Showing posts with label paleontology. Show all posts
Showing posts with label paleontology. Show all posts

Monday 18 March 2024

JAPANESE CORKSCREW AMMONITE: HYPHANTOCERAS ORIENTALE

A stunning example of the heteromorph ammonite, Hyphantoceras orientale macroconch. This beauty corresponds to 'Morphotype C' from Aiba (2017). 

The specimen is a handful at 136 mm and was lovingly prepared by the hand holding it, that of the talented José Juárez Ruiz.

This an adult specimen (not the juvenile stage) from Upper Santonian outcrops near Ashibetsu, Hokkaido, Japan.

Aiba published on a possible phylogenetic relationship of two species of Hyphantoceras (Ammonoidea, Nostoceratidae) earlier this year, proposing that a phylogenetic relationship may exist based on newly found specimens with precise stratigraphic occurrences in the Kotanbetsu and Obira areas, northwestern Hokkaido.

Two closely related species, Hyphantoceras transitorium and H. orientale, were recognized in the examined specimens from the Kotanbetsu and Obira areas. Specimens of H. transitorium show wide intraspecific variation in the whorl shape. The stratigraphic occurrences of the two species indicate that they occur successively in the Santonian–lowermost Campanian, without stratigraphic overlapping. 

The similarity of their shell surface ornamentations and the stratigraphic relationships possibly suggest that H.orientale was derived from H. transitorium. The presumed lineage is likely indigenous to the northwestern Pacific realm in the Santonian–earliest Campanian. Hyphantoceras venustum and H. heteromorphum might stand outside a H. transitorium–H. orientale lineage, judging from differences of their shell surface ornamentation.

Aiba, Daisuke. (2019). A Possible Phylogenetic Relationship of Two Species of Hyphantoceras (Ammonoidea, Nostoceratidae) in the Cretaceous Yezo Group, Northern Japan. Paleontological Research. 23. 65-80. 10.2517/2018PR010.

Saturday 16 March 2024

DRIFTWOOD CANYON FOSSIL BEDS / KUNGAX

Puffbird similar to Fossil Birds found at Driftwood Canyon 
Driftwood Canyon Provincial Park 

Driftwood Canyon Provincial Park covers 23 hectares of the Bulkley River Valley, on the east side of Driftwood Creek, a tributary of the Bulkley River, 10 km northeast of the town of Smithers in northern British Columbia. 

Wet'suwet'en First Nation

The parklands are part of the asserted traditional territory of the Wet'suwet'en First Nation which includes lands around the Bulkley River, Burns Lake, Broman Lake, and François Lake in the northwestern Central Interior of British Columbia. 

The Wetʼsuwetʼen are part of the Dakelh or Carrier First Nation, and in combination with the Babine First Nation are referred to as the Western Carrier. They speak Witsuwitʼen, a dialect of the Babine-Witsuwitʼen language which, like its sister language Carrier, is a member of the Athabaskan family.

Their oral history or kungax recounts a time when their ancestral village, Dizkle or Dzilke, once stood upstream from the Bulkley Canyon. This cluster of cedar houses on both sides of the river was said to be abandoned because of an omen of impending disaster. The exact location of the village has been lost but their stories live on. 

The neighbouring Gitxsan, collectively the People of Smooth Waters—the Gilseyhu Big Frog Clan, the Laksilyu Small Frog Clan, the Tsayu Beaver Clan, the Gitdumden Wolf and Bear Clan and the Laksamshu Fireweed and Owl Clan—each phratry or kinship group calling the Lax Yip home—33,000 km2 of land and water in northwestern ​British Columbia along the waters of the Skeena River and its tributaries—have a similar tale—though the village in their versions is referred to as Dimlahamid or Temlahan depending on which family house is sharing it and where they are located as dialects differ. 

Gitksan speak Sim'algaxthe real or true language. Within the Gitxsan communities there are two slightly different dialects. The Gyeets (Downriver) dialect spoken in Gijigyukwhla (Gitsegukla), Gitwangax, and Gitanyow—and the Gigeenix (Upriver) dialect is spoken in Ansbayaxw (Kispiox), Sik-E-Dakh and Gitanmaax.

Driftwood Canyon Fossil Beds

Driftwood Canyon's Fossil Beds record life in the earlier portion of the Eocene when British Columbia — and indeed our world — was much warmer than it is today. This site was discovered in the beginning of the 20th century and is now recognized as containing significant fossil material. 

The fossils found here—and their superb preservation—provide a fascinating opportunity to understand the area’s evolutionary processes of both geology and biology over the past fifty million years or so. The fossils themselves are 51.7 million years old and look remarkably like many of the species we recognize today. 

The park that contains these beautiful fossils is fifty-seven years old. It was created in 1967 by the generosity of the late Gordon Harvey (1913–1976). He donated the land to protect fossil resources that he truly loved and wanted to see preserved. The fossil beds are on the east side of Driftwood Creek. 

Metasequoia, the Dawn Redwood
We have found plant, fish and insect fossil here including Metasequoia, the Dawn Redwood, alder, fossil salmon, wasps, water striders and vertebrate material. Bird feathers are infrequently collected from the shales; however, two bird body fossils have been found here.

In 1968, a bird body fossil was collected in the Eocene shales of the Ootsa Lake Group in Driftwood Canyon Provincial Park by Pat Petley of Kamloops. 

Pat donated the specimen in 2000 to the Thompson Rivers University (TRU) palaeontology collections. This fossil bird specimen is tentatively identified as the puffbird, Piciformes bucconidae, of the genus Primobucco.

Primobucco is an extinct genus of bird placed in its own family, Primobucconidae. The type species, Primobucco mcgrewi, lived during the Lower Eocene of North America. It was initially described by American paleo-ornithologist Pierce Brodkorb in 1970, from a fossil right-wing, and thought to be an early puffbird. However, the discovery of a further 12 fossils in 2010 indicate that it is instead an early type of roller.

Related fossils from the European Messel deposits have been assigned to the two species P. perneri and P. frugilegus. Two specimens of P. frugilegus have been found with seeds in the area of their digestive tract, which suggests that these birds were more omnivorous than the exclusively predaceous modern rollers. The Driftwood specimen has never been thoroughly studied. If there is a grad student out there looking for a worthy thesis, head on down to the Thompson Rivers University where you'll find the specimen on display.

Another fossil bird, complete with feathers, was collected at Driftwood Canyon in 1970, This one was found by Margret and Albrecht Klöckner who were travelling from Germany. Theirs is a well-travelled specimen, having visited many sites in BC as they toured around, then to Germany and finally back to British Columbia when it was repatriated and donated to the Royal British Columbia Museum in Victoria. 

I am not sure if it is still on display or back in collections, but it was lovingly displayed back in 2008. There is a new grad student, Alexis, looking at Eocene bird feathers down at the RBCM, so perhaps it is once again doing the rounds. 

This second bird fossil is of a long-legged water bird and has been tentatively identified by Dr. Gareth Dyke of the University of Southampton as possibly from the order Charadriiformes, a diverse order of small to medium-ish water birds that include 350 species of gulls, plovers, sandpipers, terns, snipes, and waders. Hopefully, we'll hear more on this find in the future.

A Tapir showing off his prehensile nose trunk
Tapirs and Tiny Hedgehogs

The outcrops at Driftwood Canyon are also special because they record a record of some of the first fossil mammals ever to be found in British Columbia at this pivotal point in time. 

Wee proto-hedgehogs smaller than your thumb lived in the undergrowth of that fossil flora. They shared the forest floor with an extinct tapir-like herbivore in the genus Heptodon that looked remarkably similar to his modern, extant cousins (there is a rather cheeky fellow shown here so you get the idea) but lacked their pronounced snout (proboscis). I am guessing that omission made him the more fetching of his lineage.

In both cases, it was a fossilized jaw bone that was recovered from the mud, silt and volcanic ash outcrops in this ancient lakebed site. And these two cuties are significant— they are the very first fossil mammals we've ever found from the early Eocene south of the Arctic.

How can we be sure of the timing? The fossil outcrops here are found within an ancient lakebed. Volcanic eruptions 51 million years ago put loads of fine dust into the air that settled then sank to the bottom of the lake, preserving the specimens that found their way here — leaves, insects, birds, mammals.

 As well as turning the lake into a fossil making machine—water, ash, loads of steady sediment to cover specimens and stave off predation—the volcanic ash contains the very chemically inert—resistant to mechanical weathering—mineral zircon which we can date with uranium/lead (U/Pb). 

The U/Pb isotopic dating technique is wonderfully accurate and mighty helpful in dating geologic events from volcanic eruptions, continental movements to mass extinctions. This means we know exactly when these lovelies were fossilized and, in turn, their significance.

Know Before You Go

If you fancy a visit to Driftwood Canyon Park, the park is accessible from Driftwood Road from Provincial Highway 16. You are welcome to view and photograph the fossils found here but collecting is strictly forbidden. 

Driftwood Canyon is recognized as one of the world’s most significant fossil beds. It provides park users with a fascinating opportunity to understand the area’s evolutionary processes of both geology and biology. The day-use area is open from May 15 to September 2. There is a short, wheelchair-accessible interpretative trail that leads from the parking are to the fossil beds. Pets are welcome on leash. Signs along the trail provide information on fossils and local history. 

Below a cliff face at the end of the trail is a viewing area that has interpretive information and viewing area overlooking Driftwood Creek.

This park proudly operated by Mark and Anais Drydyk
Email: kermodeparks@gmail.com / Tel: 1 250 877-1482 or 1 250 877-1782

Palaeo Coordinates: Latitude: 50° 51' 59" N / Longitude: 116° 27' 37" W
Lat/Long (dec): 50.86665,-116.46042 / GUID: d3a6bd3e-68d6-42cf-9b2c-d20a30576988

Driftwood Canyon Provincial Park Brochure: 
https://bcparks.ca/explore/parkpgs/driftwood_cyn/driftwood-canyon-brochure.pdf?v=1638723136455


Sunday 3 March 2024

LATE HETTANGIAN FOSSIL FAUNA FROM THE TASEKO LAKES: BRITISH COLUMBIA

The late Hettangian ammonite fauna from Taseko Lakes is diverse and relatively well‐preserved. Over three field seasons, thirty-five taxa from the Mineralense and Rursicostatum zones were studied and three new species discovered and named: Fergusonites hendersonae, Eolytoceras constrictum and Pseudaetomoceras victoriense

This material is very important as it greatly expands our understanding of the fauna and ranges of ammonites currently included in the North American regional ammonite zonation. 

I had the very great honour of having the fellow below, Fergusonites hendersonae, a new species of nektonic carnivorous ammonite, named after me by palaeontologist Louse Longridge from the University of British Columbia. 

I'd met Louise as an undergrad and was pleased as punch to hear that she would be continuing the research by Dr. Howard Tipper, the authority on this area of the Chilcotins and Haida Gwaii — which he dearly loved. 

"Tip" was a renowned Jurassic ammonite palaeontologist and an excellent regional mapper who mapped large areas of the Cordillera. He made significant contributions to Jurassic paleobiogeography and taxonomy in collaboration with Dr. Paul Smith, Head of Earth and Ocean Science at the University of British Columbia. 

Tip’s regional mapping within BC has withstood the test of time and for many areas became the regions' base maps for future studies. The scope of Tip’s understanding of Cordilleran geology and Jurassic palaeontology will likely never be matched. He passed away on April 21, 2005. His humour, knowledge and leadership will be sorely missed. 

Fergusonites hendersonae
Before he left us, he shared that knowledge with many of whom who would help to secure his legacy for future generations. We did several trips over the years up to the Taseko Lake area of the Rockies joined by many wonderful researchers from Vancouver Island Palaeontological Society and Vancouver Paleontological Society, as well as the University of British Columbia. 

Both Dan Bowen and John Fam were instrumental in planning those expeditions and each of them benefited greatly from the knowledge of Dr. Howard Tipper. 

If not for Tipper's early work in the region, our shared understanding and much of what was accomplished in his last years and after his passing would not have been possible. 

Over the course of three field seasons, we endured elevation sickness, rain, snow, grizzly bears and very chilly nights  — we were sleeping right next to a glacier at one point — but were rewarded by the enthusiastic crew, helicopter rides — which really cut down the hiking time — excellent specimens including three new species of ammonites, along with a high-spired gastropod and lobster claw that have yet to be written up. This area of the world is wonderful to hike and explore — stunningly beautiful country. We were also blessed with access as the area is closed to all fossil collecting except with a permit.

This fauna understanding helps us to understand the correlations between different areas: (1) the Mineralense and Rursicostatum zones are present in Taseko Lakes and can be readily correlated with contemporaneous strata elsewhere in North America; (2) the Mineralense and Rursicostatum zones of North America are broadly equivalent to the Canadensis Zone and probably the Arcuatum horizon of the South American succession; (3) broad correlations are possible with middle–late Hettangian and earliest Sinemurian taxa in New Zealand; (4) the Mineralense and Rursicostatum zones are broadly equivalent to the circum‐Mediterranean Marmoreum Zone; (5) the Mineralense Zone and the lower to middle portion of the Rursicostatum Zone are probably equivalent to the Complanata Subzone whereas the upper portion of the Rursicostatum Zone may equate to the Depressa Subzone of the north‐west European succession.

Taseko Lake Area, BC
The Taseko Lakes area has yielded the best preserved and most diverse collection of late Hettangian ammonites yet discovered in British Columbia (BC). Early studies of the fauna were undertaken by Frebold (1951, 1967). At that time, eastern Pacific ammonite faunas were poorly understood and species were frequently shoehorned into established north‐west European taxa. 

Since then, knowledge of eastern Pacific Hettangian ammonite faunas has improved considerably. 

Detailed systematic studies have been completed on faunas from localities in other areas of BC, Alberta, Alaska, Oregon, Nevada, Mexico and South America (e.g. Guex 1980, 1995; Imlay 1981; Hillebrandt 1981, 1988, 1990, 1994, 2000a–d; Smith and Tipper 1986; Riccardi et al. 1991; Jakobs and Pálfy 1994; Pálfy et al. 1994, 1999; Taylor 1998; Hall et al. 2000; Taylor and Guex 2002; Hall and Pitaru 2004). 

These studies have demonstrated that Early Jurassic eastern Pacific ammonites had strong Tethyan affinities as well as a high degree of endemism (Guex 1980, 1995; Taylor et al. 1984; Smith et al. 1988; Jakobs et al. 1994; Pálfy et al. 1994). Frebold’s early studies were also hampered because they were based on small collections, which limited understanding of the diversity of the fauna and variation within populations. However, recent mapping has greatly improved our understanding of the geology of Taseko Lakes (Schiarizza et al. 1997; Smith et al. 1998; Umhoefer and Tipper 1998) and encouraged further collecting that has dramatically increased the size of the sample.

A study of the ammonite fauna from Taseko Lakes is of interest for several reasons. The data are important for increasing the precision of the late Hettangian portion of the North American Zonation. 

Owing to the principally Tethyan or endemic nature of Early Jurassic ammonites in the eastern Pacific, a separate zonation for the Hettangian and Sinemurian of the Western Cordillera of North America has been established by Taylor et al. (2001). Except for information available from the early studies by Frebold (1951, 1967), the only Taseko Lakes taxa included in the North American Zonation of Taylor et al. (2001) were species of Angulaticeras studied by Smith and Tipper (2000). 

Since then, Longridge et al. (2006) made significant changes to the zonation of the late Hettangian and early Sinemurian based on a detailed study of the Badouxia fauna from Taseko Lakes (Text‐fig. 2). An additional taxonomic study was recently completed on the late Hettangian ammonite Sunrisites (Longridge et al. 2008) and this information has not yet been included within the zonation. 

Hettangian Zonation
The systematics of the remaining ammonite fauna from Taseko Lakes are presented here. A comprehensive study of this material is important because the exceptional quality and diversity of the fauna provide important data for updating the North American Zonation, making it more comprehensive and more widely applicable, especially in Canada.

The Taseko Lakes fauna can improve Hettangian correlations within North America as well as between North America and the rest of the world. 

North‐west European ammonite successions (e.g. Dean et al. 1961; Mouterde and Corna 1997; Page 2003) are considered the primary standard for Early Jurassic biochronology (Callomon 1984). 

In north‐west Europe, the turnover from schlotheimiid dominated faunas in the late Hettangian to arietitid dominated faunas in the early Sinemurian was sharp (e.g. Dean et al. 1961; Bloos 1994; Bloos and Page 2002). In other areas, by contrast, these faunas were not so mutually exclusive and the transition was much more gradual. 

This makes correlations between north‐west Europe and other areas difficult (e.g. Bloos 1994; Bloos and Page 2000, 2002). Correlations are further impeded by endemism and provincialism. 

The Taseko Lakes fauna addresses these problems because it contains many taxa that are common throughout the eastern Pacific as well as several cosmopolitan taxa that make intercontinental correlation possible. Correlation between North America and other areas is of particular significance in that the interbedded volcanic and fossiliferous marine rocks in North America permit the calibration of geochronological and biochronological time scales (Pálfy et al. 1999, 2000). 

This correlation between the late Hettangian fauna in the Taseko Lakes area and contemporaneous faunas in other areas of North America, South America, New Zealand, western and eastern Tethys, and north‐west Europe is of particular interest to me — especially the correlation of the faunal sequences of Nevada, USA. 

Reference: PaleoDB 157367 M. Clapham GSC C-208992, Section A 09, Castle Pass Angulata - Jurassic 1 - Canada, Longridge et al. (2008)

L. M. Longridge, P. L. Smith, and H. W. Tipper. 2008. Late Hettangian (Early Jurassic) ammonites from Taseko Lakes, British Columbia, Canada. Palaeontology 51:367-404

PaleoDB taxon number: 297415; Cephalopoda - Ammonoidea - Juraphyllitidae; Fergusonites hendersonae Longridge et al. 2008 (ammonite); Average measurements (in mm): shell width 9.88, shell diameter 28.2; Age range: 201.6 to 196.5 Ma. Locality info: British Columbia, Canada (51.1° N, 123.0° W: paleo coordinates 22.1° N, 66.1° W)

Photo One: Hettangian Ammonites and Gastropods, Taseko Lakes. Photo Two: Fergusonites hendersonae, a Late Hettangian (Early Jurassic) ammonite from the Taseko Lakes area of British Columbia, Canadian Rockies, named by Dr. Louise Longridge after Heidi Henderson, Chair, Vancouver Paleontological Society who collected and subsequently donated many Hettangian specimens from Taseko Lakes to the GSC collections. Holotype. GSC 127423 from the Rursicostatum Zone, Castle Pass section A, level 06, Taseko Lakes.

Map: Localities of sections and isolated outcrops bearing late Hettangian ammonites in the Taseko Lakes map area. Figure Two: Zonation for the Hettangian showing correlation of North American zones with South America, north‐west Europe, western Tethys (circum‐Mediterranean), eastern Tethys and New Zealand. Only approximate correlations are implied. 

Friday 16 February 2024

UPTHRUSTING PLATES: WASHINGTON GEOLOGY

Two hundred million years ago, Washington was two large islands, bits of the continent on the move westward, eventually bumping up against the North American continent and calling it home. The shifting continues, subtly changing the landscape like a breath. We only notice when pockets of resistance manifest as earthquakes, some newsworthy, some all but unnoticed. For now, the more extreme movement has subsided laterally and continues vertically, pushing California towards the North Pole. Hello Baja-BC.

The upthrusting of plates move our mountain ranges skyward – the path of least resistance. And it is this dynamic movement that's created the landscape we see today.

The 3,000 meters of the stratigraphic section of the Chuckanut Formation along Chuckanut Drive span an age range of just a few million years. The lower part is late Paleocene with a radiometric age of around 56 million years. The upper part of the section is early Eocene. The fossils found here lived and died very close to where they are now but in a much warmer, wetter, swampy setting. The exposures of the Chuckanut Formation were once part of a vast river delta; imagine, if you will, the bayou country of the Lower Mississippi. The siltstones, sandstones, mudstones and conglomerates of this formation were laid down during a time of luxuriant plant growth in the subtropical flood plain that covered much of the Pacific Northwest.

This ancient wetland provided ideal conditions to preserve the many trees, shrubs and plants that thrived here giving us a lot of information about climate, temperature, the water cycle and humidity of the region. The Chuckanut flora is made up predominantly of plants whose modern relatives live in tropical areas such as Mexico and Central America. While less abundant, evidence of the animals that called this ancient swamp home are also found here. Rare bird, reptile, and mammal tracks have been immortalized in the outcrops of the Chuckanut Formation.

Sumas Eocene Shorebird Trackway
Tracks of a type of archaic mammal of the Orders Pantodonta or Dinocerata (blunt foot herbivores), footprints from a small shorebird, and tracks from an early equid or webbed bird track give evidence to the vertebrates that inhabited the swamps, lakes and riverways of the Pacific Northwest 50 million years ago.

Fossil mammals and bird trackways from Washington State have caused great excitement over the past few years. Many new trackways have been discovered since the 2009 slides near Sumas. George Mustoe and team collected these important finds, bringing them to the Burke Museum in Washington State to study and make available for display.

The movement of these vertebrates was captured in the soft mud on the banks of an ancient river, one of the only depositional environments favourable for track preservation. The terrestrial paleontological record of Washington State at sites like Chuckanut and Racehorse Creek (U-Pb 53 Ma.) is primarily made up of plant material with some wonderfully enticing mammal, shorebird (seen here) and large Diatryma bird tracks on rare occasions.

Tuesday 13 February 2024

JUVENILE HAMITES SUBROTUNDUS

A tremendously delicate juvenile Hamites subrotundus (J. Sowerby 1814) from Upper Albian outcrops in Mallorca, the largest of the three Balearic Islands in the Mediterranean at more than 3,600 square kilometers. 

Mallorca has been home to various inhabitants for thousands of years. Sitting some 200 kilometers off Spain’s southeastern, it is a idyllic setting for exploring the rich human and geologic history of this part of the world.

The island is made up of dolomite and limestone from a huge expanse of time, the Mesozoic and Cenozoic—170 million to 10 million years ago— and bookended by two parallel mountain chains top and bottom on its southern and northern coasts. 

As you walk the mountain passes northwest to northeast, you stroll across Miocene deposits 20 million to 13 million years ago that speak of the time in our Earth's recent past when part of the African continent collided with Europe. 

It is famous for its limestone mountains and Roman and Moorish remains. As you can see here, it is also home to some rather nice fossils including this specimen of Hamites subrotundus.

While H. subrotundus is generally a Middle Albian species, this specimen was found in the lower part of Upper Albian in the Cristatum zone by José Juárez Ruiz. José had to piece this lovely together from seven fragments. His labour of love was worth the effort. The final piece is sheer perfection and a beautiful specimen approximately 2.5 cm long.

Mallorca and the other Balearic Islands are geologically an extension of the Baetic Cordillera mountain chain of western Andalusia that extends to Murcia and Valencia. 

They are made up of sediments deposited in the Tethys Sea during the Mesozoic.

Exploring the islands, you can collect from deposits from the Triassic, Cretaceous, Jurassic, and Neogene periods. 

The limestone outcrops contain lovely examples of foraminifers—mainly the species Globigerina.

We also see lovely examples of Hamites (Hamites) subrotundus in the Euhoplites loricatus zone; Euhoplites meandrinus subzone from the Middle Albian (Lower Gault) of Folkestone, Kent, UK. 

Photo, preparation and in the collection of the deeply awesome José Juárez Ruiz. Wright C. W. 1996. Treatise on Invertebrate Paleontology (Part L Mollusca 4 Revised) Volume 4: Cretaceous Ammonoidea

Monday 12 February 2024

FOSSILS, LIMESTONE & SALT: HALLSTATT

Hallstatt Salt Mines, Austria / Permian Salt Diapir
The Hallstatt Limestone is the world's richest Triassic ammonite unit, yielding specimens of more than 500 ammonite species.

Along with diversified cephalopod fauna  — orthoceratids, nautiloids, ammonoids — we also see gastropods, bivalves, especially the late Triassic pteriid bivalve Halobia (the halobiids), brachiopods, crinoids and a few corals. We also see a lovely selection of microfauna represented. 

For microfauna, we see conodonts, foraminifera, sponge spicules, radiolaria, floating crinoids and holothurian sclerites —  polyp-like, soft-bodied invertebrate echinozoans often referred to as sea cucumbers because of their similarities in size, elongate shape, and tough skin over a soft interior. 

Franz von Hauer’s exhaustive 1846 tome describing Hallstatt ammonites inspired renowned Austrian geologist Eduard Suess’s detailed study of the area’s Mesozoic history. That work was instrumental in Suess being the first person to recognize the former existence of the Tethys Sea, which he named in 1893 after the sister of Oceanus, the Greek god of the ocean. As part of the Northern Limestone Alps, the Dachstein rock mass, or Hoher Dachstein, is one of the large karstic mountains of Austria and the second-highest mountain in the Northern Limestone Alps. It borders Upper Austria and Styria in central Austria and is the highest point in each of those states.

Parts of the massif also lie in the state of Salzburg, leading to the mountain being referred to as the Drei-Länder-Berg or three-state mountain. Seen from the north, the Dachstein massif is dominated by the glaciers with the rocky summits rising beyond them. By contrast, to the south, the mountain drops almost vertically to the valley floor. The karst limestones and dolomites were deposited in our Mesozoic seas. The geology of the Dachstein massif is dominated by the Dachstein-Kalk Formation — the Dachstein limestone — which dates back to the Triassic.

Hallstatt and the Hallstatt Sea, Austria
There were several phases of mountain building in this part of the world pushing the limestone deposits 3,000 metres above current sea level. The rock strata were originally deposited horizontally, then shifted, broken up and reshaped by the erosive forces of ice ages and erosion.

The Hallstatt mine exploits a Permian salt diapir that makes up some of this area’s oldest rock. 

The salt accumulated by evaporation in the newly opened, and hence shallow, Hallstatt-Meliata Ocean. This was one of several small ocean basins that formed in what is now Europe during the late Paleozoic and early Mesozoic when the world’s landmasses were welded together to form the supercontinent Pangea. 

Pangea was shaped like a crescent moon that cradled the famous Tethys Sea. Subduction of Tethyian oceanic crust caused several slivers of continental crust to separate from Pangea, forming new “back-arc basins” (small oceans formed by rifting that is associated with nearby subduction) between the supercontinent and the newly rifted ribbon continents.

The Hallstatt-Meliata Ocean was one such back-arc basin. As it continued to expand and deepen during the Triassic, evaporation ceased and reefs flourished; thick limestone deposits accumulated atop the salt. When the Hallstatt-Meliata Ocean closed in the Late Jurassic, the compression squeezed the low-density salt into a diapir that rose buoyantly, injecting itself into the Triassic limestones above.

The Hallstatt salt diapir and its overlying limestone cap came to rest in their present position in the northern Austrian Alps when they were shoved northward as nappes (thrust sheets) during two separate collision events, one in the Cretaceous and one in the Eocene, that created the modern Alps. It is from the Hallstatt salt diapir that Hallstatt, like so many cities and towns, gets its name.

Deposits of rock salt or halite, the mineral name of sodium chloride with the chemical formula of NaCl, are found and mined around the globe. These deposits mark the dried remains of ancient oceans and seas. Names of rivers, towns and cities in Europe — Salzburg, Halle, Hallstatt, Hallein, La Salle, Moselle — all pay homage to their connection to halite and salt production. The Greek word for salt is hals and the Latin is sal. The Turkish name for salt is Tuz, which we see in the naming of Tuzla, a salt-producing region of northeastern Bosnia-Herzegovina and in the names of towns that dot the coast of Turkey where it meets the Black Sea. Hallstatt with its salt diapir is no exception.

The salt-named town of Hallstatt sits on the shores of the idyllic Hallstätter Sea at the base of the Dachstein massif. Visiting it today, you experience a quaint traditional fishing village built in the typical upper Austrian style. Tourism drives the economy as much as salt as this area of the world is picture-perfect from every angle.

Space is at a minimum in the town. For centuries, every ten years the local cemetery exhumes the bones of those buried there and moves them to an ossuary to make room for new burials. The Hallstatt Ossuary is called Karner, Charnel House, or simply Beinhaus (Bone House). Karners are places of secondary burials. They were once common in the Eastern Alps, but that custom has largely disappeared.

Hallstatt Beinhaus Ossuary, Hallstatt, Austria
A collection of over 700 elaborately decorated skulls rest inside the ossuary. They are lined up on rows of wooden shelves that grace the walls of the chapel. Another 500 undecorated skulls, bare and without any kind of adornment, are stacked in the corners.

Each is inscribed and attached to a record with the deceased's name, profession and date of death. The Bone House is located in a chapel in the basement of the Church of Saint Michael. The church dates from the 12th century CE. 

Decorating the skulls was traditionally the job of the local gravedigger and an honour granted to very few. At the family's request, garlands of flowers were painted on the skulls of deceased as decorative crowns if they were female. The skulls of men and boys were painted wreaths of oak or ivy.

Every building in Hallstatt looks out over the Hallstätter Sea. This beautiful mountain lake considered one of the finest of Austria's Salzkammergut region. It lies at the northern foot of the Dachstein mountain range, sitting eight-and-a-half kilometres long and two kilometres wide. The shoreline is dotted by the villages of  Obertraun, Steeg, and Hallstatt.

The region is habitat to a variety of diverse flora and fauna, including many rare species such as native orchids, in the wetlands and moors in the south and north.

Linked by road to the cities of Salzburg and Graz, Hallstatt and its lake were declared one of the World Heritage sites in Austria in 1997 and included in the Hallstatt-Dachstein Salzkammergut Alpine UNESCO World Heritage Site. The little market village of Hallstatt takes its name from the local salt mine.

Hallstatt, Salzkammergut region, Austria
The town is a popular tourist destination with its quaint shops and terraced cafes. In the centre of town, the 19th-century Evangelical Church of Hallstatt with its tall, slender spire is a lakeside landmark. You can see it here in the photo on the left.

Above the town are the Hallstatt Salt mines located within the 1,030-meter-tall Salzburg Salt Mountain. They are accessible by cable car or a three-minute journey aboard the funicular railway. There is also a wonderful Subterranean Salt Lake.

In 1734, there was a corpse found here preserved in salt. The fellow became known as the Man in Salt. Though no archaeological analysis was performed at the time — the mummy was respectfully reburied in the Hallstatt cemetery — based on descriptions in the mine records, archaeologists suspect the miner lived during the Iron Age. This Old Father, Senos ph₂tḗr, 'ɸatīr 'father' may have been a local farmer, metal-worker, or both and chatted with his friends and family in Celtic or Proto-Celtic.

Salt mining in the area dates back to the Neolithic period, from the 8th to 5th Centuries BC. This is around the time that Roman legions were withdrawing from Britain and the Goths sacked Rome. In Austria, agricultural settlements were dotting the landscape and the alpine regions were being explored and settled for their easy access to valuable salt, chert and other raw materials.

The salt-rich mountains of Salzkammergut and the upland valley above Hallstatt were attractive for this reason. The area was once home to the Hallstatt culture, an archaeological group linked to Proto-Celtic and early Celtic people of the Early Iron Age in Europe, c.800–450 BC.
Bronze Age vessel with cow and calf

In the 19th century, a burial site was discovered with 2,000 individuals, many of them buried with Bronze Age artefacts of amber and ivory.

It was this find that helped lend the name Hallstatt to this epoch of human history. The Late Iron Age, between around 800 and 400 BC, became known as the Hallstatt Period.

For its rich history, natural beauty and breathtaking mountainous geology, Hallstatt is a truly irresistible corner of the world.

Salzbergstraße 1, 4830 Hallstatt.  https://www.salzwelten.at/en/home/

Photo: Bronze vessel with cow and calf, Hallstatt by Alice Schumacher - Naturhistorisches Museum Wien - A. Kern – K. Kowarik – A. W. Rausch – H. Reschreiter, Salz-Reich. 7000 Jahre Hallstatt, VPA 2 (Wien, 2008) Seite 133 Abbildung 6. Hallstatt Village & Ossuary Photos: P. McClure Photography ca. 2015.

Bernoulli D, Jenkyns HC (1974) Alpine, Mediterranean, and Central Atlantic Mesozoic facies in relation to the early evolution of the Tethys. Soc Econ Paleont Mineral Spec Publ 19:129–160

Bernoulli D, Jenkyns H (2009) Ancient oceans and continental margins of the Alpine-Mediterranean Tethys: deciphering clues from Mesozoic pelagic sediments and ophiolites. Sedimentology 56:149–190

Friday 9 February 2024

ANAHOPLITES PLANUS OF FRANCE

A beautiful specimen of the ammonite, Anahoplites planus (Mantell, 1822) from Albian deposits in Villemoyenne Quarry, Courcelles, Aube, north-central France.

Anahoplites (Hyatt, 1900) is a genus of compressed hoplitid ammonites with flat sides, narrow, flat or grooved venters, and flexious ribs or striae arising from weak umbilical tubercles that end in fine dense ventrolateral nodes.

This lovely has attracted some roommates — an oyster, some bryozoans and worm tubes are attached to her shell.

Anahoplites is now included in the subfamily Anahoplitinae and separated from the Hoplitinae where it was placed in the older in the 1957 edition of the Treatise on Invertebrate Paleontology, Part L (Ammonoidea). Genera of the Hoplitinae tend to be more robust, with broader whorls and stronger ribs.

Anahoplites is found in Cretaceous (Middle to the Late Albian) deposits from England, through Europe, all the way to the Transcaspian Oblast region in Russia to the east of the Caspian Sea. The Aube department, named after the local river, is the type locality of the Albian stage (d'ORBIGNY, 1842). 

A. planus from the French Coast
Two formations are recognized in the clay facies (the "Gault" auct.) of the stratotype, the Argiles tégulines de Courcelles (82 m), overlain by the Marnes de Brienne (43 m). The boundary between the two formations is well-defined at the top of an indurated bed and readily identifiable in the field.

This involute (113 mm) specimen shows evidence of cohabitation by some of his marine peers. 

We see two different bryozoa, an oyster and some serpulids making a living and leaving trace fossils on her flat sides. The top specimen was prepared with potase by José Juárez Ruiz of Spain. 

The lovely Anahoplites planus you see here to the lower right was found by Bertus op den Dries on the French coast in Albian deposits near Wissant, P5 and measures in at 8 cm. This on edge view gives you a very good sense of the keel.

Wednesday 7 February 2024

VANCOUVER ISLAND'S TRENT RIVER PALAEONTOLOGY

Dan Bowen, Chair, VIPS, Trent River
The rocks that make up the Trent River on Vancouver Island are on the move. They were laid down near of the equator as small, tropical islands. They rode across the Pacific heading north and slightly east over the past 85 million years to where we find them today.

The Pacific Plate is an oceanic tectonic plate that lies beneath the Pacific Ocean. And it is massive. At 103 million km2 (40 million sq mi), it is the largest tectonic plate and continues to grow fed by volcanic eruptions that piggyback onto its trailing edge.

This relentless expansion pushes the Pacific Plate into the North American Plate. The pressure subducts it beneath our continent where it then melts back into the earth. Plate tectonics are slow but powerful forces. 

The island chains that rode the plates across the Pacific smashed into our coastline and slowly built the province of British Columbia. And because each of those islands had a different origin, they create pockets of interesting and diverse geology.

It is these islands that make up the Insular Belt — a physio-geological region on the northwestern North American coast. It consists of three major island groups — and many smaller islands — that stretches from southern British Columbia up into Alaska and the Yukon. These bits of islands on the move arrived from the Late Cretaceous through the Eocene — and continues to this day.

The rocks that form the Insular Superterrane are allochthonous, meaning they are not related to the rest of the North American continent. The rocks we walk over along the Trent River are distinct from those we find throughout the rest of Vancouver Island, Haida Gwaii, the rest of the province of British Columbia and completely foreign to those we find next door in Alberta.

To discover what we do find on the Trent takes only a wee stroll, a bit of digging and time to put all the pieces of the puzzle together. The first geological forays to Vancouver Island were to look for coal deposits, the profitable remains of ancient forests that could be burned to the power industry.

Jim Monger and Charlie Ross of the Geological Survey of Canada both worked to further our knowledge of the complex geology of the Comox Basin. They were at the cutting edge of west coast geology in the 1970s. It was their work that helped tease out how and where the rocks we see along the Trent today were formed and made their way north.

We know from their work that by 85 million years ago, the Insular Superterrane had made its way to what is now British Columbia. 

The lands were forested much as they are now but by extinct genera and families. The fossil remains of trees similar to oak, poplar, maple and ash can be found along the Trent and Vancouver Island. We also see the lovely remains of flowering plants such as Cupanities crenularis, figs and breadfruit.

Heading up the river, you come to a delineation zone that clearly marks the contact between the dark grey marine shales and mudstones of the Haslam Formation where they meet the sandstones of the Comox Formation. Fossilized material is less abundant in the Comox sandstones but still contains some interesting specimens. Here you begin to see fossilized wood and identifiable fossil plant material.

Further upstream, there is a small tributary, Idle Creek, where you can find more of this terrestrial material in the sandy shales. As you walk up, you see identifiable fossil plants beneath your feet and jungle-like, overgrown moss-covered, snarly trees all around you.

Walking west from the Trent River Falls at the bottom, you pass the infamous Ammonite Alley, where you can find Mesopuzosia sp. and Kitchinites sp. of the Upper Cretaceous (Santonian), Haslam Formation. Minding the slippery green algae covering some of the river rocks, you can see the first of the Polytychoceras vancouverense zone.

Continuing west, you reach the first of two fossil turtle sites on the river — amazingly, one terrestrial and one marine. If you continue, you come to the Inland Island Highway.

The Trent River has yielded some very interesting marine specimens, and significant terrestrial finds. We have found a wonderful terrestrial helochelydrid turtle, Naomichelys speciosa, and the caudal vertebrae of a Hadrosauroid dinosaur. Walking down from the Hadrosaur site you come to the site of the fossil ratfish find — one of the ocean's oddest fish.

Ratfish, Hydrolagus Collie, are chimaera found in the north-eastern Pacific Ocean today. The fossil specimen from the Trent would be considered large by modern standards as it is a bruiser in comparison to his modern counterparts. 

This robust fellow had exceptionally large eyes and sex organs that dangled enticingly between them. You mock, but there are many ratfish who would differ. While inherently sexy by ratfish standards, this fellow was not particularly tasty to their ancient marine brethren (or humans today) — so not hugely sought after as a food source or prey.

A little further again from the ratfish site we reach the contact of the two Formations. The rocks here have travelled a long way to their current location. With them, we peel away the layers of the geologic history of both the Comox Valley and the province of British Columbia.

The Trent River is not far from the Puntledge, a river whose banks have also revealed many wonderful fossil specimens. The Puntledge is also the name used by the K'ómoks First Nation to describe themselves. They have lived here since time immemorial. Along with Puntledge, they refer to themselves as Sahtloot, Sasitla and Ieeksun.

References: Note on the occurrence of the marine turtle Desmatochelys (Reptilia: Chelonioidea) from the Upper Cretaceous of Vancouver Island Elizabeth L. Nicholls Canadian Journal of Earth Sciences (1992) 29 (2): 377–380. https://doi.org/10.1139/e92-033; References: Chimaeras - The Neglected Chondrichthyans". Elasmo-research.org. Retrieved 2017-07-01.

Directions: If you're keen to explore the area, park on the side of Highway 19 about three kilometres south of Courtenay and hike up to the Trent River. Begin to look for parking about three kilometres south of the Cumberland Interchange. There is a trail that leads from the highway down beneath the bridge which will bring you to the Trent River's north side.

Monday 5 February 2024

CARNOTAURUS SASTREI: FLESH EATING BULL

Carnotaurus sastrei, a genus of large theropod dinosaurs that roamed the southern tip of Argentina, South America during the Late Cretaceous, 72 to 69.9 million years ago. His name means "flesh-eating bull,' and he lives up to it.

This fellow — or at least his robust skull with the short, knobby eyebrow horns and fierce-looking teeth — is on display at the Natural History Museum in Madrid, Spain. For now, he is the only known genus of this species of bipedal predator.

The first specimen of Carnotaurus sastrei was found in Chubut on vast plains between the Andes Mountains and the Atlantic Ocean. A physician, Dr. A'ngel Tailor noticed a large concretion showing some bone fragments. A team led by José F. Bonaparte excavated the find in 1984 as part of a paleontological expedition funded by the Argentine Museum of Natural Sciences.

Sadly, Bonaparte — the Maestro del Mesozoico — passed away the 18th February 20220 at the age of 91. He spent the majority of his career as head of the Vertebrate Palaeontology Division of the Museo Argentino de Ciencias Naturales “Bernardino Rivadavia,” in Buenos Aires. Bonaparte opened up the vertebrate finds of Argentina to the world. He was instrumental in the finding, excavating and naming many iconic dinosaurs — Carnotaurus, Amargasaurus, Abelisaurus, Argentinosaurus, Noasaurus along with the finding of the first fossilised remains of Mesozoic South American mammals. He mentored many palaeontologists who will miss his keen eye and tremendous work ethic — Luis Chiappe, Rodolfo Coria, Agustín Martinelli, Fernando Novas, Jaime Powell, Guillermo Rougier, Leonardo Salgado, Sebastián Apesteguía and many others.

His excavation of Carnotaurus was the first of its kind and he recognized that the skull is quite unusual. Initially, it has a very marine reptile feel — but make no mistake this guy is clearly a terrestrial theropod. He had smallish, underdeveloped arms — teeny by theropod standards. Once you look closer you see his bull-like horns from whence he gets his name — horns that imply battle between rivals for the best meal, sexual partner and to be the one who leads the herd. 

He was covered in leathery skin lined with rows of cone-shaped nodules or bumps. These get larger as they move towards his spine. He had forward-facing eyes, similar to tyrannosaurs like T-rex and smaller theropods like Velociraptor and Troodon — who had better vision even that T-rex — which would have given him the advantage of binocular vision and depth perception. Forward-facing eyes are also quite helpful with nocturnal hunting — think owls and cats — as they take in more light and help with nighttime predation. So perhaps this flesh-eating bull fancied a late-night snack on his menu from time to time.

Species like squirrels, pigeons and crocodiles have eyes on the sides of their heads. They lack the important competitive feature of well-developed depth perception — being able to easily and estimate distance — but perhaps make up for it with a panorama that offers a wider field of view.   

Saturday 3 February 2024

OPHTHALMOPLAX BRASILIANA

Ophthalmoplax brasiliana  / Photo: José F. Ventura‎
Ventral view of the carnivorous portunoid crab Ophthalmoplax brasiliana (Maury, 1930) from the latest Maastrichtian (~66.2 Ma.) deposits near Coahuila, northern Mexico.

This marine species was originally thought to have been found only in the upper Member (Owl Creek Formation) Late/Upper Maastrichtian deposits of Tippah County in Mississippi, USA. 

Sohl and Koch published on the Mississippian finds in the USGS in 1983. Fedorov and Nyborg published on this same species again in 2017. Paleocoordinates: (34.8° N, 88.9° W: 38.3° N, 66.2° W)

Friday 2 February 2024

CHAMPAGNE-ARDENNE HOPLITES

An excellent example of the ammonite, Hoplites bennettiana (Sowby, 1826) with a pathology. This beauty is from Albian deposits near Carrière de Courcelles, Villemoyenne, laid down in the Cretaceous near la région de Troyes (Aube) Champagne in northeastern France.

L'Albien or Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It was named after Alba, the Latin name for the River Aube, a tributary of the Seine that flows through the Champagne-Ardenne region of northwestern France.

The Albian is the youngest or uppermost subdivision of the Lower Cretaceous, approximately 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma (million years ago).

At the time that this fellow was swimming in our oceans, ankylosaurs were strolling about Mongolia and stomping through the foliage in Utah, Kansas and Texas. Bony fish were swimming over what would become the strata making up Canada, the Czech Republic and Australia. Cartilaginous fish were prowling the western interior seaway of North America and a strange extinct herbivorous mammal, Eobaatar, was snuffling through Mongolia, Spain and England. Interesting times.

Hoplites maritimus / Hoplites rudis
Hoplites are amongst my favourite ammonites. I still have a difficult time telling them apart. To the right, you can see a slightly greyish, Hoplites maritimus, from Sussex England. Below him is a brownish Hoplites rudis from outcrops between Courcelles and Troyes, France. There are many Hoplites species. Each has a nicely raised tire-track ribbing. My preference is for Hoplities bennetianus (or bennettiana). I'm still sorting out the naming of that species. The difference between Hoplites bennettiana and Hoplites dentatus is seen on the venter.

Hoplites shells have compressed, rectangular and trapezoidal whorl sections. They have pronounced umbilical bullae from which their prominent ribs branch out. The ends of the ribs can be both alternate or opposite. Some species have zigzagging ribs and these usually end thickened or raised into ventrolateral tubercules.

Ammonites were predatory, squid-like creatures that lived inside coil-shaped shells. Like other cephalopods, ammonites had sharp, beaklike jaws inside a ring of tentacles that extended from their shells to snare prey such as small fish and crustaceans. Some ammonites grew more than three feet (one meter) across — possible snack food for the giant mosasaur Tylosaurus.

Ammonites constantly built new shell as they grew, but only lived in the outer chamber. They scooted through the warm, shallow seas by squirting jets of water from their bodies. A thin, tubelike structure called a siphuncle reached into the interior chambers to pump and siphon air and helped them move through the water.

Ammonites first appeared about 240 million years ago, though they descended from straight-shelled cephalopods called bacrites that date back to the Devonian, about 415 million years ago. They were prolific breeders, lived in schools, and are among the most abundant fossils found today. They went extinct with the dinosaurs 65 million years ago. Scientists use the various shapes and sizes of ammonite shells that appeared and disappeared through the ages to date other fossils.

Hoplites sp. from the Early Cretaceous of Dorset, UK
During their evolution, three catastrophic events occurred. The first during the Permian period (250million years ago), only 10% survived.  They went on to flourish throughout the Triassic period, but at the end of this period (206 million years ago), all but one species died. Then they began to thrive from the Jurassic period until the end of the Cretaceous period when all species of ammonites became extinct.

Ammonites began life very tiny, less than 1mm in diameter, and were vulnerable to attack from predators. They fed on plankton and quickly assumed a strong protective outer shell. They also grew quickly with the females growing up to 400% larger than the males; because they needed the larger shell for egg production. Most ammonites only lived for two years.  Some lived longer becoming very large. The largest ever found was in Germany (6.5 feet in diameter).

Ammonites lived in shallow waters of 100 meters or less. They moved through the water by jet propulsion expelling water through a funnel-like opening to propel themselves in the opposite direction. They were predators (cephalopods) feeding on most living marine life including mollusks, fish even other cephalopods. Ammonites would silently stalk their prey then quickly extend their tentacles to grab it.  When caught the prey would be devoured by the Ammonites' jaws located at the base of the tentacles between the eyes.

Hoplites dentalus, from Albian deposits near Troyes, France
Most ammonites have coiled shells. The chambered part of the shell is called a phragmocone.  It contains a series of progressively layered chambers called camerae, which were divided by thin walls called septae. The last chamber is the body chamber.

As the ammonite grew, it added new and larger chambers to the opened end of the shell. A thin living tube called a siphuncle passed through the septa, extending from the body to the empty shell chambers.

This allowed the ammonite to empty water out of the shell chambers by hyperosmotic active transport process. This process controlled the buoyancy of the ammonite's shell.

First Photo: Hoplites Bennettiana from near Troyes, France. Collection de Christophe Marot

Second Photo: Top: Hoplites maritimus from Sussex, UK. Bottom: Hoplites rudis from near Troyes, France. Collection of Mark O'Dell

Third Photo: Hoplites sp. from the Early Cretaceous of Dorset, UK. Natural Selection Fossils

Fourth Photo: Hoplites dentalus from Albian deposits near Troyes, France. Collection of Stéphane Rolland.

Wright, C. W. (1996). Treatise on Invertebrate Paleontology, Part L, Mollusca 4: Cretaceous Ammonoidea (with contributions by JH Calloman (sic) and MK Howarth). Geological Survey of America and University of Kansas, Boulder, Colorado, and Lawrence, Kansas, 362.

Amédro, F., Matrion, B., Magniez-Jannin, F., & Touch, R. (2014). La limite Albien inférieur-Albien moyen dans l’Albien type de l’Aube (France): ammonites, foraminifères, séquences. Revue de Paléobiologie, 33(1), 159-279.

Thursday 1 February 2024

AMMONITE OF THE RHÔNE

An exquisite specimen of the delicately ridged ammonite, Porpoceras verticosum, from Middle Toarcian outcrops adjacent the Rhône in southeastern France.

Porpoceras (Buchman, 1911) is a genus of ammonite that lived during the early and middle Toarcian stage of the Early Jurassic. We see members of this genus from the uppermost part of Serpentinum Zone to Variabilis Subzone. These beauties are found in Europe, Asia, North America and South America.

Ammonites belonging to this genus have evolute shells, with compressed to depressed whorl section. Flanks were slightly convex and venter has been low. The whorl section is sub-rectangular. 

The rib is pronounced and somewhat fibulate on the inner whorls — just wee nodes here — and tuberculate to spined on the ventrolateral shoulder. It differs from Peronoceras by not having a compressed whorl section and regular nodes or fibulation. Catacoeloceras is also similar, but it has regular ventrolateral tubercules and is missing the classic nodes or fibulation of his cousins.

This specimen hails from southern France near the Rhône, one of the major rivers of Europe. It has twice the average water level of the Loire and is fed by the Rhône Glacier in the Swiss Alps at the far eastern end of the Swiss canton of Valais then passes through Lake Geneva before running through southeastern France. This 10 cm specimen was prepared by the supremely talented José Juárez Ruiz

Wednesday 31 January 2024

H. SAVENYEI: RENE'S BEE

This is a tale of friendship, tragic loss and fossil bees and an introduction to one of the most delightful paleo enthusiasts I have ever had the pleasure to know and collect with — Rene Savenye. He and I enjoyed many years of waxing poetic about our shared love of palaeontology and natural history. 

Rene was a mountain goat in the field, stalking the hills in his signature red t-shirt. He was tremendously knowledgeable about the natural world and delighted in it. For many years, he was Chair of the White Rock and Surrey Naturalists, while I was Chair of the Vancouver Paleontological Society. Together, we would plan and often co-lead field trips to many of the wonderful fossil outcrops in British Columbia and Washington state. 

In 2002, we were planning a very exciting round of field trips. I was offered a fully paid trip to India with Karen Lund to hike to the headwaters of the Ganges, a trip which I was to forgo in favour of a hike up to the outcrops of the Cathedral Escarpment and Burgess Shale and then to yummy Lower Jurassic and Lower Cretaceous, Albian, outcrops accessed only by boat in Haida Gwaii. 

Rene and I had talked about "walking in the shoes" of Joseph Whiteaves, the GSC's chief palaeontologist in Ottawa. He published a paper in 1876 describing the Jurassic and Cretaceous faunas of Skidegate Inlet and spent a significant portion of his career working out the fossil fauna of the Burgess Shale. Combining these two sites within the same field season was a fitting homage. 

John Fam, Vancouver Paleontological Society (VanPS) and Dan Bowen, Vancouver Island Palaeontological Society (VIPS), did much of the planning for that Haida Gwaii trip, they too being inspired by Whiteaves papers and the work of James Richardson and George Dawson — as a whole, we were giddy with the prospect of the year ahead.

Rene and I had planned to do both, but in the end, I had to give up the hike to Burgess that year and Rene never made it back to join me in Haida Gwaii. 

Rene Savenye
In the days before the official trip to Burgess, Rene did some solo hiking in the mountains and hills near Field, British Columbia. He was excited to test his stamina against the steep passes that protect the majestic ridges of Wapta Mountain, Mount Field and Mount Stephen — ever mindful of collecting only with his camera. 

He walked through the hallowed footsteps of Joseph Whiteaves and Charles Doolittle Walcott over ground that should have been named La Entrada de Dios, The Gateway of God, for each footfall brought him closer to meeting the big man. While a naturalist, Rene held to the belief that once his days were done on this Earth, he would be breaking bread in heaven above. 

Rene started with clear skies and a pack full of geology hammers, maps and chisels — the hillside a sea of white and pink flecked wildflowers in the sunlight. As the day went on, the skies filled with rolling clouds, then thunder. Grey sheets of rain covered the landscape. Seeing the danger of being solo in darkening weather, he started down the slope back to his car — his shadow long and thin striking out before him in the fading light — but he never made it. On the afternoon of July 28th, he was struck and killed by lightning — a tragic loss. 

I take heart that he lived and died doing what he loved most. I got the news a few days later and cried for the loss of a great friend. I am sharing my memory of him with you so that you can remember him, too, and share in the delight and loss of one of the loveliest men to ever walk our planet. His years of teaching, mentoring, encouragement and generosity have helped shape natural science and those who have gone on to make it their passion or career — or happily, both.   

Rene's name will not be forgotten to science. His namesake, H. Savenyei, is a lovely fossil halictine bee from Early Eocene deposits near Quilchena, British Columbia — and the first bee body-fossil known from the Okanagan Highlands — and indeed from Canada. 

As a school teacher, Rene once taught the, then student, now SFU biology instructor, Rolf Mathewes. Rene passed his scientifically valuable specimen to Mathews, knowing it was important to science. Mathewes brought it to the attention of Bruce Archibald and Michael Engel, who described Rene's bee in the Canadian Journal of Zoology. Their work is a lovely legacy to a wonderful man and a specimen from one of his favourite collecting sites — Quilchena — a small road-cut exposure of the Coldwater beds of the Princeton Group, one of several depositional basins in the Merritt region of south-central British Columbia.

Rene is also remembered in spirit by the British Columbia Paleontological Alliance (BCPA) Rene Savenye Award. It was established in 2003 to honour those who have demonstrated outstanding service to the science of palaeontology or to palaeontological education in British Columbia. 

Notable past recipients are a veritable who's who from the Pacific Northwest — Graham Beard of Qualicum in 2005, Charles Helm of Tumbler Ridge in 2011, Pat Trask of Courtenay in 2014, Rod Bartlett in 2016, and Joseph "Joe" Haegert in 2018. I'll share a link to the award below so you can read more at your leisure about Rene and those who bear the award with his name.

About H. Savenyei, (Engel & Archibald, 2003): The type specimen is a fairly well preserved complete adult female preserved with portions of the fore-wings and hind-wings. The specimen is 7.04 millimetres (0.277 in) long with the possibility of alteration in length during fossilization. The sections of the forewing which are preserved are approximately 4.8 millimetres (0.19 in) long and show dark brown to black colouration. The presence of a pygidial plate bordered by setae on the fifth metasomal tergum supports the placement into the Halictidae subfamily Halictinae. Placement into the tribe Halictini is based on the lack of a medial cleft in the fifth tergum.

References:

Archibald, B. & R. W. Mathewes. 2000. “Early Eocene Insects from Quilchena, BC, and their Paleoclimatic Implications.” Canadian Journal of Zoology, Volume 78, Number 6: pp 1441-1462.

Grimaldi, D. 1999. “The Co-radiations of Pollinating Insects and Angiosperms in the Cretaceous.” Annals of the Missouri Botanical Garden. 86: 373-406.

Photo: Halictidae sp.; Archibald and Mathewes 2000: 1453.

Rene Savenye Award: https://bcfossils.ca/rene-savenye-award

Monday 29 January 2024

FOSSILS AND FAUNA OF MADAGASGAR

Aioloceras besairiei (Collingnon, 1949)
A stunning example of the internal suturing with calcite infill in this sliced Aioloceras besairiei (Collingnon, 1949) ammonite from the Upper Cretaceous (Lower Albian) Boeny region of Madagascar. 

This island country is 400 kilometres off the coast of East Africa in the Indian Ocean and a wonderful place to explore off the beaten track.

Madagascar has some of the most spectacular of all the fossil specimens I have ever seen. This beauty is no exception. The shell has a generally small umbilicus, arched to acute venter, and typically at some growth stage, falcoid ribs that spring in pairs from umbilical tubercles, usually disappearing on the outer whorls. I had originally had this specimen marked as a Cleoniceras besairiei, except Cleoniceras and Grycia are not present in Madagascar. 

This lovely, seen in cross-section, is now far from home and in the collection of a wonderful friend. It is an especially lovely example of the ammonite, Aioloceras besairiei, making it a beudanticeratinae. Cleoniceras and Grycia are the boreal genera. If you'd like to see (or argue) the rationale on the name, consider reading Riccardi and Medina's riveting work from back in 2002, or Collingnon from 1949.

The beauty you see here measures in at a whopping 22 cm, so quite a handful. This specimen is from the youngest or uppermost subdivision of the Lower Cretaceous. I'd originally thought this locality was older, but dating reveals it to be from the Lower Albian, so approximately 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma.

Aioloceras are found in the Cretaceous of Madagascar at geo coordinates 16.5° S, 45.9° E: paleo-coordinates 40.5° S, 29.3° E.; and in four localities in South Africa: at locality 36, near the Mzinene River at 28.0° S, 32.3° E: paleo-coordinates 48.6° S, 7.6° E. 

We find them near the Mziene River, at a second locality north of Hluhluwe where the Mzinene Formation overlies the Aptian-Albian Makatini Formation at 28.0° S, 32.3° E: paleo-coordinates 48.6° S, 7.6° E; and at Haughton Z18, on the Pongola River in the Albian III, Tegoceras mosense beds at 27.3° S, 32.2° E: paleo-coordinates 48.0° S, 7.8° E.

If you happen to be trekking to Madagascar, know that it's big. It’s 592,800 square kilometres (or  226,917 square miles), making it the fourth-largest island on the planet — bigger than Spain, Thailand, Sweden and Germany. The island has an interesting geologic history.

Although there has been a geological survey, which was active extending back well into French colonial times, in the non-French-speaking world our geological understanding of the island is still a bit of a mystery. 

Plate tectonic theory had its beginnings in 1915 when Alfred Wegener proposed his theory of "continental drift." 

Wegener proposed that the continents ploughed through the crust of ocean basins, which would explain why the outlines of many coastlines (like South America and Africa) look like they fit together like a puzzle. Half a century after Wegener there is still no agreement as to whether in continental reconstructions Madagascar should be placed adjacent to the Tanzanian coast to the north (e.g., McElhinny and Embleton,1976), against the Mozambique-Natal coast (Flores 1970), or basically left where it is (Kent 1974, Nairn 1978).

There have been few attempts apart from McKinley’s (1960) comparison of the Karoo succession of southwestern Tanzania with that of Madagascar to follow the famous geological precept of “going to sea.” One critical reason is that although there may be a bibliography of several thousand items dealing with Madagascan geology as Besairie (1971) claims, they are items not generally available to the general public. The vital information gained of the geology of the offshore area by post-World War II petroleum exploration has remained largely proprietary. 

Without this data to draw upon, our understanding remains incomplete. I don't actually mind a bit of a mystery here. It is interesting to speculate on how these geologic puzzle pieces fit together and wait for the big reveal. Still, we have good old Besairie from his 1971, Geologie de Madagascar, and a later précis (Besairie, 1973).

We do know that Madagascar was carved off from the African-South American landmass early on. The prehistoric breakup of the supercontinent Gondwana separated the Madagascar–Antarctica–India landmass from the Africa–South America landmass around 135 million years ago. Madagascar later split from India about 88 million years ago, during the Late Cretaceous, so the native plants and animals on the island evolved in relative isolation. 

It is a green and lush island country with more than its fair share of excellent fossil exposures. Along the length of the eastern coast runs a narrow and steep escarpment containing much of the island's remaining tropical lowland forest. If you could look beneath this lush canopy, you'd see rocks of the Precambrian age stretching from the east coast all the way to the centre of the island. The western edge is made up of sedimentary rock from the Carboniferous to the Quaternary.

Red-Tailed Lemurs, Waiwai & Hedgehog
Madagascar is a biodiversity hotspot. Just as Darwin's finches on the Galápagos were isolated, evolving into distinct species (hello, adaptive radiation), over 90% of the wildlife from Madagascar is found nowhere else. 

The island's diverse ecosystems, like so many on this planet, are threatened by Earth's most deadly species, homo sapien sapiens. 

We arrived back in 490 CE and have been chopping down trees and eating our way through the island's tastier populations ever since. Still, they have cuties like this Red-Tailed Lemur. Awe, right?

Today, beautiful outcrops of wonderfully preserved fossil marine fauna hold appeal for me. The material you see from Madagascar is distinctive — and prolific.

Culturally, you'll see a French influence permeating the language, architecture and legal process. There is a part of me that pictures these lovely Lemurs chatting away in French. "Ah, la vache! Regarde le beau fossile, Hérissonne!"

We see the French influence because good 'ol France invaded sleepy Madagascar back in 1883, during the first Franco-Hova War. Malagasy (the local Madagascarian residents) were enlisted as troops, fighting for France in World War I.  During the Second World War, the island was the site of the Battle of Madagascar between the Vichy government and the British. By then, the Malagasy had had quite enough of colonization and after many hiccuping attempts, reached full independence in 1960. Colonization had ended but the tourist barrage had just begun. You can't stop progress.

If you're interested in learning more about this species, check out the Treatise on Invertebrate Paleontology, Part L (Ammonoidea). R.C. Moore (ed). Geological Soc of America and Univ. Kansas Press (1957), p L394. Or head over to look at the 2002 paper from Riccardi and Medina. 2002. Riccardi, A., C. & Medina, F., A. The Beudanticeratinae and Cleoniceratinae (Ammonitina) from the Lower Albian of Patagonia in Revue de Paléobiologie - 21(1) - Muséum d’Histoire Naturelle de la ville de Genève, p 313-314 (=Aioloceras besairiei (COLLIGNON, 1949). You have Bertrand Matrion to thank for the naming correction. Good to have friends in geeky places!

Collignon, M., 1933, Fossiles cenomaniens d’Antmahavelona (Province d’ Analalave, Madagascar), Ann. Geol. Serv. Min. Madagascar, III, 1934 Les Cephalopods du Trias inferieur de Madagascar, Ann. Paleont. XXII 3 and 4, XXII 1.

Besairie, H., 1971, Geologie de Madagascar, 1. Les terrains sedimentaires, Ann. Geol. Madagascar, 35, p. 463.

J. Boast A. and E. M. Nairn collaborated on a chapter in An Outline of the Geology of Madagascar, that is very readable and cites most of the available geologic research papers. It is an excellent place to begin a paleo exploration of the island.

If you happen to parle français, check out: Madagascar ammonites: http://www.ammonites.fr/Geo/Madagascar.htm

Sunday 28 January 2024

THE SLOW RACE OF TIME: LIVING FOSSILS

Horseshoe crabs are marine and brackish water arthropods of the order Xiphosura — a slowly evolving, conservative taxa.

Much like (slow) Water Striders (Aquarius remigis), (relatively sluggish) Coelacanth (Latimeria chalumnae) and (the current winner on really slow evolution) Elephant Sharks (Callorhinchus milii), these fellows have a long history in the fossil record with very few anatomical changes. 

But slow change provides loads of great information. It makes our new friend, Yunnanolimulus luoingensis, an especially interesting and excellent reference point for how this group evolved. 

We can examine their genome today and make comparisons all the way back to the Middle Triassic (with this new find) and other specimens from further back in the Ordovician — 445 million years ago. 

These living fossils have survived all five mass extinction events. They are generalists who can live in shallow or deep water and will eat pretty much anything they can find on the seafloor.

The oldest horseshoe crab fossil, Lunataspis aurora, is found in outcrops in Manitoba, Canada. Charmingly, the name means crescent moon shield of the dawn. It was palaeontologist Dave Rudkin and team who chose that romantic name. Finding them as fossils is quite remarkable as their shells are made of protein which does not mineralized like typical fossils.

Even so, the evolution of their exoskeleton is well-documented by fossils, but appendage and soft-tissue preservation are extremely rare. 

A new study analyzes details of the appendage and soft-tissue preservation in Yunnanolimulus luoingensis, a Middle Triassic (ca. 244 million years old) horseshoe crab from Yunnan Province, SW China. The remarkable anatomical preservation includes the chelicerae, five pairs of walking appendages, opisthosomal appendages with book gills, muscles, and fine setae permits comparison with extant horseshoe crabs.

The close anatomical similarity between the Middle Triassic horseshoe crabs and their recent analogues documents anatomical conservatism for over 240 million years, suggesting persistence of lifestyle.

The occurrence of Carcinoscorpius-type claspers on the first and second walking legs in male individuals of Y. luoingensis tells us that simple chelate claspers in males are plesiomorphic for horseshoe crabs, and the bulbous claspers in Tachypleus and Limulus are derived.

As an aside, if you hadn't seen an elephant shark before and were shown a photo, you would likely say, "that's no freaking shark." You would be wrong, of course, but it would be a very clever observation.

Callorhinchus milii look nothing like our Great White friends and they are not true sharks at all. Rather, they are ghost sharks that belong to the subclass Holocephali (chimaera), a group lovingly known as ratfish. They diverged from the shark lineage about 400 million years ago.

If you have a moment, do a search for Callorhinchus milii. The odd-looking fellow with the ironic name, kallos, which means beautiful in Greek, sports black blotches on a pale silver elongate body. And their special feature? It is the fishy equivalent of business in the front, party in the back, with a dangling trunk-like projection at the tip of their snout and well-developed rectal glands near the tail.

As another small point of interest with regards to horseshoe crabs, John McAllister collected several of these while working on his MSc to see if they had microstructures similar to trilobites (they do) and whether their cuticles were likewise calcified. He found no real calcification in their cuticles, in fact, he had a rather frustrating time getting anything measurable to dissolve in acid in his hunt for trace elements. 

Likewise, when looking at oxygen isotopes (16/18) to get a handle on water salinity and temperature, his contacts at the University of Waterloo had tons of fun getting anything at all to analyze. It made for some interesting findings. Sadly, for a number of reasons, he abandoned the work, but you can read his very interesting thesis here: https://dr.library.brocku.ca/handle/10464/1959

Ref: Hu, Shixue & Zhang, Qiyue & Feldmann, Rodney & Benton, Michael & Schweitzer, Carrie & Huang, Jinyuan & Wen, Wen & Zhou, Changyong & Xie, Tao & Lü, Tao & Hong, Shuigen. (2017). Exceptional appendage and soft-tissue preservation in a Middle Triassic horseshoe crab from SW China. Scientific Reports. 7. 10.1038/s41598-017-13319-x.