BARNACLES: CUVIER TO DARWIN

Barnacles All Closed Up

One of the most interesting and enigmatic little critters we find at the seashore are barnacles. 

They cling to rocks at the waters' edge, closed to our curiosity, their domed mounds like little closed beaks shut to the water and the world.

They choose their permanent homes as larvae, sticking to hard substrates that will become their permanent homes for the rest of their lives. It has taken us a long time to find how they actually stick or what kind of "glue" they were using.

A clever fellow from Duke University's Marine Laboratory in Durhan, North Carolina finally cracked that puzzle. Instead of chopping up barnacles to see what makes them stick, he observed and collected the oozing glue from some Amphibalanus amphitrite as they secreted it.

Remarkably, the barnacle glue sticks to rocks in a similar way to how red cells bind together. Red blood cells bind and clot with a little help from some enzymes. These work to create long protein fibres that first blind, clot then form a scab. The mechanism barnacles use, right down to the enzyme, is very similar. That's especially interesting as about a billion years separate our evolutionary path from theirs.

So, with the help of their clever enzymes, they can affix to most anything – ship hulls, rocks, and even the skin of whales. If you find them in tidepools, you begin to see their true nature as they open up, their delicate feathery finger-like projections flowing back and forth in the surf.

Barnacle Cirri Seeking Tasty Plankton

Those wee feather-like bits you see are called cirri. Eight pairs of these thoracic limbs help barnacles to filter tasty bits of plankton from the surrounding water into their mouths.

Barnacles are cirripedes, a kind of crustacean that is covered with hard plates of calcium carbonate. Named for their cirri, they live stuck to hard surfaces in and around our world's oceans. While they do not look like crustaceans, they are definitely part of this taxonomic grouping that includes crab, lobster, crayfish, prawn, krill, and woodlice.

BARNACLES IN KWAK'WALA

In the Kwak̓wala language of the Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, barnacles are known as k̕wit̕a̱'a and broken barnacle shells are known as t̕sut̕su'ma.

BARNACLES IN THE FOSSIL RECORD

They have an old history. Their ancestors can be traced back to animals such as Priscansermarinus that lived during the Middle Cambrian – some 510 to 500 million years ago. I found my first barnacle fossil at a fossil site called Muir Creek on the south end of Vancouver Island. The fossil exposures at Muir are Oligocene, 20-25 million years old. This is about the time that barnacles can be found more readily as skeletal remains.

One of the reasons for the limited number of barnacle remains in the fossil record is their preferred habitat – high energy, shallow ocean environments. These tend to see a lot of tidal action that leads to erosion and barnacles being broken apart, slowly eroded down to bits too small to recognize for what they are.

One of the fossil remains we do find are not the barnacles themselves, but trace fossils of acrothoracican barnacle borings from Rogerella. These are commonly found in the fossil record beginning in the Devonian right up to today. Rogerella is a small pouch-shaped boring (a type of trace fossil) with a slit-like aperture currently produced by acrothoracican barnacles. 

These crustaceans extrude their legs upwards through the opening for filter-feeding (Seilacher, 1969; Lambers and Boekschoten, 1986). They are known in the fossil record as borings in carbonate substrates (shells and hardgrounds) from the Devonian to the Recent (Taylor and Wilson, 2003).

Barnacle Ancestry Goes Back to the Middle Cambrian

FROM MOLLUSCA TO ARTICULATA

Barnacles were originally classified by Linnaeus and Cuvier as Mollusca, but in 1830 John Vaughan Thompson published observations showing the metamorphosis of the nauplius and cypris larvae into adult barnacles. He noted how these larvae were similar to those of crustaceans.

In 1834 Hermann Burmeister published further information, reinterpreting these findings. The effect was to move barnacles from the phylum of Mollusca to Articulata, showing naturalists that detailed study was needed to reevaluate their taxonomy.

Charles Darwin took up this challenge in 1846 and developed his initial interest in a major study published as a series of monographs in 1851 and 1854. Darwin undertook this study, at the suggestion of his friend Joseph Dalton Hooker, to thoroughly understand at least one species before making the generalizations needed for his theory of evolution by natural selection.

BARNACLES IN A NUT SHELL

Barnacles are suspension feeders, sweeping small food into their mouth with their curved 'feet'. They are cemented to rock (usually), and covered with hard calcareous plates, which they shut firmly when the tide goes out. The barnacles reproduce sexually and produce little nauplius larvae that disperse in the plankton. Eventually, the larvae change into cypris form and attach on other hard surfaces to form new barnacles.

UPPER CRETACEOUS MOTORCROSS SITE: VANCOUVER ISLAND

Steller's Jay, Cyanocitta stelleri

One of the classic Vancouver Island fossil localities is the Santonian-Maastrichtian, Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake, Nanaimo, British Columbia, Canada.

The quarry is no longer active as such though there is a busy little gravel quarry a little way down the road closer to Ammonite falls near Benson Creek Falls.

Today it is an active motocross site and remains one of the classic localities of the Nanaimo Group. We find well-preserved nautiloids and ammonites — Canadoceras, Pseudoschloenbachia, Epigoniceras — the bivalves — Inoceramus, Sphenoceramus— gastropods, and classic Nanaimo Group decapods — Hoploparia, Linuparus. We also find fossil fruit and seeds which tell the story of the terrestrial history of Vancouver Island.

Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake

It was John Fam, Vice-Chair, Vancouver Island Paleontological Society (VanPS), who originally told me about the locality. John is one of the most delightful and knowledgeable people you'd be well-blessed to meet.

While he lived on Vancouver Island, he was an active member of the VanPS back when I was Chair. Several of the best joint VIPS/VanPS paleontological expeditions were planned with or instigated by his passion for fossils. I tip my hat to him for his passion and shared love of all things paleo.

John grew up 15 minutes from the motocross locality and used to collect there a few times a week with his father. John has wonderful parents and since marrying his childhood sweetheart, the amazing Grace, those excellent genetics, curiosity and love of fossils are now being passed to a new generation. It's lovely to see John and Grace continuing tradition with two boys of their own.

I met John way back then and did an overnight at his parent's house the Friday before a weekend field trip to Jurassic Point. It was a joy to have him walk me through his collections and tell his stories from earlier years. After learning about the site from John, I headed up to the Motocross Pit with my Uncle Doug. He was a delightful man who grew up on the coast and had explored much of it but not the fossil site just 10-minutes from his home. It was wonderful to walk through time with him so many years ago and then again solo this past year with sadness in my belly that one of the best I've ever known has left this Earth.

Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake

There were some no trespassing signs up but no people around, so I walked the periphery looking for the bedrock of the Haslam.

The rocks we find here were laid down south of the equator as small, tropical islands. They rode across the Pacific heading north and slightly east over the past 80 million years to where we find them today.

Jim Haggart and Peter Ward have done much to increase our understanding of the molluscan fauna of the Nanaimo Group. Personally, both personify the charming Indiana Jones school of rugged manly palaeontologists you picture in popular film. Professionally, their singular contributions and collaborative efforts have helped shape our understanding of the correlation of Nanaimo Group fauna to those we find in the Gulf Islands of British Columbia and down in the San Juan Islands of Washington State.

Their work builds on the work of Usher (1952), Matsumoto (1959a, 1959b) and Mallory (1977). A healthy nod goes out to the work of Muller and Jeletzky (1970) for untangling the lithostratigraphic and biostratigraphic foundation for our knowledge of the Nanaimo Group.

Candoceras yokoyama, Photo: John Fam, VanPS

As I walked along the bedrock of the Haslam, a Steller's Jay, Cyanocitta stelleri, followed me from tree to tree making his guttural shook, shook, shook call. Instructive, he seemed to be encouraging me, timing his hoots to the beat of my hammer. Vancouver Island truly has glorious flora and fauna.

Fancy some additional reading? Check out a paper published in the Journal of Paleontology back in 1989 by Haggard and Ward on new Nanaimo Group Ammonites from British Columbia and Washington State.

In it, they look at the ammonite species Puzosia (Mesopuzosia) densicostata Matsumoto, Kitchinites (Neopuzosia) japonicus Spath, Anapachydiscus cf. A. nelchinensis Jones, Menuites cf. M. menu (Forbes), Submortoniceras chicoense (Trask), and Baculites cf. B. boulei Collignon are described from Santonian--Campanian strata of western Canada and northwestern United States.

Stratigraphic occurrences and ranges of the species are summarized and those taxa important for correlation with other areas in the north Pacific region and Late Cretaceous ammonite fauna of the Indo-Pacific region. Here's the link: https://www.jstor.org/stable/1305358?seq=1

Peter Ward is a prolific author, both of scientific papers and more popularized works. I highly recommend his book Gorgon: Paleontology, Obsession, and the Greatest Catastrophe in Earth's History. It is an engaging romp through a decade's research in South Africa's Karoo Desert.

Photo: Candoceras yokoyamai from Upper Cretaceous Haslam formation (Lower Campanian) near Nanaimo, British Columbia. One of the earliest fossils collected by John Fam (1993). Prepared using only a cold chisel and hammer. Photo & collection of John Fam, VIPS.

SHONISAURUS OF NEVADA

The beauties you see here are ichthyosaurs. The largest of their lineage is the genus Shonisaurus who ruled our ancient seas 217 million years ago.

At least 37 incomplete fossil specimens of the marine reptile have been found in hard limestone deposits of the Luning Formation, in far northwestern Nye County of Nevada. This formation dates to the late Carnian age of the late Triassic period when present-day Nevada and parts of the western United States were covered by an ancient ocean.

The first researcher to recognize the Nevada fossil specimens as ichthyosaurs was Siemon W. Muller of Stanford University. He had the work of Sir Richard Owen and others to build on. That being said, there are very few contenders for a species that boasts vertebrae over a foot wide and weighing in at almost 10 kg or 21 lbs. Muller contacted the University of California Museum of Paleontology at Berkeley. Surface collecting by locals continued at the site but no major excavation was planned.

Sir Richard Owen, the British biologist, comparative anatomist and paleontologist, coined the name ichthyopterygia, or "fish flippers," one hundred and fourteen years earlier, but that wee bit of scientific knowledge hadn't made its way west to the general population. The finds at Luning were still, "marine monsters."

Owen, too, was building on research going back to 1699, the very first recorded fossil fragments found of these beasties in Wales. Shortly thereafter, fossil vertebrae were published in 1708 from the Lower Jurassic.

The first complete skeleton was discovered in the early 19th century by Mary Anning and her brother Joseph along the Dorset Jurassic Coast. Mary's find was described by a British surgeon, Sir Everard Home, an elected Fellow of the Royal Society, in 1814. The specimen is now on display at the Natural History Museum in London bearing the name Temnodontosaurus platyodon, or “cutting-tooth lizard.”

In 1821, William Conybeare and Henry De La Beche, a friend of Mary's, published a paper describing three new species of unknown marine reptiles based on the Anning's finds. The Rev. William Buckland would go on to describe two small ichthyosaurs from the Lias of Lyme Regis, Ichthyosaurus communis and Ichthyosaurus intermedius. All of this early work was instrumental in aiding the researchers who would join the project at Luning.

Owen is considered to have been an outstanding naturalist with a remarkable gift for interpreting fossils. Contrary to common belief, advanced study does help with identifying fossils, but what is truly needed is a keen eye. The finds at Luning were blessed to be seen by an enthusiastic local with just that right kind of keen eye.

Almost a quarter of a century after Muller's initial reports, Dr. Charles L. Camp from UCMP received correspondence further detailing the finds from a lovely Mrs. Margaret Wheat of Fallon. She wrote to Camp in September of 1928 to say that she'd been giving the quarry section a bit of a sweep, as you do, and had uncovered a nice aligned section of vertebrae with her broom. The following year, Dr. Charles L. Camp went out to survey the finds and began working on the specimens, his first field season of many, in 1954.

Back in the 1950s, these large marine reptiles were rumoured to be "marine monsters," as the concept of an ichthyosaur was not well understood by the local townsfolk. Excitement soon hit West Union Canyon as the quarry began to reveal the sheer size of these mighty beasts. Four of the specimens were fully excavated. Most of the ichthyosaur bones were left in situ, partially because the work was tremendously difficult, and partially to allow others to see how the specimens were laid down over 200 million years ago.

Camp continued to work with Wheat at the site and brought on Sam Welles and a host of students to help with excavations. The team understood the need for protection at the site. They canvassed the Nevada Legislature to establish the Ichthyosaur Paleontological State Monument. You can see one of the Park Rangers above giving a tour within the lovely Fossil Hut building they built on the site to protect the fossils.

In 1957, the site was incorporated into the State Park System and Berlin-Ichthyosaur State Park was born. The park Twenty years later, in 1977, the population of Nevada weighed in and the Legislature designated Shonisaurus popularis as the State Fossil of Nevada. Visitors are welcome to collect fossils from the exposures of the Upper Triassic (Early Norian, Kerri Zone) of the Luning Formation, West Union Canyon, just outside Berlin-Ichthyosaur State Park.

Address: State route 844, Austin, NV 89310, United States. Area: 4.58 km². Open 24 hours;
Elevation: 6,975 ft (2,126 m); Tel: +1 775-964-2440; http://parks.nv.gov/parks/berlin-ichthyosaur

BRONZE BEAUTY: EIFELIAN PARALEJURUS

This bronzed beauty is the Middle Devonian, Eifelian (~395 mya) trilobite, Paralejurus rehamnanus (Alberti, 1970) from outcrops near Issoumour, Alnif, Morocco in North Africa. 

It was the colour of this amazing trilobite that captured the eye of David Appleton in whose collection it now resides. He is an avid collector and coming into his own as a macro photographer. I have shared three of his delightful photos for you here.

It initially thought that the gold we see here was added during prep, particularly considering the colouration of the matrix, but macro views of the surface show mineralization and the veins running right through the specimen into the matrix. There is certainly some repairs but that is common in the restoration of these specimens. Many of the trilobites I have seen from Morocco have bronze on black colouring but not usually this pronounced. Even so, there is a tremendous amount of fine anatomy to explore and enjoy in this wonderfully preserved specimen.  

Paralejurus is a genus of trilobite in the phylum Arthropoda from the Late Silurian to the Middle Devonian of Africa and Europe. These lovelies grew to be up to nine centimetres, though the fellow you see here is a wee bit over half that size at 5.3 cm. 

Paralejurus specimens are very pleasing to the eye with their long, oval outline and arched exoskeletons. 

Their cephalon or head is a domed half circle with a smooth surface.  The large facet eyes have very pleasing crescent-shaped lids. You can see this rather well in the first of the photos here. The detail is quite remarkable.

As you move down from his head towards the body, there is an almost inconspicuous occipital bone behind the glabella in the transition to his burnt bronze thorax.

The body or thorax has ten narrow segments with a clearly arched and broad axial lobe or rhachis. The pygidium is broad, smooth and strongly fused in contrast to the genus Scutellum in the family Styginidae, which has a pygidium with very attractive distinct furrows that I liken to the look of icing ridges on something sweet — though that may just be me and my sweet tooth talking. In Paralejurus, they look distinctly fused — or able to fuse — to add posterior protection against predators with both the look and function of Roman armour.

In Paralejurus, the axillary lobe is rounded off and arched upwards. It is here that twelve to fourteen fine furrows extend radially to complete the poetry of his body design. 

Trilobites were amongst the earliest fossils with hard skeletons and they come in many beautiful forms. While they are extinct today, they were the dominant life form at the beginning of the Cambrian. 

As a whole, they were amongst some of the most successful of all early animals — thriving and diversifying in our ancient oceans for almost 300 million years. The last of their brethren disappeared at the end of the Permian — 252 million years ago. Now, we enjoy their beauty and the scientific mysteries they reveal about our Earth's ancient history.

Photos and collection of the deeply awesome David Appleton. Specimen: 5.3 cm. 

NEVADA: AMMONOIDS AND CONODONTS

Nevada is a wonderful place to explore our palaeontological history. The state spans a broad spectrum of exposures showcasing the depth of geologic time. It is an interesting cross-section of young and old — and interestingly, a lovely comparison to the Triassic outcrops in British Columbia.

Exposures of the Upper Triassic, Early Norian, Kerri zone, Luning formation, West Union Canyon, just outside Berlin-Ichthyosaur State Park, Nevada.

The Berlin-Ichthyosaur State Park in central Nevada is a very important locality for the understanding of the Carnian-Norian boundary (CNB) in North America.

Rich ammonoid faunas from this site within the Luning Formation were studied by Silberling (1959) and provided support for the definition of the Schucherti and Macrolobatus zones of the latest Carnian, which are here overlain by well-preserved faunas of the earliest Norian Kerri Zone. Despite its importance, no further investigations have been done at this site during the last 50 years.

Jim Haggart, Mike Orchard and Paul Smith collaborated on a project that took them down to Nevada to look at the conodonts and ammonoids; the group then published a paper, "Towards the definition of the Carnian/Norian Boundary: New data on Ammonoids and Conodonts from central Nevada," which you can find in the proceedings of the 21st Canadian Paleontology Conference; by Haggart, J W (ed.); Smith, P L (ed.); Canadian Paleontology Conference Proceedings no. 9, 2011 p. 9-10.

They conducted a bed-by-bed sampling of ammonoids and conodonts in West Union Canyon during October 2010. The eastern side of the canyon provides the best record of the Macrolobatus Zone, which is represented by several beds yielding ammonoids of the Tropites group, together with Anatropites div. sp. conodont faunas from both these and higher beds are dominated by ornate 'metapolygnthids' that would formerly have been collectively referred to Metapolygnathus primitius, a species long known to straddle the CNB. Within this lower part of the section, they resemble forms that have been separated as Metapolygnathus mersinensis. Slightly higher, forms close to 'Epigondolella' orchardi and a single 'Orchardella' n. sp. occur. This association can be correlated with the latest Carnian in British Columbia.

Ammonoids of the Luning Formation

Higher in the section, the ammonoid fauna shows a sudden change and is dominated by Tropithisbites. Few tens of metres above, but slightly below the first occurrence of Norian ammonoids Guembelites jandianus and Stikinoceras, two new species of conodonts (Gen et sp. nov. A and B) appear that also occur close to the favoured Carnian/Norian boundary at Black Bear Ridge, British Columbia. Stratigraphically higher collections continue to be dominated by forms close to M. mersinensis and 'E.' orchardi.

The best exposure of the Kerri Zone is on the western side of the West Union Canyon. Ammonoids, dominated by Guembelites and Stikinoceras div. sp., have been collected from several fossil-bearing levels. Conodont faunas replicate those of the east section. The collected ammonoids fit perfectly well with the faunas described by Silberling in 1959, but they differ somewhat from the coeval faunas of the Tethys and Canada.

The genus Gonionotites, very common in the Tethys and British Columbia, is for the moment unknown in Nevada. More in general, the Upper Carnian faunas are dominated by Tropitidae, while Juvavitidae are lacking.

After years of reading about the correlation between British Columbia and Nevada, I had the very great pleasure of walking through these same sections in October 2019 with members of the Vancouver Paleontological Society and Vancouver Island Palaeontological Society. It was with that same crew that I had originally explored fossil sites in the Canadian Rockies in the early 2000s. Those early trips led to paper after paper and the exciting revelations that inspired our Nevada adventure.

TIKTAALIK: FOSSIL FISHAPODS

Qikiqtania wakei, a fishapod & relative to tetrapods

You will likely recall the amazing tetrapodomorpha fossil found on Ellesmere Island in the Canadian Arctic in 2004, Tiktaalik roseae. 

These were advanced forms transitional between fish and the early labyrinthodonts playfully referred to as fishapods — half-fish, half-tetrapod in appearance and limb morphology. 

Up to that point, the relationship of limbed vertebrates (tetrapods) to lobe-finned fish (sarcopterygians) was well known, but the origin of significant tetrapod features remained obscure for the lack of fossils that document the sequence of evolutionary changes — until Tiktaalik. 

While Tiktaalik is technically a fish, this fellow is as far from fish-like as you can be and still be a card-carrying member of the group. 

Interestingly, while Neil Shubin and crew were combing the icy tundra for Tiktaalik, another group was trying their luck just a few kilometres away. 

A week before the eureka moment of Tiktaalik's discovery, Tom Stewart and Justin Lemberg unearthed material that we now know to be a relative of Tiktaalik's. 

Meet Qikiqtania wakei, a fishapod and close relative to our dear tetrapods — and cousin to Tiktaalik — who shares features in the flattened triangular skull, shoulders and elbows in the fin. 

Qikiqtania (pronounced kick-kick-TAN-ee-ya)

But, and here’s the amazing part, its upper arm bone (humerus) is specialised for open water swimming, not walking. 

The story gets wilder when we look at Qikiqtania’s position on the evolutionary tree— all the features for this type of swimming are newly evolved, not primitive. 

This means that Qikiqtania secondarily reentered open water habitats from ancestors that had already had some aspect of walking behaviour. 

And, this whole story was playing out 365 million years ago — the transition from water to land was going both ways in the Devonian.

Why is this exciting? You and I descend from those early tetrapods. We share the legacy of their water-to-land transition and the wee bony bits in their wrists and paddles that evolved to become our hands. I know, mindblowing!

Thomas Stewart and Justin Lemberg put in thousands of hours bringing Qikiqtania to life. 

The analysis consisted of a long path of wild events— from a haphazard moment when it was first spotted, a random collection of a block that ended up containing an articulated fin, to a serendipitous discovery three days before Covid lockdowns in March 2020.

Both teams acknowledge the profound debt owed to the individuals, organizations and indigenous communities where they had the privilege to work — Grise Fiord and Resolute Bay— Ellesmere Island in Nunavut, the largest and northernmost territory of Canada. 

Part of that debt is honoured in the name chosen for this new miraculous species. 

Aerial View of Ellesmere Island

The generic name, Qikiqtania (pronounced kick-kick-TAN-ee-ya), is derived from the Inuktitut words Qikiqtaaluk and Qikiqtani which are the traditional place name of the region where the fossil was discovered. 

The specific name, wakei, is in memory of the evolutionary biologist David Wake — colleague, mentor and friend. 

He was a professor of integrative biology and Director and curator of herpetology at the Museum of Vertebrate Zoology at the University of California, Berkeley who passed away in April 2021. 

Wake is known for his work on the biology and evolution of salamanders and vertebrate evolutionary biology. 

If you look at the photo on the left you can imagine visiting these fossil localities in Canada's far north.

Qikiqtania was found on Inuit land and belongs to the community. Thomas Stewart and his colleagues were able to conduct this research because of the generosity and support of individuals in the hamlets of Resolute Bay and Grise Fiord, the Iviq Hunters and Trappers of Grise Fiord, and the Department of Heritage and Culture, Nunavut.

To them, on behalf of the larger scientific community — Nakurmiik. Thank you! 

Here is the link to Tom Stewart's article in The Conversation & paper in Nature that dropped yesterday:

1. Stewart, Thomas A.; Lemberg, Justin B.; Daly, Ailis; Daeschler, Edward B.; Shubin, Neil H. (2022-07-20). "A new elpistostegalian from the Late Devonian of the Canadian Arctic". Nature. doi:10.1038/s41586-022-04990-w. ISSN 0028-0836.
2. Stewart, Thomas. "Meet Qikiqtania, a fossil fish with the good sense to stay in the water while others ventured onto land" The Conversation. Retrieved 2022-07-20.

Image One: An artist’s vision of Qikiqtania enjoying its fully aquatic, free-swimming lifestyle. Alex Boersma, CC BY-ND

Image Two: A new elpistostegalian from the Late Devonian of the Canadian Arctic, T. A. Stewart, J. B. Lemberg, A. Daly, E. B. Daeschler, & N. H. Shubin.

A huge shout out to the deeply awesome Neil Shubin who shared that the paper had been published and offered his insights on what played out behind the scenes!

WASHINGTON STATE PALEONTOLOGY

North Cascades National Park, Washington State, USA

Over vast expanses of time, powerful tectonic forces have massaged the western edge of the continent, smashing together a seemingly endless number of islands to produce what we now know as North America and the Pacific Northwest.

Washington is home to a wide variety of fossils—from new species of fossil crabs to marine mollusks and the fossil palm fronds that symbolize the Chuckanut formation.

We also find fossil whales, bird trackways, fossil sockeye salmon, mammal footprints, mammoth bones & the trace fossil remains of ancient rhino. In the time expanse in which we live our very short human lives, the Earth's crust appears permanent.

A fixed outer shell – terra firma. Aside from the rare event of an earthquake or the eruption of Mount St. Helen’s in 1980, our world seems unchanging, the landscape constant. In fact, it has been on the move for billions of years and continues to shift each day. As the earth’s core began cooling, some 4.5 billion years ago, plates, small bits of continental crust, have become larger and smaller as they are swept up in or swept under their neighbouring plates. Large chunks of the ocean floor have been uplifted, shifted and now find themselves thousands of miles in the air, part of mountain chains far from the ocean today or carved by glacial ice into valleys and basins.

Two hundred million years ago, Washington was two large islands, bits of the continent on the move westward, eventually bumping up against the North American continent and calling it home. Even with their new fixed address, the shifting continues; the more extreme movement has subsided laterally and continues vertically. The upthrusting of plates continue to move our mountain ranges skyward, the path of least resistance.

Fossil Palm Front, Washington State

This dynamic movement has created the landscape we see today and helped form the fossil record that tells much of Washington’s relatively recent history – the past 50 million years. Chuckanut Drive is much younger than other parts of Washington.

The fossils found there lived and died some 40-55 million years ago, very close to where they are now, but in a much warmer, swampy setting. The exposures of the Chuckanut Formation were once part of a vast river delta; imagine, if you will, the bayou country of the Lower Mississippi.

The siltstones, sandstones, mudstones and conglomerates of this formation were laid down about 40-54 million years ago during the Eocene epoch, a time of luxuriant plant growth in the subtropical flood plain that covered much of the Pacific Northwest.

This ancient wetland provided ideal conditions to preserve the many trees, shrubs, and plants that thrived here. Plants are important in the fossil record because they are more abundant and can give us a lot of information about climate, temperature, the water cycle, and humidity of the region. The Chuckanut flora is made up predominantly of plants whose modern relatives live in tropical areas such as Mexico and Central America.

Shore Bird Trackway, Washington State

While less abundant, evidence of the animals that called this ancient swamp home are also found here. Rare bird, reptile, and mammal tracks have been immortalized in the outcrops of the Chuckanut Formation.

Tracks of a type of archaic mammal of the Orders Pantodonta or Dinocerata (blunt foot herbivores), footprints from a small shorebird, and tracks from an early equid or webbed bird track give evidence to the vertebrates that inhabited the swamps, lakes and riverways of the Pacific Northwest 50 million years ago.

Fossil mammals from Washington do get most of the press. The movement of these celebrity vertebrates captured in the soft mud on the banks of a river, one of the depositional environments favourable for track preservation.

The bone record is actually far less abundant than the plant record, except near shell middens, given the preserving qualities of calcium and an alkaline environment. While calcium-rich bones and teeth fossilize well, they often do not get laid down in a situation that makes this possible. Hence the terrestrial paleontological record of Washington State at sites like Chuckanut is primarily made up of plant material.

FRACTAL BUILDING: AMMONITES

Argonauticeras besairei, Collection of José Juárez Ruiz.

An exceptional example of fractal building of an ammonite septum, in this clytoceratid Argonauticeras besairei from the awesome José Juárez Ruiz.

Ammonites were predatory, squid-like creatures that lived inside coil-shaped shells.

Like other cephalopods, ammonites had sharp, beak-like jaws inside a ring of squid-like tentacles that extended from their shells. They used these tentacles to snare prey, — plankton, vegetation, fish and crustaceans — similar to the way a squid or octopus hunt today.

Catching a fish with your hands is no easy feat, as I'm sure you know. But the Ammonites were skilled and successful hunters. They caught their prey while swimming and floating in the water column. Within their shells, they had a number of chambers, called septa, filled with gas or fluid that were interconnected by a wee air tube. By pushing air in or out, they were able to control their buoyancy in the water column.

They lived in the last chamber of their shells, continuously building new shell material as they grew. As each new chamber was added, the squid-like body of the ammonite would move down to occupy the final outside chamber.

They were a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids — octopuses, squid, and cuttlefish) than they are to shelled nautiloids such as the living Nautilus species.

The Ammonoidea can be divided into six orders:

• Agoniatitida, Lower Devonian - Middle Devonian
• Clymeniida, Upper Devonian
• Goniatitida, Middle Devonian - Upper Permian
• Prolecanitida, Upper Devonian - Upper Triassic
• Ceratitida, Upper Permian - Upper Triassic
• Ammonitida, Lower Jurassic - Upper Cretaceous

Ammonites have intricate and complex patterns on their shells called sutures. The suture patterns differ across species and tell us what time period the ammonite is from. If they are geometric with numerous undivided lobes and saddles and eight lobes around the conch, we refer to their pattern as goniatitic, a characteristic of Paleozoic ammonites.

If they are ceratitic with lobes that have subdivided tips; giving them a saw-toothed appearance and rounded undivided saddles, they are likely Triassic. For some lovely Triassic ammonites, take a look at the specimens that come out of Hallstatt, Austria and from the outcrops in the Humboldt Mountains of Nevada.

Hoplites bennettiana (Sowby, 1826).

If they have lobes and saddles that are fluted, with rounded subdivisions instead of saw-toothed, they are likely Jurassic or Cretaceous. If you'd like to see a particularly beautiful Lower Jurassic ammonite, take a peek at Apodoceras. Wonderful ridging in that species.

One of my favourite Cretaceous ammonites is the ammonite, Hoplites bennettiana (Sowby, 1826). This beauty is from Albian deposits near Carrière de Courcelles, Villemoyenne, near la région de Troyes (Aube) Champagne in northeastern France.

At the time that this fellow was swimming in our oceans, ankylosaurs were strolling about Mongolia and stomping through the foliage in Utah, Kansas and Texas. Bony fish were swimming over what would become the strata making up Canada, the Czech Republic and Australia. Cartilaginous fish were prowling the western interior seaway of North America and a strange extinct herbivorous mammal, Eobaatar, was snuffling through Mongolia, Spain and England.

In some classifications, these are left as suborders, included in only three orders: Goniatitida, Ceratitida, and Ammonitida. Once you get to know them, ammonites in their various shapes and suturing patterns make it much easier to date an ammonite and the rock formation where is was found at a glance.

Ammonites first appeared about 240 million years ago, though they descended from straight-shelled cephalopods called bacrites that date back to the Devonian, about 415 million years ago, and the last species vanished in the Cretaceous–Paleogene extinction event.

They were prolific breeders that evolved rapidly. If you could cast a fishing line into our ancient seas, it is likely that you would hook an ammonite, not a fish. They were prolific back in the day, living (and sometimes dying) in schools in oceans around the globe. We find ammonite fossils (and plenty of them) in sedimentary rock from all over the world.

In some cases, we find rock beds where we can see evidence of a new species that evolved, lived and died out in such a short time span that we can walk through time, following the course of evolution using ammonites as a window into the past.

For this reason, they make excellent index fossils. An index fossil is a species that allows us to link a particular rock formation, layered in time with a particular species or genus found there. Generally, deeper is older, so we use the sedimentary layers rock to match up to specific geologic time periods, rather the way we use tree-rings to date trees. A handy way to compare fossils and date strata across the globe.

References: Inoue, S., Kondo, S. Suture pattern formation in ammonites and the unknown rear mantle structure. Sci Rep 6, 33689 (2016). https://doi.org/10.1038/srep33689
https://www.nature.com/articles/srep33689?fbclid=IwAR1BhBrDqhv8LDjqF60EXdfLR7wPE4zDivwGORTUEgCd2GghD5W7KOfg6Co#citeas

Photo: Hoplites Bennettiana from near Troyes, France. Collection de Christophe Marot

ANCIENT OCTOPUS: KEUPPIA

A sweet as you please example of Keuppia levante (Fuchs, Bracchi & Weis, 2009), an extinct genus of octopus that swam our ancient seas back in the Cretaceous. 

The dark black and brown area you see here is his ink sac which has been preserved for a remarkable 95 million years.

This cutie is in the family Palaeoctopodidae, and one of the earliest representatives of the order Octopoda — and perhaps my favourite fossil. It was this perfect specimen that inspired the logo for the Fossil Huntress brand.  

These ancient marine beauties are in the class Cephalopoda making them relatives of our modern octopus, squid and cuttlefish.

There are two species of Keuppia, Keuppia hyperbolaris and Keuppia levante, both of which we find as fossils. We find their remains, along with those of the genus Styletoctopus, in Cretaceous-age Hâqel and Hjoula localities in Lebanon. 

For many years, Palaeoctopus newboldi (Woodward, 1896) from the Santonian limestones at Sâhel Aalma, Lebanon, was the only known pre‐Cenozoic coleoid cephalopod believed to have an unambiguous stem‐lineage representative of Octobrachia fioroni. 

With the unearthing of some extraordinary specimens with exquisite soft‐part preservation in the Lebanon limestones, our understanding of ancient octopus morphology has blossomed. The specimens are from the sub‐lithographical limestones of Hâqel and Hâdjoula, in northwestern Lebanon. These localities are about 15 km apart, 45 km away from Beirut and 15 km away from the coastal city of Jbail. Fuchs et al. put a nice little map in their 2009 paper that I've included and referenced here.

Palaeoctopus newboldi had a spherical mantle sac, a head‐mantle fusion, eight equal arms armed with suckers, an ink sac, a medially isolated shell vestige, and a pair of (sub‐) terminal fins. The bipartite shell vestige suggests that Palaeoctopus belongs to the octopod stem‐lineage, as the sister taxon of the Octopoda, the Cirroctopoda, is characterized by an unpaired clasp‐like shell vestige (Engeser 1988; Haas 2002; Bizikov 2004).

It is from the comparisons of Canadian fauna combined with those from Lebanon and Japan that things really started to get interesting with Octobrachia. Working with fossil specimens from the Campanian of Canada, Fuchs et al. (2007a ) published on the first record of an unpaired, saddle‐shaped shell vestige that might have belonged to a cirroctopod. 

Again from the Santonian–Campanian of Canada and Japan, Tanabe et al. (2008) reported on at least four different jaw morphotypes. Two of them — Paleocirroteuthis haggarti (Tanabe et al., 2008) and Paleocirroteuthis Pacifica  (Tanabe et al ., 2008) — have been interpreted as being of cirroctopod type, one of octopod type, and one of uncertain octobrachiate type. 

Interestingly Fuchs et al. have gone on to describe the second species of Palaeoctopus, the Turonian Palaeoctopus pelagicus from limestones at Vallecillo, Mexico. While more of this fauna will likely be recovered in time, their work is based solely on a medially isolated shell vestige.

Five new specimens have been found in the well-known Upper Cenomanian limestones at Hâqel and Hâdjoula in Lebanon that can be reliably placed within the Octopoda. Fuchs et al. described these exceptionally well‐preserved specimens and discuss their morphology in the context of phylogeny and evolution in their 2008 paper (2009 publishing) in the Palaeontology Association Journal, Volume 51, Issue 1.

The presence of a gladius vestige in this genus shows a transition from squid to octopus in which the inner shell has divided into two parts in early forms to eventually be reduced to lateralized stylets, as can be seen in Styletoctopus.

The adorable fellow you see here with his remarkable soft-bodied preservation and inks sack and beak clearly visible is Keuppia levante. He hails from Late Cretaceous (Upper Cenomanian) limestone deposits near Hâdjoula, northwestern Lebanon. The vampyropod coleoid, Glyphiteuthis abisaadiorum n. sp. is also found at this locality. This specimen is about 5 cm long.

Fuchs, D.; Bracchi, G.; Weis, R. (2009). "New octopods (Cephalopoda: Coleoidea) from the Late Cretaceous (Upper Cenomanian) of Hâkel and Hâdjoula, Lebanon". Palaeontology. 52: 65–81. doi:10.1111/j.1475-4983.2008.00828.x.

Photo one: Fossil Huntress. Figure Two: Topographic map of north‐western Lebanon with the outcrop area in the upper right-hand corner. Fuchs et al, 2009.  

UNESCOCERATOPS BY JULIUS CSOTONYI

Unescoceratops koppelhusae, Julius Csotonyi

A very sweet small leptoceratopsid dinosaur, Unescoceratops koppelhusae — a new species in the collections of the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta.

The colourful and beautifully detailed painting you see here is by the very talented Julius Csotonyi who captured the magnificence of form, texture and palette to bring this small leptoceratopsid dinosaur to life.

The Royal Tyrrell Museum of Palaeontology, named in honour of Joseph Burr Tyrrell, is a palaeontology museum and research facility in Drumheller, Alberta, Canada. 

This jaw is the holotype specimen of this small leptoceratopsid dinosaur. Only a handful of isolated fossils have been found from this species, including a jaw that is the holotype specimen now in collections at the Royal Tyrell. 

The Royal Tyrrell Museum of Palaeontology, named in honour of Joseph Burr Tyrrell, is a palaeontology museum and research facility in Drumheller, Alberta, Canada. 

Unescoceratops koppelhusae, RTMP Collections

The rusty chocolate jaw bone you see here is the puzzle piece that helped all of the research come together and help us to better understand more about the diminutive leptoceratopsid dinosaurs from Alberta. 

The Cleveland Museum of Natural History's Michael Ryan and David Evans of the Royal Ontario Museum in Toronto recently determined that the specimen was a new genus and species. 

Unescoceratops is a genus of leptoceratopsid ceratopsian dinosaurs known from the Late Cretaceous (about 76.5-75 million years ago) of Alberta, Canada. Unescoceratops is thought to have been between one and two meters long and less than 91 kilograms. A plant-eater, its teeth were the roundest of all Leptocertopsids.

Dinosaur Provincial Park, Alberta, Canada

The genus name acknowledges the UNESCO  World Heritage Site, Dinosaur Provincial Park, where the fossil was found. 

In addition to its particularly beautiful scenery, Dinosaur Provincial Park – located at the heart of the province of Alberta's badlands – is unmatched in terms of the number and variety of high-quality specimens.

To date, they represent more than 44 species, 34 genera and 10 families of dinosaurs, dating back 75-77 million years. This provides us with remarkable insight into life millions of years ago.

The park contains exceptional riparian habitat features as well as badlands of outstanding aesthetic value.

The creamy honey, beige and rust coloured hills around the fossil locality are outstanding examples of major geological processes and fluvial erosion patterns in semi-arid steppes — think glorious! 

The scenic badlands stretch along 26 kilometres of high quality and virtually undisturbed riparian habitat, presenting a landscape of stark but exceptional natural beauty.

The species name honours Dr. Eva Koppelhus, who has made significant contributions to vertebrate palaeontology and palynology. 

The genus is named to honour the UNESCO World Heritage Site designation for the locality where the specimen was found and from the Greek “ceratops,” which means 'horned face'. 

Dr Michael Ryan explained that he meant to honour UNESCO's efforts to increase understanding of natural history sites around the world.

© Julius T. Csotonyi An illustration of Unescoceratops koppelhusae, a plant-eating dinosaur from the Late Cretaceous period that lived approximately 75 million years ago shared with his gracious permission. 

ABOUT THE ARTIST

Dr. Julius Csotonyi is a Vancouver-based scientific illustrator and natural history fine artist. He is a featured paleoartist on Season One and Season Two of BC's Fossil Bounty. Julius has a scientific background in ecology (MSc) and microbiology (PhD) which has taken him to study sensitive ecosystems, from sand dunes in the Rocky Mountain parks to hydrothermal vents at the bottom of the Pacific Ocean. 

These experiences have fuelled his strong resolve to work toward preserving our Earth’s biota. Painting biological subjects is one means that he uses to both enhance public awareness of biological diversity and to motivate concern for its welfare.   

He paints murals and panels that have appeared in numerous museums including the Smithsonian’s National Museum of Natural History, press release images for scientific publications, books, stamp sets — including the outstanding 2018 “Sharks of Canada” set for Canada Post — and coins for the Royal Canadian Mint. To view more of Julius Csotonyi's exquisite work visit: https://csotonyi.com/

JUVENILE HAMITES SUBROTUNDUS

A tremendously delicate juvenile Hamites subrotundus (J. Sowerby 1814) from Upper Albian outcrops in Mallorca, the largest of the three Balearic Islands in the Mediterranean at more than 3,600 square kilometers. 

Mallorca has been home to various inhabitants for thousands of years. Sitting some 200 kilometers off Spain’s southeastern, it is a idyllic setting for exploring the rich human and geologic history of this part of the world.

The island is made up of dolomite and limestone from a huge expanse of time, the Mesozoic and Cenozoic—170 million to 10 million years ago— and bookended by two parallel mountain chains top and bottom on its southern and northern coasts. 

As you walk the mountain passes northwest to northeast, you stroll across Miocene deposits 20 million to 13 million years ago that speak of the time in our Earth's recent past when part of the African continent collided with Europe. 

It is famous for its limestone mountains and Roman and Moorish remains. As you can see here, it is also home to some rather nice fossils including this specimen of Hamites subrotundus.

While H. subrotundus is generally a Middle Albian species, this specimen was found in the lower part of Upper Albian in the Cristatum zone by José Juárez Ruiz. José had to piece this lovely together from seven fragments. His labour of love was worth the effort. The final piece is sheer perfection and a beautiful specimen approximately 2.5 cm long.

Mallorca and the other Balearic Islands are geologically an extension of the Baetic Cordillera mountain chain of western Andalusia that extends to Murcia and Valencia. 

They are made up of sediments deposited in the Tethys Sea during the Mesozoic.

Exploring the islands, you can collect from deposits from the Triassic, Cretaceous, Jurassic, and Neogene periods. 

The limestone outcrops contain lovely examples of foraminifers—mainly the species Globigerina.

We also see lovely examples of Hamites (Hamites) subrotundus in the Euhoplites loricatus zone; Euhoplites meandrinus subzone from the Middle Albian (Lower Gault) of Folkestone, Kent, UK. 

Photo, preparation and in the collection of the deeply awesome José Juárez Ruiz. Wright C. W. 1996. Treatise on Invertebrate Paleontology (Part L Mollusca 4 Revised) Volume 4: Cretaceous Ammonoidea

JURASSIC SEA URCHIN: AM'DA'MA

This lovely little biscuit is a Holectypus sea urchin from 120 million-year-old deposits from the Lagniro Formation of Madagascar.

The specimen you see here is in the collections of my beautiful friend Ileana. She and I were blessed to meet in China many years ago and formed an unbreakable bond that happens so few times in one's life. 

Holectypus are a genus of extinct echinoids related to modern sea urchins and sand dollars. They were abundant from the Jurassic to the Cretaceous (between 200 million and 65.5 million years ago).

This specimen is typical of Holectypus with his delicate five-star pattern adorning a slightly rounded test and flattened bottom. The specimen has been polished and was harvested both for its scientific and aesthetic value. 

I have many wonderful memories of collecting their modern cousins that live on the north end of Vancouver Island and along the beaches of Balaklava Island. In the Kwak̓wala language of the Kwakiutl or Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, sea urchins are known as a̱m'da̱'ma and it is this name that I hear in my head when I think of them.

In echinoids, the skeleton is almost always made up of tightly interlocking plates that form a rigid structure or test — in contrast with the more flexible skeletal arrangements of starfish, brittle stars, and sea cucumbers. Test shapes range from nearly globular, as in some sea urchins, to highly flattened, as in sand dollars. 

Sea Urchin Detail

Living echinoids are covered with spines, which are movable and anchored in sockets in the test. These spines may be long and prominent, as in typical sea urchins and most have lovely raised patterns on their surface. 

In sand dollars and heart urchins, however, the spines are very short and form an almost felt-like covering. The mouth of most echinoids is provided with five hard teeth arranged in a circle, forming an apparatus known as Aristotle’s lantern.

Echinoids are classified by the symmetry of the test, the number and arrangement of plate rows making up the test, and the number and arrangement of respiratory pore rows called petals. Echinoids are divided into two subgroups: regular echinoids, with nearly perfect pentameral (five-part) symmetry; and irregular echinoids with altered symmetry.

Because most echinoids have rigid tests, their ability to fossilize is greater than that of more delicate echinoderms such as starfish, and they are common fossils in many deposits. The oldest echinoids belong to an extinct regular taxon called the Echinocystitoidea. 

They first appeared in the fossil record in the Late Ordovician. Cidaroids or pencil urchins appear in the Mississippian (Early Carboniferous) and were the only echinoids to survive the mass extinction at the Permo-Triassic boundary. Echinoids did not become particularly diverse until well after the Permo-Triassic mass extinction event, evolving the diverse forms we find them in today. 

True sea urchins first appear in the Late Triassic, cassiduloids in the Jurassic, and spatangoids or heart urchins in the Cretaceous. Sand dollars, a common and diverse group today, do not make an appearance in the fossil record until the Paleocene. They remain one of my favourite echinoderms and stand tall amongst the most pleasing of the invertebrates.

CHUCKANUT DRIVE: EOCENE TROPICAL PARADISE

A trip along Chuckanut Drive, in northwestern Washington is a chance to view incredible diversity from sea to sky.

An amazing array of plants and animals call this coastline home. 

For the fossil enthusiast, it is a chance to slip back in time and have a bird’s eye view of a tropical paradise preserved in the Eocene strata of various fossil sites. 

Snug up against the Pacific Ocean, this 6000m thick exposure yields a vast number of tropical and flowering plants that you might see in Mexico today. Easily accessible by car, this rich natural playground makes for an enjoyable daytrip just one hour south of the US Border.

Over vast expanses of time, powerful tectonic forces have massaged the western edge of the continent, smashing together a seemingly endless number of islands to produce what we now know as North America and the Pacific Northwest. Intuition tells us that the earth’s crust is a permanent, fixed outer shell – terra firma.

Aside from the rare event of an earthquake or the eruption of Mount St. Helen’s, our world seems unchanging, the landscape constant. In fact, it has been on the move for billions of years and continues to shift each day. As the earth’s core began cooling, some 4.5 billion years ago, plates, small bits of continental crust, have become larger and smaller as they are swept up in or swept under their neighboring plates. 

Large chunks of the ocean floor have been uplifted, shifted and now find themselves thousands of miles in the air, part of mountain chains far from the ocean today or carved by glacial ice into valleys and basins.

Two hundred million years ago, Washington was two large islands, bits of continent on the move westward, eventually bumping up against the North American continent and calling it home. Even with their new fixed address, the shifting continues; the more extreme movement has subsided laterally and continues vertically. 

The upthrusting of plates continues to move our mountain ranges skyward – the path of least resistance. This dynamic movement has created the landscape we see today and helped form the fossil record that tells much of Washington’s relatively recent history – the past 50 million years.

Chuckanut Drive is much younger than other parts of Washington. The fossils found there lived and died some 40-55 million years ago, very close to where they are now, but in a much warmer, swampy setting. The exposures of the Chuckanut Formation were once part of a vast river delta; imagine, if you will, the bayou country of the Lower Mississippi. The siltstones, sandstones, mudstones and conglomerates of the Chuckanut Formation were laid down about 40-54 million years ago during the Eocene epoch, a time of luxuriant plant growth in the subtropical flood plain that covered much of the Pacific Northwest.

This ancient wetland provided ideal conditions to preserve the many trees, shrubs & plants that thrived here. Plants are important in the fossil record because they are more abundant and can give us a lot of information about climate, temperature, the water cycle and humidity of the region. 

The Chuckanut flora is made up predominantly of plants whose modern relatives live in tropical areas such as Mexico and Central America. If you are interesting in viewing a tropical paradise in your own backyard, look no further than the Chuckanut. 

Images and tag lines: Glyptostrobus, the Chinese swamp cypress, is perhaps the most common plant found here. Also abundant are fossilized remains of the North American bald cypress, Taxodium; Metasequoia (dawn redwood), Lygodium (climbing fern), large Sabal (palm) and leaves from a variety of broad leaf angiosperm plants such as (witch hazel), Laurus (laurel), Ficus (fig) and Platanus (sycamore), and several other forms.

While less abundant, evidence of the animals that called this ancient swamp home are also found here. Rare bird, reptile, and mammal tracks have been immortalized in the outcrops of the Chuckanut Formation. Tracks of a type of archaic mammal of the Orders Pantodonta or Dinocerata (blunt foot herbivores), footprints from a small shorebird, and tracks from an early equid or webbed bird track give evidence to the vertebrates that inhabited the swamps, lakes and river ways of the Pacific Northwest 50 million years ago.

The movement of these celebrity vertebrates was captured in the soft mud on the banks of a river, one of the only depositional environments favorable for track preservation.

INKY BEAUTY: AMMONITE OF PONGO DE MANSERICHE

This inky beauty is Prolyelliceras ulrichi (Knechtel, 1947) a fast-moving nektonic carnivorous ammonite from Cretaceous lithified, black, carbonaceous limestone outcrops in the Pongo de Manseriche gorge in northwest Peru. 

If you look closely, you can see that this specimen shows a pathology, a slight deviation to the side of the siphonal of the ammonite. We see Prolyelliceras from the Albian to Middle Albian from five localities in Peru.

The canyons of the Amazon River system in the eastern ranges of the Andes of Peru are known by the Indian name pongo. 

The most famous of these is the Pongo de Manseriche, cut by the Marañon River through the eastern range of the Andes, where it emerges from the cordillera into the flat terrane of the Amazon Basin. The fossil exposures here are best explored by boat. The reality of the collecting is similar to the imagined. I was chatting with Betty Franklin, VIPS, about this. They float along and pick up amazing specimen after amazing specimen. When the water rises, the ammonites are aided in their erosion out of the cliffs.  

The Pongo de Manseriche lies nearly 500 miles upstream from Iquitos, and consequently nearly 3,000 miles above the mouth of the Amazon River. It is situated in the heart of the montaña, in a vast region the ownership of which has long been in dispute between Peru and Ecuador, but over which neither country exercises any police or other governmental control. There is an ancient tradition of the indigenous people of the vicinity that one of their gods descended the Marañón and another ascended the Amazon to communicate with him. Together they opened the pass called the Pongo de Manseriche.

Reference: M. M. Knechtel. 1947. Cephalopoda. In: Mesozoic fossils of the Peruvian Andes, Johns Hopkins University Studies in Geology 15:81-139

W. J. Kennedy and H. C. Klinger. 2008. Cretaceous faunas from Zululand and Natal, South Africa. The ammonite subfamily Lyelliceratinae Spath, 1921. African Natural History 4:57-111. The beauty you see here is in the collection of the deeply awesome José Juárez Ruiz.

EXPLORING WRANGELLIA: HAIDA GWAII

Misty shores, moss covered forests, a rich cultural history, dappled light, fossils and the smell of salt air—these are my memories of Haida Gwaii.

The archipelago of Haida Gwaii lays at the western edge of the continental shelf due west of the central coast of British Columbia.

They form part of Wrangellia, an exotic tectonostratigraphic terrane that includes Vancouver Island, parts western British Columbia and Alaska.

The Geological Survey of Canada sponsored many expeditions to these remote islands and has produced numerous reference papers on this magnificent terrain, exploring both the geology and palaeontology of the area.

Joseph Whiteaves, the GSC's chief palaeontologist in Ottawa, published a paper in 1876 describing the Jurassic and Cretaceous faunas of Skidegate Inlet, furthering his reputation globally as both a geologist, palaeontologist as well as a critical thinker in the area of science.

The praise was well-earned and foreshadowed his significant contributions to come. Sixteen years later, he wrote up and published his observations on a strange Mount Stephen fossil that resembled a kind of headless shrimp with poorly preserved appendages. 

Because of the unusual pointed shape of the supposed ventral appendages and the position of the spines near the posterior of the animal, Whiteaves named it Anomalocaris canadensis. The genus name "Anomalocaris" means "unlike other shrimp" and the species name "canadensis" refers to the country of origin.

Whiteaves work on the palaeontology of Haida Gwaii provided excellent reference tools, particularly his work on the Cretaceous exposures and fauna that can be found there.

One of our fossil field trips was to the ruggedly beautiful Cretaceous exposures of Lina Island. We had planned this expedition as part of our “trips of a lifetime.” 

Both John Fam, the Vice Chair of the Vancouver Paleontological Society and Dan Bowen, the Chair of both the British Columbia Paleontological Alliance and Vancouver Island Palaeontological Society, can be congratulated for their efforts in researching the area and ably coordinating a warm welcome by the First Nations community and organizing fossil field trips to some of the most amazing fossil localities in the Pacific Northwest.

With great sandstone beach exposures, the fossil-rich (Albian to Cenomanian) Haida formation provided ample specimens, some directly in the bedding planes and many in concretion. Many of the concretions contained multiple specimens of typical Haida Formation fauna, providing a window into this Cretaceous landscape.

It is always interesting to see who was making a living and co-existing in our ancient oceans at the time these fossils were laid down. We found multiple beautifully preserved specimens of the spiny ammonite, Douvelleiceras spiniferum along with Brewericeras hulenense, Cleoniceras perezianum and many cycads in concretion.

Douvelliceras spiniferum, Cretaceous Haida Formation

Missing from this trip log are tales of Rene Savenye, who passed away in the weeks just prior. While he wasn't there in body, he was with us in spirit. I thought of him often on the mist-shrouded days of collecting. 

Many of the folk on who joined me on those outcrops were friends of Rene's and would go on to receive the Rene Savenye Award for their contributions to palaeontology. There is a certain poetry in that. 

The genus Douvilleiceras range from Middle to Late Cretaceous and can be found in Asia, Africa, Europe and North and South America. 

We have beautiful examples in the early to mid-Albian from the archipelago of Haida Gwaii in British Columbia. Joseph F. Whiteaves was the first to recognize the genus from Haida Gwaii when he was looking over the early collections of James Richardson and George Dawson.

My collections from Haida Gwaii will all be lovingly prepped and donated to the Haida Gwaii Museum in Skidegate, British Columbia.