for the ammonites, their shells becoming warped and unevenly weighted. Over time, both the flourishing bivalves and the ammonite shells growing up and over them produced some of the most interesting pathology specimens I have ever seen.    

In the photo here from Emil Black, you can see some of the distorted shapes of Quenstedtoceras sp. Look closely and you see a trochospiral or flattened appearance on one side while they are rounded on the other. 

All of these beauties hail from the Dubki Quarry near Saratov, Russia. The ammonites were collected in marl or clay used in brick making. The clay particles suggest a calm, deep marine environment. One of the lovely features of the preservation here is the amount of pyrite filling and replacement. It looks like these ammonites were buried in an oxygen-deficient environment. 

The ammonites were likely living higher in the water column, well above the oxygen-poor bottom. An isotopic study would be interesting to prove this hypothesis. There's certainly enough of these ammonites that have been recovered to make that possible. It's estimated that over a thousand specimens have been recovered from the site but that number is likely much higher. But these are not complete specimens. We mostly find the phragmocones and partial body chambers. Given the numbers, this may be a site documenting a mass spawning death over several years or generations.

If you fancy a read on all things cephie, consider picking up a copy of Cephalopods Present and Past: New Insights and Fresh Perspectives edited by Neil Landman and Richard Davis. Figure 16.2 is from page 348 of that publication and shows the hosting predation quite well. 

Photos: Courtesy of the deeply awesome Emil Black. These are in his personal collection that I hope to see in person one day. 

It was his sharing of the top photo and the strange anomaly that had me explore more about the fossils from Dubki and the weird and wonderful hosting relationship between ammonites and bivalves. Thank you, my friend!

INKY BEAUTY: AMMONITE OF PONGO DE MANSERICHE

This inky beauty is Prolyelliceras ulrichi (Knechtel, 1947) a fast-moving nektonic carnivorous ammonite from Cretaceous lithified, black, carbonaceous limestone outcrops in the Pongo de Manseriche gorge in northwest Peru. 

If you look closely, you can see that this specimen shows a pathology, a slight deviation to the side of the siphonal of the ammonite. We see Prolyelliceras from the Albian to Middle Albian from five localities in Peru.

The canyons of the Amazon River system in the eastern ranges of the Andes of Peru are known by the Indian name pongo. 

The most famous of these is the Pongo de Manseriche, cut by the Marañon River through the eastern range of the Andes, where it emerges from the cordillera into the flat terrane of the Amazon Basin. The fossil exposures here are best explored by boat. The reality of the collecting is similar to the imagined. I was chatting with Betty Franklin, VIPS, about this. They float along and pick up amazing specimen after amazing specimen. When the water rises, the ammonites are aided in their erosion out of the cliffs.  

The Pongo de Manseriche lies nearly 500 miles upstream from Iquitos, and consequently nearly 3,000 miles above the mouth of the Amazon River. It is situated in the heart of the montaña, in a vast region the ownership of which has long been in dispute between Peru and Ecuador, but over which neither country exercises any police or other governmental control. There is an ancient tradition of the indigenous people of the vicinity that one of their gods descended the Marañón and another ascended the Amazon to communicate with him. Together they opened the pass called the Pongo de Manseriche.

Reference: M. M. Knechtel. 1947. Cephalopoda. In: Mesozoic fossils of the Peruvian Andes, Johns Hopkins University Studies in Geology 15:81-139

W. J. Kennedy and H. C. Klinger. 2008. Cretaceous faunas from Zululand and Natal, South Africa. The ammonite subfamily Lyelliceratinae Spath, 1921. African Natural History 4:57-111. The beauty you see here is in the collection of the deeply awesome José Juárez Ruiz.

BACK IN THE USSR: BEADANTICERAS OF THE NORTHERN CAUCASUS

This lovely oil in water coloured ammonite is the beauty Beudanticeras sp. from the Lower Cretaceous (Upper Aptian), Krasnodar region, Northern Caucasus, southern Russia. 

This area of the world has beautiful fossil specimens with their distinct colouring. The geology and paleontological history of the region are fascinating as is its more recent history. 

The territory of present Krasnodar Krai was inhabited as early as the Paleolithic, about 2 million years ago. It was inhabited by various tribes and peoples since ancient times. There were several Greek colonies on the Black Sea coast, which later became part of the Kingdom of the Bosporus. In 631, the Great Bulgaria state was founded in the Kuban. In the 8th-10th centuries, the territory was part of Khazaria.

In 965, the Kievan Prince Svyatoslav defeated the Khazar Khanate and this region came under the power of Kievan Rus, Tmutarakan principality was formed. At the end of the 11th century, in connection with the strengthening of the Polovtsy and claims of Byzantium, Tmutarakan principality came under the authority of the Byzantine emperors (until 1204).

In 1243-1438, this land was part of the Golden Horde. After its collapse, Kuban was divided between the Crimean Khanate, Circassia, and the Ottoman Empire, which dominated in the region. Russia began to challenge the protectorate over the territory during the Russian-Turkish wars.

In 1783, by decree of Catherine II, the right-bank Kuban and Taman Peninsula became part of the Russian Empire after the liquidation of the Crimean Khanate. In 1792-1793, Zaporozhye (Black Sea) Cossacks resettled here to protect new borders of the country along the Kuban River. 

During the military campaign to establish control over the North Caucasus (Caucasian War of 1763-1864), in the 1830s, the Ottoman Empire for forced out of the region and Russia gained access to the Black Sea coast.

Prior to the revolutionary events of 1917, most of the territory of present Krasnodar Krai was occupied by the Kuban region, founded in 1860. In 1900, the population of the region was about 2 million people. In 1913, it ranked 2nd by the gross harvest of grain, 1st place for the production of bread in the Russian Empire.

The Kuban was one of the centres of resistance after the Bolshevik revolution of 1917. In 1918-1920, there was a non-Bolshevik Kuban People’s Republic. In 1924, North-Caucasian krai was founded with the centre in Rostov-on-Don. In 1934, it was divided into Azov-Black Sea krai (Rostov-on-Don) and North Caucasus krai (Stavropol).

September 13, 1937, the Azov-Black Sea region was divided into the Rostov region and Krasnodar Krai that included Adygei autonomous oblast. During the Second World War, the region was captured by the Germans. After the battle for the Caucasus, it was liberated. There are about 1,500 monuments and memorials commemorating heroes of the war on the territory of Krasnodar Krai.

The lovely block you see here is in the collections of the awesome John Fam, Vice-Chair of the Vancouver Paleontological Society in British Columbia, Canada.

PHYLLOCERAS VELLEDAE OF MADAGASCAR

This specimen of Phylloceras velledae (Michelin) has a shell with a small umbilicus, arched, acute venter, and at some growth stage, falcoid ribs that spring in pairs from umbilical tubercles, disappearing on the outer whorls.

This specimen has been polished to show the sutures to great effect. These ammonites are common in rock shops and plentiful on the internet.

These ammonites make a lovely addition to any teaching collection as they provide a lot of detail and can be handled quite well by small, less gentle hands. 

Like other cephalopods, ammonites had sharp, beak-like jaws inside a ring of squid-like tentacles that extended from their shells. 

They used these tentacles to snare prey, — plankton, vegetation, fish and crustaceans — similar to the way a squid or octopus hunt today.

Catching a fish with your hands is no easy feat, as I'm sure you know. But the Ammonites were skilled and successful hunters. They caught their prey while swimming and floating in the water column. 

Within their shells, they had a number of chambers, called septa, filled with gas or fluid that were interconnected by a wee air tube. By pushing air in or out, they were able to control their buoyancy in the water column.

They lived in the last chamber of their shells, continuously building new shell material as they grew. As each new chamber was added, the squid-like body of the ammonite would move down to occupy the final outside chamber.

They were a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids — octopuses, squid, and cuttlefish) than they are to shelled nautiloids such as the living Nautilus species.

DESMOCERAS OF MAHAJANGA

This lovely dark rust chunky monkey is the ammonite Desmoceras (Pseudouhligella) latidorsatum from the Lower Cretaceous, Lower Albian, Douvilliceras inequinodum Zone, Ambarimaninga, Mahajanga Province, Madagascar.

Ammonites were predatory, squid-like creatures that lived inside coil-shaped shells. Like other cephalopods, ammonites had sharp, beak-like jaws inside a ring of squid-like tentacles. 

They used their tentacles to snare prey, — plankton, vegetation, fish and crustaceans — similar to the way a squid or octopus hunt today.

JAPANESE CORKSCREW AMMONITE: HYPHANTOCERAS ORIENTALE

A stunning example of the heteromorph ammonite, Hyphantoceras orientale macroconch. This beauty corresponds to 'Morphotype C' from Aiba (2017). 

The specimen is a handful at 136 mm and was lovingly prepared by the hand holding it, that of the talented José Juárez Ruiz.

This an adult specimen (not the juvenile stage) from Upper Santonian outcrops near Ashibetsu, Hokkaido, Japan.

Aiba published on a possible phylogenetic relationship of two species of Hyphantoceras (Ammonoidea, Nostoceratidae) earlier this year, proposing that a phylogenetic relationship may exist based on newly found specimens with precise stratigraphic occurrences in the Kotanbetsu and Obira areas, northwestern Hokkaido.

Two closely related species, Hyphantoceras transitorium and H. orientale, were recognized in the examined specimens from the Kotanbetsu and Obira areas. Specimens of H. transitorium show wide intraspecific variation in the whorl shape. The stratigraphic occurrences of the two species indicate that they occur successively in the Santonian–lowermost Campanian, without stratigraphic overlapping. 

The similarity of their shell surface ornamentations and the stratigraphic relationships possibly suggest that H.orientale was derived from H. transitorium. The presumed lineage is likely indigenous to the northwestern Pacific realm in the Santonian–earliest Campanian. Hyphantoceras venustum and H. heteromorphum might stand outside a H. transitorium–H. orientale lineage, judging from differences of their shell surface ornamentation.

Aiba, Daisuke. (2019). A Possible Phylogenetic Relationship of Two Species of Hyphantoceras (Ammonoidea, Nostoceratidae) in the Cretaceous Yezo Group, Northern Japan. Paleontological Research. 23. 65-80. 10.2517/2018PR010.

MEET FERGUSONITES HENDERSONAE: HETTANGIAN AMMONITE

Fergusonites hendersonae (Longridge, 2008)

Meet Fergusonites hendersonae, a Late Hettangian (Early Jurassic) ammonite from the Taseko Lakes area of British Columbia, Canadian Rockies.

I had the very great honour of having this fellow, a new species of nektonic carnivorous ammonite, named after me by paleontologist Louse Longridge from the University of British Columbia. I'd met Louise as an undergrad and was pleased as punch to hear that she would be continuing the research by Dr. Howard Tipper.

We did several trips over the years up to the Taseko Lake area of the Rockies joined by many wonderful researchers from Vancouver Island Palaeontological Society and Vancouver Paleontological Society, as well as the University of British Columbia. Both Dan Bowen and John Fam were instrumental in planning those expeditions. We endured elevation sickness, rain, snow, grizzly bears and very chilly nights (we were sleeping right next to a glacier at one point) but were rewarded by the enthusiastic crew, helicopter rides (which really cut down the hiking time) excellent specimens and stunningly beautiful country. We were also blessed with excellent access as the area is closed to collecting except with a permit.

Reference: PaleoDB 157367 M. Clapham GSC C-208992, Section A 09, Castle Pass Angulata - Jurassic 1 - Canada, Longridge et al. (2008)

Full reference: L. M. Longridge, P. L. Smith, and H. W. Tipper. 2008. Late Hettangian (Early Jurassic) ammonites from Taseko Lakes, British Columbia, Canada. Palaeontology 51:367-404

PaleoDB taxon number: 297415; Cephalopoda - Ammonoidea - Juraphyllitidae; Fergusonites hendersonae Longridge et al. 2008 (ammonite); Average measurements (in mm): shell width 9.88, shell diameter 28.2; Age range: 201.6 to 196.5 Ma. Locality info: British Columbia, Canada (51.1° N, 123.0° W: paleo coordinates 22.1° N, 66.1° W)

CHAMPAGNE-ARDENNE HOPLITES

An excellent example of the ammonite, Hoplites bennettiana (Sowby, 1826) with a pathology. This beauty is from Albian deposits near Carrière de Courcelles, Villemoyenne, laid down in the Cretaceous near la région de Troyes (Aube) Champagne in northeastern France.

L'Albien or Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It was named after Alba, the Latin name for the River Aube, a tributary of the Seine that flows through the Champagne-Ardenne region of northwestern France.

The Albian is the youngest or uppermost subdivision of the Lower Cretaceous, approximately 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma (million years ago).

At the time that this fellow was swimming in our oceans, ankylosaurs were strolling about Mongolia and stomping through the foliage in Utah, Kansas and Texas. Bony fish were swimming over what would become the strata making up Canada, the Czech Republic and Australia. Cartilaginous fish were prowling the western interior seaway of North America and a strange extinct herbivorous mammal, Eobaatar, was snuffling through Mongolia, Spain and England. Interesting times.

Hoplites maritimus / Hoplites rudis

Hoplites are amongst my favourite ammonites. I still have a difficult time telling them apart. To the right, you can see a slightly greyish, Hoplites maritimus, from Sussex England. Below him is a brownish Hoplites rudis from outcrops between Courcelles and Troyes, France. There are many Hoplites species. Each has a nicely raised tire-track ribbing. My preference is for Hoplities bennetianus (or bennettiana). I'm still sorting out the naming of that species. The difference between Hoplites bennettiana and Hoplites dentatus is seen on the venter.

Hoplites shells have compressed, rectangular and trapezoidal whorl sections. They have pronounced umbilical bullae from which their prominent ribs branch out. The ends of the ribs can be both alternate or opposite. Some species have zigzagging ribs and these usually end thickened or raised into ventrolateral tubercules.

Ammonites were predatory, squid-like creatures that lived inside coil-shaped shells. Like other cephalopods, ammonites had sharp, beaklike jaws inside a ring of tentacles that extended from their shells to snare prey such as small fish and crustaceans. Some ammonites grew more than three feet (one meter) across — possible snack food for the giant mosasaur Tylosaurus.

Ammonites constantly built new shell as they grew, but only lived in the outer chamber. They scooted through the warm, shallow seas by squirting jets of water from their bodies. A thin, tubelike structure called a siphuncle reached into the interior chambers to pump and siphon air and helped them move through the water.

Ammonites first appeared about 240 million years ago, though they descended from straight-shelled cephalopods called bacrites that date back to the Devonian, about 415 million years ago. They were prolific breeders, lived in schools, and are among the most abundant fossils found today. They went extinct with the dinosaurs 65 million years ago. Scientists use the various shapes and sizes of ammonite shells that appeared and disappeared through the ages to date other fossils.

Hoplites sp. from the Early Cretaceous of Dorset, UK

During their evolution, three catastrophic events occurred. The first during the Permian period (250million years ago), only 10% survived.  They went on to flourish throughout the Triassic period, but at the end of this period (206 million years ago), all but one species died. Then they began to thrive from the Jurassic period until the end of the Cretaceous period when all species of ammonites became extinct.

Ammonites began life very tiny, less than 1mm in diameter, and were vulnerable to attack from predators. They fed on plankton and quickly assumed a strong protective outer shell. They also grew quickly with the females growing up to 400% larger than the males; because they needed the larger shell for egg production. Most ammonites only lived for two years.  Some lived longer becoming very large. The largest ever found was in Germany (6.5 feet in diameter).

Ammonites lived in shallow waters of 100 meters or less. They moved through the water by jet propulsion expelling water through a funnel-like opening to propel themselves in the opposite direction. They were predators (cephalopods) feeding on most living marine life including mollusks, fish even other cephalopods. Ammonites would silently stalk their prey then quickly extend their tentacles to grab it.  When caught the prey would be devoured by the Ammonites' jaws located at the base of the tentacles between the eyes.

Hoplites dentalus, from Albian deposits near Troyes, France

Most ammonites have coiled shells. The chambered part of the shell is called a phragmocone.  It contains a series of progressively layered chambers called camerae, which were divided by thin walls called septae. The last chamber is the body chamber.

As the ammonite grew, it added new and larger chambers to the opened end of the shell. A thin living tube called a siphuncle passed through the septa, extending from the body to the empty shell chambers.

This allowed the ammonite to empty water out of the shell chambers by hyperosmotic active transport process. This process controlled the buoyancy of the ammonite's shell.

First Photo: Hoplites Bennettiana from near Troyes, France. Collection de Christophe Marot

Second Photo: Top: Hoplites maritimus from Sussex, UK. Bottom: Hoplites rudis from near Troyes, France. Collection of Mark O'Dell

Third Photo: Hoplites sp. from the Early Cretaceous of Dorset, UK. Natural Selection Fossils

Fourth Photo: Hoplites dentalus from Albian deposits near Troyes, France. Collection of Stéphane Rolland.

Wright, C. W. (1996). Treatise on Invertebrate Paleontology, Part L, Mollusca 4: Cretaceous Ammonoidea (with contributions by JH Calloman (sic) and MK Howarth). Geological Survey of America and University of Kansas, Boulder, Colorado, and Lawrence, Kansas, 362.

Amédro, F., Matrion, B., Magniez-Jannin, F., & Touch, R. (2014). La limite Albien inférieur-Albien moyen dans l’Albien type de l’Aube (France): ammonites, foraminifères, séquences. Revue de Paléobiologie, 33(1), 159-279.

AMMONITE OF THE RHÔNE

An exquisite specimen of the delicately ridged ammonite, Porpoceras verticosum, from Middle Toarcian outcrops adjacent the Rhône in southeastern France.

Porpoceras (Buchman, 1911) is a genus of ammonite that lived during the early and middle Toarcian stage of the Early Jurassic. We see members of this genus from the uppermost part of Serpentinum Zone to Variabilis Subzone. These beauties are found in Europe, Asia, North America and South America.

Ammonites belonging to this genus have evolute shells, with compressed to depressed whorl section. Flanks were slightly convex and venter has been low. The whorl section is sub-rectangular. 

The rib is pronounced and somewhat fibulate on the inner whorls — just wee nodes here — and tuberculate to spined on the ventrolateral shoulder. It differs from Peronoceras by not having a compressed whorl section and regular nodes or fibulation. Catacoeloceras is also similar, but it has regular ventrolateral tubercules and is missing the classic nodes or fibulation of his cousins.

This specimen hails from southern France near the Rhône, one of the major rivers of Europe. It has twice the average water level of the Loire and is fed by the Rhône Glacier in the Swiss Alps at the far eastern end of the Swiss canton of Valais then passes through Lake Geneva before running through southeastern France. This 10 cm specimen was prepared by the supremely talented José Juárez Ruiz

OIL IN WATER BEAUTY: FOSSILS OF FOLKSTONE

Sheer beauty — a beautiful Euhoplites ammonite from Folkstone, UK. I've been really enjoying looking at all oil-in-water colouring and chunkiness of these ammonites.

Euhoplites is an extinct ammonoid cephalopod from the Lower Cretaceous, characterized by strongly ribbed, more or less evolute, compressed to inflated shells with flat or concave ribs, typically with a deep narrow groove running down the middle.

In some, ribs seem to zigzag between umbilical tubercles and parallel ventrolateral clavi. In others, the ribs are flexious and curve forward from the umbilical shoulder and lap onto either side of the venter.

Its shell is covered in the lovely lumps and bumps we associate with the genus. The function of these adornments are unknown. I wonder if they gave them greater strength to go deeper into the ocean to hunt for food. 

They look to have been a source of hydrodynamic drag, likely preventing Euhoplites from swimming at speed. Studying them may give some insight into the lifestyle of this ancient marine predator. Euhoplites had shells ranging in size up to a 5-6cm. 

We find them in Lower Cretaceous, middle to upper Albian age strata. Euhoplites has been found in Middle and Upper Albian beds in France where it is associated respectively with Hoplites and Anahoplites, and Pleurohoplites, Puzosia, and Desmoceras; in the Middle Albian of Brazil with Anahoplites and Turrilites; and in the Cenomanian of Texas.

This species is the most common ammonite from the Folkstone Fossil Beds in southeastern England where a variety of species are found, including this 37mm beauty from the collections of José Juárez Ruiz.

QUENSTEDTOCERAS WITH PATHOLOGY

What you are seeing here is a protuberance extruding from the venter of Quenstedtoceras cf. leachi (Sowerby). It is a pathology in the shell from hosting immature bivalves that shared the seas with these Middle Jurassic, Upper Callovian, Lamberti zone fauna from the Volga River basin. The collecting site is the now inactive Dubki commercial clay quarry and brickyard near Saratov, Russia. 

The site has produced thousands of ammonite specimens. A good 1,100 of those ended up at the Black Hills Institute of Geological Research in Hill City, South Dakota. 

Roughly 1,000 of those are Quenstedtoceras (Lamberticeras) lamberti and the other 100 are a mix of other species found in the same zone. These included Eboraciceras, Peltoceras, Kosmoceras, Grossouvria, Proriceras, Cadoceras and Rursiceras. 

What is especially interesting is the volume of specimens — 167 Quenstedtoceras (Lamberticeras) lamberti and 89 other species in the Black Hills collection — with healed predation injuries. It seems Quenstedtoceras (Lamberticeras) lamberti are the most common specimens found here and so not surprisingly the most common species found injured. Of the 1,000, 655 of the Quenstedtoceras (Lamberticeras) lamberti displayed some sort of deformation or growth on the shell or had grown in a tilted manner. 

Again, some of the Q. lamberti had small depressions in the centre likely due to a healed bite and hosting infestations of the immature bivalve Placunopsis and some Ostrea. 

The bivalves thrived on their accommodating hosts and the ammonites carried on, growing their shells right up and over their bivalve guests. This relationship led to some weird and deformities of their shells. They grow in, around, up and over nearly every surface of the shell and seem to have lived out their lives there. It must have gotten a bit unworkable for the ammonites, their shells becoming warped and unevenly weighted. Over time, both the flourishing bivalves and the ammonite shells growing up and over them produced some of the most interesting pathology specimens I have ever seen.    

In the photo here from Emil Black, you can see some of the distorted shapes of Quenstedtoceras sp. Look closely and you see a trochospiral or flattened appearance on one side while they are rounded on the other. 

All of these beauties hail from the Dubki Quarry near Saratov, Russia. The ammonites were collected in marl or clay used in brick making. The clay particles suggest a calm, deep marine environment. One of the lovely features of the preservation here is the amount of pyrite filling and replacement. It looks like these ammonites were buried in an oxygen-deficient environment. 

The ammonites were likely living higher in the water column, well above the oxygen-poor bottom. An isotopic study would be interesting to prove this hypothesis. There's certainly enough of these ammonites that have been recovered to make that possible. It's estimated that over a thousand specimens have been recovered from the site but that number is likely much higher. But these are not complete specimens. We mostly find the phragmocones and partial body chambers. Given the numbers, this may be a site documenting a mass spawning death over several years or generations.

If you fancy a read on all things cephie, consider picking up a copy of Cephalopods Present and Past: New Insights and Fresh Perspectives edited by Neil Landman and Richard Davis. Figure 16.2 is from page 348 of that publication and shows the hosting predation quite well. 

Photos: Courtesy of the deeply awesome Emil Black. These are in his personal collection that I hope to see in person one day. 

It was his sharing of the top photo and the strange anomaly that had me explore more about the fossils from Dubki and the weird and wonderful hosting relationship between ammonites and bivalves. Thank you, my friend!