FERNIE'S GIANT AMMONITE: TITANITES

Titanites occidentalis, Fernie Ammonite

The Fernie ammonite, Titanites occidentalis, from outcrops on Coal Mountain near Fernie, British Columbia, Canada. 

This beauty is the remains of a carnivorous cephalopod within the family Dorsoplanitidae that lived and died in a shallow sea some 150 million years ago.

If you would like to get off the beaten track and hike up to see this ancient beauty, you will want to head to the town of Fernie in British Columbia close to the Alberta border. 

This is the traditional territory of the the Yaq̓it ʔa·knuqⱡi ‘it First Nation who have lived here since time immemorial. There was some active logging along the hillside in 2021, so if you are looking at older directions on how to get to the site be mindful that many of the trailheads have been altered and a fair bit of bushwhacking will be necessary to get to the fossil site proper. That being said, the loggers from CanWel may have clear-cut large sections of the hillside but they did give the ammonite a wide berth and have left it intact.

Wildsight, a non-profit environmental group out of the Kimberly Cranbrook area has been trying to gain grant funding to open up the site as an educational hike with educational signage for folks visiting the Fernie area. It is likely the province of British Columbia would top up those funds if they are able to place the ammonite under the Heritage Conservation Act. CanWel would remain the owners of the land but the province could assume the liability for those visiting this iconic piece of British Columbia's palaeontological history. 

Driving to the trail base is along an easy access road just east of town along Fernie Coal Road. There are some nice exposures of Cretaceous plant material on the north side (left-hand side) of the road as you head from Fernie towards Coal Creek. I recently drove up to Fernie to look at Cretaceous plant material and locate the access point to the now infamous Late Jurassic (Tithonian) Titanites (S.S. Buckman, 1921) site. While the drive out of town is on an easy, well-maintained road, the slog up to the ammonite site is often a wet, steep push. This time of year, it is buried in a blanket of crisp, white snow.

Fernie, British Columbia, Canada

The first Titanites occidentalis was about one-third the size and was incorrectly identified as Lytoceras, a fast-moving nektonic carnivore. The specimen you see here is significantly larger at 1.4 metres (about four and a half feet) and rare in North America. 

Titanites occidentalis, the Western Giant, is the second known specimen of this extinct fossil species. 

The first was discovered in 1947 in nearby Coal Creek by a British Columbia Geophysical Society mapping team. When they first discovered this marine fossil high up on the hillside, they could not believe their eyes — both because it is clearly marine at the top of a mountain and the sheer size of this ancient beauty.

In the summer of 1947, on a warm summer's day, a field crew was mapping coal outcrops for the BC Geological Survey (GSC) east of Fernie. One of the students reported finding “a fossil truck tire.” Fair enough. The similarity of size and optics are pretty close to your average Goodridge. 

A few years later, GSC Paleontologist Hans Frebold described and named the fossil Titanites occidentalis after the large Jurassic ammonites from Dorset, England. The name comes from Greek mythology. Tithonus, as you may recall, was the Prince of Troy. He fell in love with Eos, the Greek Goddess of the Dawn. Eos begged Zeus to make her mortal lover immortal. Zeus granted her wish but did not grant Tithonus eternal youth. He did indeed live forever — ageing hideously. Ah, Zeus, you old trickster. It is a clever play on time placement. Dawn is the beginning of the day and the Tithonian being the latest age of the Late Jurassic. Clever Hans!

HIKING TO THE FERNIE AMMONITE

From the town of Fernie, British Columbia, head east along Coal Creek Road towards Coal Creek. The site is 3.81 km from the base of Coal Creek Road to the trailhead as the crow flies. I have mapped it here for you in yellow and added the wee purple GPS marker for the ammonite site proper. There is a nice, dark grey to black roadcut exposure of Cretaceous plants on the north side of the dirt road that is your cue to pull over and park.  

You access what is left of the trailhead on the south side of the road. You will need to cross the creek to begin your ascent. There is no easy way across the creek and you'll want to tackle this one with a friend when the water level is low. 

The beginning of the trail is not clear but a bit of searching will reveal the trailhead with its telltale signs of previous hikers. This is a moderate 6.3-kilometre hike up & back bushwhacking through scrub and fallen trees. Heading up, you will make about a 246-metre elevation gain. You will likely not have a cellular signal up here but if you download the Google Map to your mobile, you will have GPS to guide you. The area has been recently logged so much of the original trail has been destroyed. There may now be easier vehicle access up the logging roads but I have not driven them since the logging and new road construction.

If you are coming in from out of town, the closest airport is Cranbrook. Then it is about an hour and change to Fernie and another 15-minutes or so to park near the site.

You will want to leave your hammers with your vehicle (no need to carry the weight and this lovely should never be struck with anything more than a raindrop) as this site is best enjoyed with a camera. 

This is a site you will want to wear hiking boots to access. Know that these will get wet as you cross the creek. 

If you would like to see the ammonite but are not keen on the hike, a cast has been made by fossil preparator Rod Bartlett is on display at the Courtenay Museum in Courtenay, Vancouver Island, Canada. 

Respect for the Land / Leave No Trace

As your feet move up the hillside, you can imagine this land 10,000 years ago, rising above great glaciers. Where footfalls trace the steps of those that came before you. This land has been home to the Yaq̓it ʔa·knuqⱡi ‘it First Nation and Ktunaxa or Kukin ʔamakis First Nations whose oral history have them living here since time immemorial. Like them, take only what you need and no more than the land offers — packing out anything that you packed in. 

Fernie Ammonite Palaeo Coordinates: 49°29'04"N 115°00'49"W

LOWER LIAS LYTOCERAS

A superbly prepped and extremely rare Lytoceras (Suess, 1865) ammonite found as a green ammonite nodule by Matt Cape in the Lower Lias of Dorset. 

Lytoceras are rare in the Lower Lias of Dorset — apart from the Belemnite Stone horizon — so much so that Paul Davis, whose skilled prep work you see here, initially thought it might be a Becheiceras hidden within the large, lumpy nodule. 

One of the reasons these lovelies are rarely found from here is that they are a Mediterranean Tethyian genus. The fossil fauna we find in the United Kingdom are dominated by Boreal Tethyian genera. 

We do find Lytoceras sp. in the Luridum subzone of the Pliensbachian showing that there was an influx of species from the Mediterranean realm during this time. This is the first occurrence of a Lytoceras that he has ever seen in a green nodule and Paul's seen quite a few. 

This absolutely cracking specimen was found and is in the collections of the awesome Matt Cape. Matt recognized that whatever was hidden in the nodule would take skilled and careful preparation using air scribes. Indeed it did. It took more than five hours of time and skill to unveil the lovely museum-worthy specimen you see here. 

We find Lytoceras in more than 1,000 outcrops around the globe ranging from the Jurassic through to the Cretaceous, some 189.6 to 109.00 million years ago. Once this specimen is fully prepped with the nodule material cut or scraped away, you can see the detailed crinkly growth lines or riblets on the shell and none of the expected coarse ribbing. 

Lytoceras sp. Photo: Craig Chivers

If you imagine running your finger along these, you would be tracing the work of decades of growth of these cephalopods. 

While we cannot know their actual lifespans, but we can make a healthy guess. 

The nautilus, their closest living cousins live upwards of 20 years — gods be good — and less than three years if conditions are poor.

The flanges, projecting flat ribs or collars, develop at the edge of the mouth border on the animal's mantle as they grow each new chamber. 

Each delicate flange grows over the course of the ammonites life, marking various points in time and life stages as the ammonite grew. There is a large variation within Lytoceras with regards to flanges. They provide both ornamentation and strength to the shell to protect it from water pressure as they moved into deeper seas.

The concretion prior to prep

This distinctive genus with its evolute shells are found in the Cretaceous marine deposits of: 

Antarctica (5 collections), Austria (19), Colombia (1), the Czech Republic (3), Egypt (2), France (194), Greenland (16), Hungary (25), Italy (11), Madagascar (2), Mexico (1), Morocco (4), Mozambique (1), Poland (2), Portugal (1), Romania (1), the Russian Federation (2), Slovakia (3), South Africa (1), Spain (24), Tanzania (1), Trinidad and Tobago (1), Tunisia (25); and the United States of America (17: Alaska, California, North Carolina, Oregon).

We also find them in Jurassic marine outcrops in:

Austria (15), Canada (9: British Columbia), Chile (6), France (181), Germany (11), Greenland (1), Hungary (189), India (1), Indonesia (1), Iran (1), Italy (50), Japan (14), Kenya (2), Luxembourg (4), Madagascar (2), Mexico (1), Morocco (43), New Zealand (15), Portugal (1), Romania (5), the Russian Federation (1), Slovakia (1), Spain (6), Switzerland (2), Tunisia (11), Turkey (12), Turkmenistan (1), Ukraine (5), the United Kingdom (12), United States (11: Alaska, California) — in at least 977 known collections. 

References:

Sepkoski, Jack (2002). "A compendium of fossil marine animal genera (Cephalopoda entry)". Bulletins of American Paleontology. 363: 1–560. Archived from the original on 2008-05-07. Retrieved 2017-10-18.

Paleobiology Database - Lytoceras. 2017-10-19.

Systematic descriptions, Mesozoic Ammonoidea, by W.J Arkell, Bernhard Kummel, and C.W. Wright. 1957. Treatise on Invertebrate Paleontology, Part L. Geological Society of America and University of Kansas press.

MEET FERGUSONITES HENDERSONAE

Fergusonites hendersonae (Longridge, 2008)

Meet Fergusonites hendersonae, a Late Hettangian (Early Jurassic) ammonite from the Taseko Lakes area of British Columbia, Canadian Rockies.

I had the very great honour of having this fellow, a new species of nektonic carnivorous ammonite, named after me by paleontologist Louse Longridge from the University of British Columbia. I'd met Louise as an undergrad and was pleased as punch to hear that she would be continuing the research by Dr. Howard Tipper.

We did several trips over the years up to the Taseko Lake area of the Rockies joined by many wonderful researchers from Vancouver Island Palaeontological Society and Vancouver Paleontological Society, as well as the University of British Columbia. Both Dan Bowen and John Fam were instrumental in planning those expeditions. We endured elevation sickness, rain, snow, grizzly bears and very chilly nights (we were sleeping right next to a glacier at one point) but were rewarded by the enthusiastic crew, helicopter rides (which really cut down the hiking time) excellent specimens and stunningly beautiful country. We were also blessed with excellent access as the area is closed to collecting except with a permit.

Reference: PaleoDB 157367 M. Clapham GSC C-208992, Section A 09, Castle Pass Angulata - Jurassic 1 - Canada, Longridge et al. (2008)

Full reference: L. M. Longridge, P. L. Smith, and H. W. Tipper. 2008. Late Hettangian (Early Jurassic) ammonites from Taseko Lakes, British Columbia, Canada. Palaeontology 51:367-404

PaleoDB taxon number: 297415; Cephalopoda - Ammonoidea - Juraphyllitidae; Fergusonites hendersonae Longridge et al. 2008 (ammonite); Average measurements (in mm): shell width 9.88, shell diameter 28.2; Age range: 201.6 to 196.5 Ma. Locality info: British Columbia, Canada (51.1° N, 123.0° W: paleo coordinates 22.1° N, 66.1° W)

PTEROSAURS AND SARODONTID FISH OF HORNBY ISLAND

Quetzalcoatlus

If you could travel through time and go back to observe our ancient skies, you would see massive pterosaurs — huge, winged flying reptiles of the extinct order Pterosauria — cruising along with you.

They soared our skies during most of the Mesozoic — from the late Triassic to the end of the Cretaceous (228 to 66 million years ago). 

By the end of the Cretaceous, they had grown to giants and one of their brethren, Quetzalcoatlus, a member of the family Azhdarchidae, boasts being the largest known flying animal that ever lived. They were the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger.

We divide their lineage into two major types: basal pterosaurs and pterodactyloids. Basal pterosaurs — also called 'non-pterodactyloid pterosaurs' or ‘rhamphorhynchoids’ — were smaller with fully toothed jaws and longish tails. Their wide wing membranes connected to their hind legs giving them some maneuverability on the ground, but with an awkward sprawling posture. Picture a bat trying to walk or crawl along — doable but painful to watch. They were better climbers with flexible joint anatomy and strong claws. Basal pterosaurs preferred to dine on insects and small vertebrates.

Later pterosaurs (pterodactyloids) evolved many sizes, shapes, and lifestyles. Pterodactyloids had narrower wings with free hind limbs, highly reduced tails, and long necks with large heads. On the ground, pterodactyloids walked better than their earlier counterparts, maneuvering all four limbs smoothly with an upright posture. 

They walked standing plantigrade on the hind feet and folding the wing finger upward to walk on the three-fingered "hand." These later pterosaurs were more nimble. They could take off from the ground, run and wade and swim. Their jaws had horny beaks and some of these later groups lacked the teeth of earlier lineages. Some groups developed elaborate head crests that were likely used to attract mates' sexy-pterosaur style.

So can we or have we found pterosaurs on Hornby Island? The short answer is yes.

Collishaw Point, known locally as Boulder Point, Hornby Island

Hornby Island is a lovely lush, island in British Columbia's northern Gulf Islands. It was formed from sediments of the upper Nanaimo Group which are also widely exposed on adjacent Denman Island and the southern Gulf Islands.

Peter Mustard, a geologist from the Geologic Survey of Canada, did considerable work on the geology of the island. It has a total stratigraphic thickness of 1350 m of upper Nanaimo Group marine sandstone, conglomerate and shale. 

These are partially exposed in the Campanian to the lower Maastrichtian outcrops at Collishaw Point on the northwest side of Hornby Island. Four formations underlie the island from oldest to youngest, and from west to east: the Northumberland, Geoffrey, Spray and Gabriola.

During the upper Cretaceous, between ~90 to 65 Ma, sediments derived from the Coast Belt to the east and the Cascades to the southeast poured seaward to the west and northwest into what was the large ancestral Georgia Basin. This major forearc basin was situated between Vancouver Island and the mainland of British Columbia. The rocks you find here originated far to the south in Baja California and are the right age and type of sediment for a pterosaur find. But are we California dreaming?

Upper Cretaceous Nanaimo Group Fossil Concretion

Well, truth be told, we were with one of the potential pterosaur finds from Hornby. 

It wasn't just hopeful thinking that had the west coast in a paleo uproar many ago when Sharon Hubbard of the Vancouver Island Palaeontological Society found what looked very much like a pterosaur.

Right time period. Right location. And, we have found them here in the past. Sandy McLachlan found the first definitive pterosaur, an azhdarchid, back in 2008.

But was Sharon's find a pterosaur? Victoria Arbour, a Canadian evolutionary biologist and palaeontologist working as a Natural Sciences and Engineering Research Council of Canada postdoctoral fellow at the University of Toronto and Royal Ontario Museum, certainly thought so. 

While Arbour is an expert on ankylosaurs, our lumbering armoured dinosaurs friends, she has studied pterosaurs and participated in the naming of Gwawinapterus from Hornby Island. But here's the thing — bony material encased in stone and let to cement for millions of years can be tricky.

While this fossil find was initially described as a very late-surviving member of the pterosaur group Istiodactylidae, further examination cast doubt on the identification. Once more detail was revealed the remains were published as being those of a saurodontid fish, an ambush predator with very sharp serrated teeth and elongate, torpedo-like bodies that grew up to two meters. 

Not a pterosaur but still a massively exciting find. Arbour was very gracious at the renaming, taking it in stride. She has since gone on to name a partial ornithischian dinosaur from Sustut Basin, as well as the ankylosaurs Zuul, Zaraapelta, Crichtonpelta, and Ziapelta. But she may have another shot at a pterosaur.

Dan Bowen, Chair, VIPS. Photo: Deanna Steptoe Graham

In 2019, Dan Bowen, Chair of the Vancouver Island Palaeontological Society and a truly awesome possum, found some very interesting bones in concretion on Hornby. 

The concretion was nestled amongst the 72 million-year-old grey shales of the Northumberland Formation, Campanian to the lower Maastrichtian, part of the Cretaceous Nanaimo Group from Collishaw Point.

The site is known as Boulder Point to the locals and it has been a popular fossil destination for many years. It is the same site where Sharon made her find years earlier.

The concretion contains four articulated vertebrae that looked to be fish at first glance. Jay Hawley, a local fossil enthusiast was asked to prep the block to reveal more details. Once the matrix was largely removed the vertebrae inside were revealed to be bird bones, not fish and not another saurodontid as originally thought. Palaeontologist Victoria Arbour was called back in to put her keen lens on the discovery. 

You will appreciate that she took a good long look at the specimen and confirmed it to be a bird or a pterosaur. We still do not have confirmation on which it is as yet. The delicate bony material is very flattened with a very shallow u-shape on the bottom but will need additional study to confirm if the skies above California were once home to a great pterosaur who died, was fossilized then rode our tectonic plates to now call Hornby home. It is a great story and one that I am keen to follow.

References: To learn more about the azhdarchid remains found by Sandy McLachlan, check out the paper by Martin-Silverston et al. 2016.

BRITISH COLUMBIA'S GIANT HIDDEN AMMONITE

Titanites occidentalis, Fernie Ammonite

The Fernie ammonite, Titanites occidentalis, from outcrops on Coal Mountain near Fernie, British Columbia, Canada. 

This beauty is the remains of a carnivorous cephalopod within the family Dorsoplanitidae that lived and died in a shallow sea some 150 million years ago.

If you would like to get off the beaten track and hike up to see this ancient beauty, you will want to head to the town of Fernie in British Columbia close to the Alberta border. 

This is the traditional territory of the the Yaq̓it ʔa·knuqⱡi ‘it First Nation who have lived here since time immemorial. There was some active logging along the hillside in 2021, so if you are looking at older directions on how to get to the site be mindful that many of the trailheads have been altered and a fair bit of bushwhacking will be necessary to get to the fossil site proper. That being said, the loggers from CanWel may have clear-cut large sections of the hillside but they did give the ammonite a wide berth and have left it intact.

Wildsight, a non-profit environmental group out of the Kimberly Cranbrook area has been trying to gain grant funding to open up the site as an educational hike with educational signage for folks visiting the Fernie area. It is likely the province of British Columbia would top up those funds if they are able to place the ammonite under the Heritage Conservation Act. CanWel would remain the owners of the land but the province could assume the liability for those visiting this iconic piece of British Columbia's palaeontological history. 

Driving to the trail base is along an easy access road just east of town along Fernie Coal Road. There are some nice exposures of Cretaceous plant material on the north side (left-hand side) of the road as you head from Fernie towards Coal Creek. I recently drove up to Fernie to look at Cretaceous plant material and locate the access point to the now infamous Late Jurassic (Tithonian) Titanites (S.S. Buckman, 1921) site. While the drive out of town is on an easy, well-maintained road, the slog up to the ammonite site is often a wet, steep push. This time of year, it is buried in a blanket of crisp, white snow.

Fernie, British Columbia, Canada

The first Titanites occidentalis was about one-third the size and was incorrectly identified as Lytoceras, a fast-moving nektonic carnivore. The specimen you see here is significantly larger at 1.4 metres (about four and a half feet) and rare in North America. 

Titanites occidentalis, the Western Giant, is the second known specimen of this extinct fossil species. 

The first was discovered in 1947 in nearby Coal Creek by a British Columbia Geophysical Society mapping team. When they first discovered this marine fossil high up on the hillside, they could not believe their eyes — both because it is clearly marine at the top of a mountain and the sheer size of this ancient beauty.

In the summer of 1947, on a warm summer's day, a field crew was mapping coal outcrops for the BC Geological Survey (GSC) east of Fernie. One of the students reported finding “a fossil truck tire.” Fair enough. The similarity of size and optics are pretty close to your average Goodridge. 

A few years later, GSC Paleontologist Hans Frebold described and named the fossil Titanites occidentalis after the large Jurassic ammonites from Dorset, England. The name comes from Greek mythology. Tithonus, as you may recall, was the Prince of Troy. He fell in love with Eos, the Greek Goddess of the Dawn. Eos begged Zeus to make her mortal lover immortal. Zeus granted her wish but did not grant Tithonus eternal youth. He did indeed live forever — ageing hideously. Ah, Zeus, you old trickster. It is a clever play on time placement. Dawn is the beginning of the day and the Tithonian being the latest age of the Late Jurassic. Clever Hans!

HIKING TO THE FERNIE AMMONITE

From the town of Fernie, British Columbia, head east along Coal Creek Road towards Coal Creek. The site is 3.81 km from the base of Coal Creek Road to the trailhead as the crow flies. I have mapped it here for you in yellow and added the wee purple GPS marker for the ammonite site proper. There is a nice, dark grey to black roadcut exposure of Cretaceous plants on the north side of the dirt road that is your cue to pull over and park.  

You access what is left of the trailhead on the south side of the road. You will need to cross the creek to begin your ascent. There is no easy way across the creek and you'll want to tackle this one with a friend when the water level is low. 

The beginning of the trail is not clear but a bit of searching will reveal the trailhead with its telltale signs of previous hikers. This is a moderate 6.3-kilometre hike up & back bushwhacking through scrub and fallen trees. Heading up, you will make about a 246-metre elevation gain. You will likely not have a cellular signal up here but if you download the Google Map to your mobile, you will have GPS to guide you. The area has been recently logged so much of the original trail has been destroyed. There may now be easier vehicle access up the logging roads but I have not driven them since the logging and new road construction.

If you are coming in from out of town, the closest airport is Cranbrook. Then it is about an hour and change to Fernie and another 15-minutes or so to park near the site.

You will want to leave your hammers with your vehicle (no need to carry the weight and this lovely should never be struck with anything more than a raindrop) as this site is best enjoyed with a camera. 

This is a site you will want to wear hiking boots to access. Know that these will get wet as you cross the creek. 

If you would like to see the ammonite but are not keen on the hike, a cast has been made by fossil preparator Rod Bartlett is on display at the Courtenay Museum in Courtenay, Vancouver Island, Canada. 

Respect for the Land / Leave No Trace

As your feet move up the hillside, you can imagine this land 10,000 years ago, rising above great glaciers. Where footfalls trace the steps of those that came before you. This land has been home to the Yaq̓it ʔa·knuqⱡi ‘it First Nation and Ktunaxa or Kukin ʔamakis First Nations whose oral history have them living here since time immemorial. Like them, take only what you need and no more than the land offers — packing out anything that you packed in. 

Fernie Ammonite Palaeo Coordinates: 49°29'04"N 115°00'49"W

RAINBOW AMMONITE: THE UNLIKELY LANDLORD

What you are seeing here is a protuberance extruding from the venter of Quenstedtoceras cf. leachi (Sowerby). It is a pathology in the shell from hosting immature bivalves that shared the seas with these Middle Jurassic, Upper Callovian, Lamberti zone fauna from the Volga River basin. The collecting site is the now inactive Dubki commercial clay quarry and brickyard near Saratov, Russia. 

The site has produced thousands of ammonite specimens. A good 1,100 of those ended up at the Black Hills Institute of Geological Research in Hill City, South Dakota. 

Roughly 1,000 of those are Quenstedtoceras (Lamberticeras) lamberti and the other 100 are a mix of other species found in the same zone. These included Eboraciceras, Peltoceras, Kosmoceras, Grossouvria, Proriceras, Cadoceras and Rursiceras. 

What is especially interesting is the volume of specimens — 167 Quenstedtoceras (Lamberticeras) lamberti and 89 other species in the Black Hills collection — with healed predation injuries. It seems Quenstedtoceras (Lamberticeras) lamberti are the most common specimens found here and so not surprisingly the most common species found injured. Of the 1,000, 655 of the Quenstedtoceras (Lamberticeras) lamberti displayed some sort of deformation or growth on the shell or had grown in a tilted manner. 

Again, some of the Q. lamberti had small depressions in the centre likely due to a healed bite and hosting infestations of the immature bivalve Placunopsis and some Ostrea. 

The bivalves thrived on their accommodating hosts and the ammonites carried on, growing their shells right up and over their bivalve guests. This relationship led to some weird and deformities of their shells. They grow in, around, up and over nearly every surface of the shell and seem to have lived out their lives there. It must have gotten a bit unworkable for the ammonites, their shells becoming warped and unevenly weighted. Over time, both the flourishing bivalves and the ammonite shells growing up and over them produced some of the most interesting pathology specimens I have ever seen.    

In the photo here from Emil Black, you can see some of the distorted shapes of Quenstedtoceras sp. Look closely and you see a trochospiral or flattened appearance on one side while they are rounded on the other. 

All of these beauties hail from the Dubki Quarry near Saratov, Russia. The ammonites were collected in marl or clay used in brick making. The clay particles suggest a calm, deep marine environment. One of the lovely features of the preservation here is the amount of pyrite filling and replacement. It looks like these ammonites were buried in an oxygen-deficient environment. 

The ammonites were likely living higher in the water column, well above the oxygen-poor bottom. An isotopic study would be interesting to prove this hypothesis. There's certainly enough of these ammonites that have been recovered to make that possible. It's estimated that over a thousand specimens have been recovered from the site but that number is likely much higher. But these are not complete specimens. We mostly find the phragmocones and partial body chambers. Given the numbers, this may be a site documenting a mass spawning death over several years or generations.

If you fancy a read on all things cephie, consider picking up a copy of Cephalopods Present and Past: New Insights and Fresh Perspectives edited by Neil Landman and Richard Davis. Figure 16.2 is from page 348 of that publication and shows the hosting predation quite well. 

Photos: Courtesy of the deeply awesome Emil Black. These are in his personal collection that I hope to see in person one day. 

It was his sharing of the top photo and the strange anomaly that had me explore more about the fossils from Dubki and the weird and wonderful hosting relationship between ammonites and bivalves. Thank you, my friend!

FOSSIL HUNTING: NOT FOR THE FAINT OF HEART

Coroniceras sp. from Sayward, British Columbia

This yummy Lower Jurassic ammonite with the creamy dark chocolate colouring is from an all but inaccessible outcrop of the Upper Sinemurian, Bonanza Group,  Harbledownense Zone, Memekay River area, near Sayward, Vancouver Island, British Columbia, Canada. 

This area is home to the We Wai Kai and Wei Wai Kum First Nations and lands of the K'omoks whose culture thrives and reflects the natural rugged beauty of the central island region.

I passed through Sayward earlier this month on the way to northern Vancouver Island. 

It is rugged, remote and beautiful. Think trees and valleys for as far as the eye can see. Some of those hillsides on the horizon contain wonderful fossils, including this Coroniceras sp. with the truly marvellous keel.

By the time these ammonites were being buried in sediment, Wrangellia, the predominately volcanic terrane that now forms Vancouver Island and Haida Gwaii, had made its way to the northern mid-latitudes.

Within the basal part of the sequence, sedimentary beds are found interbedded with lapilli and crystal tuffs. Here you'll see maroon tuffaceous sandstone, orange-grey sandstone, granule sandstone and conglomerate. Within them we find ammonites nestled in with gastropods and pelecypods. 

While the fossiliferous outcrop is quite small, the Bonanza group is much larger, estimated to be at least 1000 metres thick. The site is quite small and in an active logging area, so the window to collect was limited. The drive up the mountain was thrilling as there had just been heavy rains and the road was washed out and narrowed until it was barely the width of our wheelbase and very, very steep. Closer to the top it narrowed to be just shy of the width of the vehicle — thrilling, to say the least. 

So scary that my passengers all got out as there was a high probability of going head-first over the edge. I was navigating by some handwritten field notes and a wee map on a paper napkin that should have read, "park at the bottom and hike up." 

Did we park at the bottom and hike up? No, we did not. 

The torrential rains of the Pacific Northwest had been working their magic on the hillside and slowly washing out the road until it slowly became more of a trail.

At the base of the hillside all looked well. Giddy for the fossils to come, we ventured off with a truck full of enthusiasm. Within 15 minutes of steep elevation gain, we had a wonderful view of the valley below. We were halfway up the mountain before I realized the error of my ways. The road twisted and turned then slowly narrowed to the width of my tires. Too narrow to turn around, so the only way was up. 

Graham Beard from Qualicum Beach was the fellow who showed me the site and drew the wee map for me. I cannot recall everyone on the trip, but Perry Poon was there — he shot a video of the drive up that he described as thrilling. I have never seen it but would like to one day — and so was Patricia Coutts with her lovely Doberman. 

She and I had just done a trip up to Goldbridge where the cliff we were on had turned into a landslide into a ravine so she was feeling understandably cautious about the power of Mother Nature. 

Picture the angle, the hood of my jeep riding high and hiding what remained of the road beneath and a lovely stick shift that made you roll backwards a wee bit with every move to put it into gear. So, without being able to see the very narrow path beneath, I had to just keep going. 

Both Perry and Patricia helped with filling in the potholes so my tires would have something to grip. 

I bent the frame on the jeep heading up and had some explaining to do when I returned it to the car rental place. 

As I recall, I wasn't in my ordinary vehicle but a rental because my car had been stolen the weekend before when I was away with John Fam and Dan Bowen collecting at Jurassic Point, an epic fossil site accessible only by boat on our wild west coast.

Fortuitous timing really, as they stole my car but I had unloaded my precious fossil collecting gear out of the trunk just days before.  

In the end, we found what we were looking for. Memekay yields a mix of ammonites, gastropods and bivalves. 

Many of them are poorly preserved. It was a hell of a ride but well worth the effort as we found some great fossils and with them more information on the palaeontology and geology of Vancouver Island. Just look at the keel on this beauty.

I would share the site information but it is now covered over with debris and inaccessible. One day, this whole region will be developed and the site will be opened up again. Until then, we'll have to enjoy what has been unearthed.

AMMONITE TIME KEEPERS: INDEX FOSSILS

Argonauticeras besairei, José Juárez Ruiz

An exceptional example of the fractal building of an ammonite septum, in this clytoceratid Argonauticeras besairei from the awesome José Juárez Ruiz.

Ammonites were predatory, squidlike creatures that lived inside coil-shaped shells.

Like other cephalopods, ammonites had sharp, beak-like jaws inside a ring of squid-like tentacles that extended from their shells. 

They used these tentacles to snare prey, — plankton, vegetation, fish and crustaceans — similar to the way a squid or octopus hunt today.

Catching a fish with your hands is no easy feat, as I am sure you know. But the Ammonites were skilled and successful hunters. They caught their prey while swimming and floating in the water column. 

Within their shells, they had a number of chambers, called septa, filled with gas or fluid that were interconnected by a wee air tube. By pushing air in or out, they were able to control their buoyancy in the water column.

They lived in the last chamber of their shells, continuously building new shell material as they grew. As each new chamber was added, the squid-like body of the ammonite would move down to occupy the final outside chamber.

They were a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. 

These molluscs, commonly referred to as ammonites, are more closely related to living coleoids — octopuses, squid, and cuttlefish) than they are to shelled nautiloids such as the living Nautilus species.

The Ammonoidea can be divided into six orders:

• Agoniatitida, Lower Devonian - Middle Devonian
• Clymeniida, Upper Devonian
• Goniatitida, Middle Devonian - Upper Permian
• Prolecanitida, Upper Devonian - Upper Triassic
• Ceratitida, Upper Permian - Upper Triassic
• Ammonitida, Lower Jurassic - Upper Cretaceous

Ammonites have intricate and complex patterns on their shells called sutures. The suture patterns differ across species and tell us what time period the ammonite is from. If they are geometric with numerous undivided lobes and saddles and eight lobes around the conch, we refer to their pattern as goniatitic, a characteristic of Paleozoic ammonites.

If they are ceratitic with lobes that have subdivided tips; giving them a saw-toothed appearance and rounded undivided saddles, they are likely Triassic. For some lovely Triassic ammonites, take a look at the specimens that come out of Hallstatt, Austria and from the outcrops in the Humboldt Mountains of Nevada.

Hoplites bennettiana (Sowby, 1826) Christophe Marot

If they have lobes and saddles that are fluted, with rounded subdivisions instead of saw-toothed, they are likely Jurassic or Cretaceous. If you'd like to see a particularly beautiful Lower Jurassic ammonite, take a peek at Apodoceras. Wonderful ridging in that species.

One of my favourite Cretaceous ammonites is the ammonite, Hoplites bennettiana (Sowby, 1826). This beauty is from Albian deposits near Carrière de Courcelles, Villemoyenne, near la région de Troyes (Aube) Champagne in northeastern France.

At the time that this fellow was swimming in our oceans, ankylosaurs were strolling about Mongolia and stomping through the foliage in Utah, Kansas and Texas. Bony fish were swimming over what would become the strata making up Canada, the Czech Republic and Australia. Cartilaginous fish were prowling the western interior seaway of North America and a strange extinct herbivorous mammal, Eobaatar, was snuffling through Mongolia, Spain and England.

In some classifications, these are left as suborders, included in only three orders: Goniatitida, Ceratitida, and Ammonitida. Once you get to know them, ammonites in their various shapes and suturing patterns make it much easier to date an ammonite and the rock formation where it is found.

Ammonites first appeared about 240 million years ago, though they descended from straight-shelled cephalopods called bacrites that date back to the Devonian, about 415 million years ago, and the last species vanished in the Cretaceous–Paleogene extinction event.

They were prolific breeders that evolved rapidly. If you could cast a fishing line into our ancient seas, it is likely that you would hook an ammonite, not a fish. They were prolific back in the day, living (and sometimes dying) in schools in oceans around the globe. We find ammonite fossils (and plenty of them) in sedimentary rock from all over the world.

In some cases, we find rock beds where we can see evidence of a new species that evolved, lived and died out in such a short time span that we can walk through time, following the course of evolution using ammonites as a window into the past.

For this reason, they make excellent index fossils. An index fossil is a species that allows us to link a particular rock formation, layered in time with a particular species or genus found there. 

Generally, deeper is older, so we use the sedimentary layers of rock to match up to specific geologic time periods, rather like the way we use tree rings to date trees. A handy way to compare fossils and date strata across the globe.

References: Inoue, S., Kondo, S. Suture pattern formation in ammonites and the unknown rear mantle structure. Sci Rep 6, 33689 (2016). https://doi.org/10.1038/srep33689

https://www.nature.com/articles/srep33689?fbclid=IwAR1BhBrDqhv8LDjqF60EXdfLR7wPE4zDivwGORTUEgCd2GghD5W7KOfg6Co#citeas

Photos: Argonauticeras besairei from the awesome José Juárez Ruiz.

Photo: Hoplites bennettiana from near Troyes, France. Collection de Christophe Marot

FOSSIL FLORA AND FAUNA OF MADAGASCAR

Aioloceras besairiei (Collingnon, 1949)

A stunning example of the internal suturing with calcite infill in this sliced Aioloceras besairiei (Collingnon, 1949) ammonite from the Upper Cretaceous (Lower Albian) Boeny region of Madagascar. 

This island country is 400 kilometres off the coast of East Africa in the Indian Ocean and a wonderful place to explore off the beaten track.

Madagascar has some of the most spectacular of all the fossil specimens I have ever seen. This beauty is no exception. The shell has a generally small umbilicus, arched to acute venter, and typically at some growth stage, falcoid ribs that spring in pairs from umbilical tubercles, usually disappearing on the outer whorls. I had originally had this specimen marked as a Cleoniceras besairiei, except Cleoniceras and Grycia are not present in Madagascar. 

This lovely, seen in cross-section, is now far from home and in the collection of a wonderful friend. It is an especially lovely example of the ammonite, Aioloceras besairiei, making it a beudanticeratinae. Cleoniceras and Grycia are the boreal genera. If you'd like to see (or argue) the rationale on the name, consider reading Riccardi and Medina's riveting work from back in 2002, or Collingnon from 1949.

The beauty you see here measures in at a whopping 22 cm, so quite a handful. This specimen is from the youngest or uppermost subdivision of the Lower Cretaceous. I'd originally thought this locality was older, but dating reveals it to be from the Lower Albian, so approximately 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma.

Aioloceras are found in the Cretaceous of Madagascar at geo coordinates 16.5° S, 45.9° E: paleo-coordinates 40.5° S, 29.3° E.; and in four localities in South Africa: at locality 36, near the Mzinene River at 28.0° S, 32.3° E: paleo-coordinates 48.6° S, 7.6° E. 

We find them near the Mziene River, at a second locality north of Hluhluwe where the Mzinene Formation overlies the Aptian-Albian Makatini Formation at 28.0° S, 32.3° E: paleo-coordinates 48.6° S, 7.6° E; and at Haughton Z18, on the Pongola River in the Albian III, Tegoceras mosense beds at 27.3° S, 32.2° E: paleo-coordinates 48.0° S, 7.8° E.

If you happen to be trekking to Madagascar, know that it's big. It’s 592,800 square kilometres (or  226,917 square miles), making it the fourth-largest island on the planet — bigger than Spain, Thailand, Sweden and Germany. The island has an interesting geologic history.

Although there has been a geological survey, which was active extending back well into French colonial times, in the non-French-speaking world our geological understanding of the island is still a bit of a mystery. 

Plate tectonic theory had its beginnings in 1915 when Alfred Wegener proposed his theory of "continental drift." 

Wegener proposed that the continents ploughed through the crust of ocean basins, which would explain why the outlines of many coastlines (like South America and Africa) look like they fit together like a puzzle. Half a century after Wegener there is still no agreement as to whether in continental reconstructions Madagascar should be placed adjacent to the Tanzanian coast to the north (e.g., McElhinny and Embleton,1976), against the Mozambique-Natal coast (Flores 1970), or basically left where it is (Kent 1974, Nairn 1978).

There have been few attempts apart from McKinley’s (1960) comparison of the Karoo succession of southwestern Tanzania with that of Madagascar to follow the famous geological precept of “going to sea.” One critical reason is that although there may be a bibliography of several thousand items dealing with Madagascan geology as Besairie (1971) claims, they are items not generally available to the general public. The vital information gained of the geology of the offshore area by post-World War II petroleum exploration has remained largely proprietary. 

Without this data to draw upon, our understanding remains incomplete. I don't actually mind a bit of a mystery here. It is interesting to speculate on how these geologic puzzle pieces fit together and wait for the big reveal. Still, we have good old Besairie from his 1971, Geologie de Madagascar, and a later précis (Besairie, 1973).

We do know that Madagascar was carved off from the African-South American landmass early on. The prehistoric breakup of the supercontinent Gondwana separated the Madagascar–Antarctica–India landmass from the Africa–South America landmass around 135 million years ago. Madagascar later split from India about 88 million years ago, during the Late Cretaceous, so the native plants and animals on the island evolved in relative isolation. 

It is a green and lush island country with more than its fair share of excellent fossil exposures. Along the length of the eastern coast runs a narrow and steep escarpment containing much of the island's remaining tropical lowland forest. If you could look beneath this lush canopy, you'd see rocks of the Precambrian age stretching from the east coast all the way to the centre of the island. The western edge is made up of sedimentary rock from the Carboniferous to the Quaternary.

Red-Tailed Lemurs, Waiwai & Hedgehog

Madagascar is a biodiversity hotspot. Just as Darwin's finches on the Galápagos were isolated, evolving into distinct species (hello, adaptive radiation), over 90% of the wildlife from Madagascar is found nowhere else. 

The island's diverse ecosystems, like so many on this planet, are threatened by Earth's most deadly species, homo sapien sapiens. 

We arrived back in 490 CE and have been chopping down trees and eating our way through the island's tastier populations ever since. Still, they have cuties like this Red-Tailed Lemur. Awe, right?

Today, beautiful outcrops of wonderfully preserved fossil marine fauna hold appeal for me. The material you see from Madagascar is distinctive — and prolific.

Culturally, you'll see a French influence permeating the language, architecture and legal process. There is a part of me that pictures these lovely Lemurs chatting away in French. "Ah, la vache! Regarde le beau fossile, Hérissonne!"

We see the French influence because good 'ol France invaded sleepy Madagascar back in 1883, during the first Franco-Hova War. Malagasy (the local Madagascarian residents) were enlisted as troops, fighting for France in World War I.  During the Second World War, the island was the site of the Battle of Madagascar between the Vichy government and the British. By then, the Malagasy had had quite enough of colonization and after many hiccuping attempts, reached full independence in 1960. Colonization had ended but the tourist barrage had just begun. You can't stop progress.

If you're interested in learning more about this species, check out the Treatise on Invertebrate Paleontology, Part L (Ammonoidea). R.C. Moore (ed). Geological Soc of America and Univ. Kansas Press (1957), p L394. Or head over to look at the 2002 paper from Riccardi and Medina. 2002. Riccardi, A., C. & Medina, F., A. The Beudanticeratinae and Cleoniceratinae (Ammonitina) from the Lower Albian of Patagonia in Revue de Paléobiologie - 21(1) - Muséum d’Histoire Naturelle de la ville de Genève, p 313-314 (=Aioloceras besairiei (COLLIGNON, 1949). You have Bertrand Matrion to thank for the naming correction. Good to have friends in geeky places!

Collignon, M., 1933, Fossiles cenomaniens d’Antmahavelona (Province d’ Analalave, Madagascar), Ann. Geol. Serv. Min. Madagascar, III, 1934 Les Cephalopods du Trias inferieur de Madagascar, Ann. Paleont. XXII 3 and 4, XXII 1.

Besairie, H., 1971, Geologie de Madagascar, 1. Les terrains sedimentaires, Ann. Geol. Madagascar, 35, p. 463.

J. Boast A. and E. M. Nairn collaborated on a chapter in An Outline of the Geology of Madagascar, that is very readable and cites most of the available geologic research papers. It is an excellent place to begin a paleo exploration of the island.

If you happen to parle français, check out: Madagascar ammonites: http://www.ammonites.fr/Geo/Madagascar.htm

FERNIE GIANT: A MASSIVE AMMONITE HIDDEN IN THE TREES

Titanites occidentalis, Fernie Ammonite

The Fernie ammonite, Titanites occidentalis, from outcrops on Coal Mountain near Fernie, British Columbia, Canada. 

This beauty is the remains of a carnivorous cephalopod within the family Dorsoplanitidae that lived and died in a shallow sea some 150 million years ago.

If you would like to get off the beaten track and hike up to see this ancient beauty, you will want to head to the town of Fernie in British Columbia close to the Alberta border. 

This is the traditional territory of the the Yaq̓it ʔa·knuqⱡi ‘it First Nation who have lived here since time immemorial. There was some active logging along the hillside in 2021, so if you are looking at older directions on how to get to the site be mindful that many of the trailheads have been altered and a fair bit of bushwhacking will be necessary to get to the fossil site proper. That being said, the loggers from CanWel may have clear-cut large sections of the hillside but they did give the ammonite a wide berth and have left it intact.

Wildsight, a non-profit environmental group out of the Kimberly Cranbrook area has been trying to gain grant funding to open up the site as an educational hike with educational signage for folks visiting the Fernie area. It is likely the province of British Columbia would top up those funds if they are able to place the ammonite under the Heritage Conservation Act. CanWel would remain the owners of the land but the province could assume the liability for those visiting this iconic piece of British Columbia's palaeontological history. 

Driving to the trail base is along an easy access road just east of town along Fernie Coal Road. There are some nice exposures of Cretaceous plant material on the north side (left-hand side) of the road as you head from Fernie towards Coal Creek. I recently drove up to Fernie to look at Cretaceous plant material and locate the access point to the now infamous Late Jurassic (Tithonian) Titanites (S.S. Buckman, 1921) site. While the drive out of town is on an easy, well-maintained road, the slog up to the ammonite site is often a wet, steep push.

Fernie, British Columbia, Canada

The first Titanites occidentalis was about one-third the size and was incorrectly identified as Lytoceras, a fast-moving nektonic carnivore. The specimen you see here is significantly larger at 1.4 metres (about four and a half feet) and rare in North America. 

Titanites occidentalis, the Western Giant, is the second known specimen of this extinct fossil species. 

The first was discovered in 1947 in nearby Coal Creek by a British Columbia Geophysical Society mapping team. When they first discovered this marine fossil high up on the hillside, they could not believe their eyes — both because it is clearly marine at the top of a mountain and the sheer size of this ancient beauty.

In the summer of 1947, a field crew was mapping coal outcrops for the BC Geological Survey east of Fernie. One of the students reported finding “a fossil truck tire.” Fair enough. The similarity of size and optics are pretty close to your average Goodridge. 

A few years later, GSC Paleontologist Hans Frebold described and named the fossil Titanites occidentalis after the large Jurassic ammonites from Dorset, England. The name comes from Greek mythology. Tithonus, as you may recall, was the Prince of Troy. He fell in love with Eos, the Greek Goddess of the Dawn. Eos begged Zeus to make her mortal lover immortal. Zeus granted her wish but did not grant Tithonus eternal youth. He did indeed live forever — ageing hideously. Ah, Zeus, you old trickster. It is a clever play on time placement. Dawn is the beginning of the day and the Tithonian being the latest age of the Late Jurassic. Clever Hans!

HIKING TO THE FERNIE AMMONITE

From the town of Fernie, British Columbia, head east along Coal Creek Road towards Coal Creek. The site is 3.81 km from the base of Coal Creek Road to the trailhead as the crow flies. I have mapped it here for you in yellow and added the wee purple GPS marker for the ammonite site proper. There is a nice, dark grey to black roadcut exposure of Cretaceous plants on the north side of the dirt road that is your cue to pull over and park.  

You access what is left of the trailhead on the south side of the road. You will need to cross the creek to begin your ascent. There is no easy way across the creek and you'll want to tackle this one with a friend when the water level is low. 

The beginning of the trail is not clear but a bit of searching will reveal the trailhead with its telltale signs of previous hikers. This is a moderate 6.3-kilometre hike up & back bushwhacking through scrub and fallen trees. Heading up, you will make about a 246-metre elevation gain. You will likely not have a cellular signal up here but if you download the Google Map to your mobile, you will have GPS to guide you. The area has been recently logged so much of the original trail has been destroyed. There may now be easier vehicle access up the logging roads but I have not driven them since the logging and new road construction.

If you are coming in from out of town, the closest airport is Cranbrook. Then it is about an hour and change to Fernie and another 15-minutes or so to park near the site.

You will want to leave your hammers with your vehicle (no need to carry the weight and this lovely should never be struck with anything more than a raindrop) as this site is best enjoyed with a camera. 

This is a site you will want to wear hiking boots to access. Know that these will get wet as you cross the creek. 

If you would like to see the ammonite but are not keen on the hike, a cast has been made by fossil preparator Rod Bartlett is on display at the Courtenay Museum in Courtenay, Vancouver Island, Canada. 

Respect for the Land / Leave No Trace

As your feet move up the hillside, you can imagine this land 10,000 years ago, rising above great glaciers. Where footfalls trace the steps of those that came before you. This land has been home to the Yaq̓it ʔa·knuqⱡi ‘it First Nation and Ktunaxa or Kukin ʔamakis First Nations whose oral history have them living here since time immemorial. Like them, take only what you need and no more than the land offers — packing out anything that you packed in. 

Fernie Ammonite Palaeo Coordinates: 49°29'04"N 115°00'49"W

LOWER LIAS LYTOCERAS

A superbly prepped and extremely rare Lytoceras (Suess, 1865) ammonite found as a green ammonite nodule by Matt Cape in the Lower Lias of Dorset. 

Lytoceras are rare in the Lower Lias of Dorset — apart from the Belemnite Stone horizon — so much so that Paul Davis, whose skilled prep work you see here, initially thought it might be a Becheiceras hidden within the large, lumpy nodule. 

One of the reasons these lovelies are rarely found from here is that they are a Mediterranean Tethyian genus. The fossil fauna we find in the United Kingdom are dominated by Boreal Tethyian genera. 

We do find Lytoceras sp. in the Luridum subzone of the Pliensbachian showing that there was an influx of species from the Mediterranean realm during this time. This is the first occurrence of a Lytoceras that he has ever seen in a green nodule and Paul's seen quite a few. 

This absolutely cracking specimen was found and is in the collections of the awesome Matt Cape. Matt recognized that whatever was hidden in the nodule would take skilled and careful preparation using air scribes. Indeed it did. It took more than five hours of time and skill to unveil the lovely museum-worthy specimen you see here. 

We find Lytoceras in more than 1,000 outcrops around the globe ranging from the Jurassic through to the Cretaceous, some 189.6 to 109.00 million years ago. Once this specimen is fully prepped with the nodule material cut or scraped away, you can see the detailed crinkly growth lines or riblets on the shell and none of the expected coarse ribbing. 

Lytoceras sp. Photo: Craig Chivers

If you imagine running your finger along these, you would be tracing the work of decades of growth of these cephalopods. 

While we cannot know their actual lifespans, but we can make a healthy guess. 

The nautilus, their closest living cousins live upwards of 20 years — gods be good — and less than three years if conditions are poor.

The flanges, projecting flat ribs or collars, develop at the edge of the mouth border on the animal's mantle as they grow each new chamber. 

Each delicate flange grows over the course of the ammonites life, marking various points in time and life stages as the ammonite grew. There is a large variation within Lytoceras with regards to flanges. They provide both ornamentation and strength to the shell to protect it from water pressure as they moved into deeper seas.

The concretion prior to prep

This distinctive genus with its evolute shells are found in the Cretaceous marine deposits of: 

Antarctica (5 collections), Austria (19), Colombia (1), the Czech Republic (3), Egypt (2), France (194), Greenland (16), Hungary (25), Italy (11), Madagascar (2), Mexico (1), Morocco (4), Mozambique (1), Poland (2), Portugal (1), Romania (1), the Russian Federation (2), Slovakia (3), South Africa (1), Spain (24), Tanzania (1), Trinidad and Tobago (1), Tunisia (25); and the United States of America (17: Alaska, California, North Carolina, Oregon).

We also find them in Jurassic marine outcrops in:

Austria (15), Canada (9: British Columbia), Chile (6), France (181), Germany (11), Greenland (1), Hungary (189), India (1), Indonesia (1), Iran (1), Italy (50), Japan (14), Kenya (2), Luxembourg (4), Madagascar (2), Mexico (1), Morocco (43), New Zealand (15), Portugal (1), Romania (5), the Russian Federation (1), Slovakia (1), Spain (6), Switzerland (2), Tunisia (11), Turkey (12), Turkmenistan (1), Ukraine (5), the United Kingdom (12), United States (11: Alaska, California) — in at least 977 known collections. 

References:

Sepkoski, Jack (2002). "A compendium of fossil marine animal genera (Cephalopoda entry)". Bulletins of American Paleontology. 363: 1–560. Archived from the original on 2008-05-07. Retrieved 2017-10-18.

Paleobiology Database - Lytoceras. 2017-10-19.

Systematic descriptions, Mesozoic Ammonoidea, by W.J Arkell, Bernhard Kummel, and C.W. Wright. 1957. Treatise on Invertebrate Paleontology, Part L. Geological Society of America and University of Kansas press.

AN AMMONITE HOSTING WEE BIVALVES

What you are seeing here is a protuberance extruding from the venter of Quenstedtoceras cf. leachi (Sowerby). It is a pathology in the shell from hosting immature bivalves that shared the seas with these Middle Jurassic, Upper Callovian, Lamberti zone fauna from the Volga River basin. The collecting site is the now inactive Dubki commercial clay quarry and brickyard near Saratov, Russia. 

The site has produced thousands of ammonite specimens. A good 1,100 of those ended up at the Black Hills Institute of Geological Research in Hill City, South Dakota. 

Roughly 1,000 of those are Quenstedtoceras (Lamberticeras) lamberti and the other 100 are a mix of other species found in the same zone. These included Eboraciceras, Peltoceras, Kosmoceras, Grossouvria, Proriceras, Cadoceras and Rursiceras. 

What is especially interesting is the volume of specimens — 167 Quenstedtoceras (Lamberticeras) lamberti and 89 other species in the Black Hills collection — with healed predation injuries. It seems Quenstedtoceras (Lamberticeras) lamberti are the most common specimens found here and so not surprisingly the most common species found injured. Of the 1,000, 655 of the Quenstedtoceras (Lamberticeras) lamberti displayed some sort of deformation or growth on the shell or had grown in a tilted manner. 

Again, some of the Q. lamberti had small depressions in the centre likely due to a healed bite and hosting infestations of the immature bivalve Placunopsis and some Ostrea. 

The bivalves thrived on their accommodating hosts and the ammonites carried on, growing their shells right up and over their bivalve guests. This relationship led to some weird and deformities of their shells. They grow in, around, up and over nearly every surface of the shell and seem to have lived out their lives there. It must have gotten a bit unworkable for the ammonites, their shells becoming warped and unevenly weighted. Over time, both the flourishing bivalves and the ammonite shells growing up and over them produced some of the most interesting pathology specimens I have ever seen.    

In the photo here from Emil Black, you can see some of the distorted shapes of Quenstedtoceras sp. Look closely and you see a trochospiral or flattened appearance on one side while they are rounded on the other. 

All of these beauties hail from the Dubki Quarry near Saratov, Russia. The ammonites were collected in marl or clay used in brick making. The clay particles suggest a calm, deep marine environment. One of the lovely features of the preservation here is the amount of pyrite filling and replacement. It looks like these ammonites were buried in an oxygen-deficient environment. 

The ammonites were likely living higher in the water column, well above the oxygen-poor bottom. An isotopic study would be interesting to prove this hypothesis. There's certainly enough of these ammonites that have been recovered to make that possible. It's estimated that over a thousand specimens have been recovered from the site but that number is likely much higher. But these are not complete specimens. We mostly find the phragmocones and partial body chambers. Given the numbers, this may be a site documenting a mass spawning death over several years or generations.

If you fancy a read on all things cephie, consider picking up a copy of Cephalopods Present and Past: New Insights and Fresh Perspectives edited by Neil Landman and Richard Davis. Figure 16.2 is from page 348 of that publication and shows the hosting predation quite well. 

Photos: Courtesy of the deeply awesome Emil Black. These are in his personal collection that I hope to see in person one day. 

It was his sharing of the top photo and the strange anomaly that had me explore more about the fossils from Dubki and the weird and wonderful hosting relationship between ammonites and bivalves. Thank you, my friend!

OH, WHAT WE DO FOR FOSSILS

Coroniceras sp. from Sayward, British Columbia

This yummy Lower Jurassic ammonite with the creamy dark chocolate colouring is from an all but inaccessible outcrop of the Upper Sinemurian, Bonanza Group,  Harbledownense Zone, Memekay River area, near Sayward, Vancouver Island, British Columbia, Canada. 

This area is home to the We Wai Kai and Wei Wai Kum First Nations and lands of the K'omoks whose culture thrives and reflects the natural rugged beauty of the central island region.

I passed through Sayward earlier this month on the way to northern Vancouver Island. 

It is rugged, remote and beautiful. Think trees and valleys for as far as the eye can see. Some of those hillsides on the horizon contain wonderful fossils, including this Coroniceras sp. with the truly marvellous keel.

By the time these ammonites were being buried in sediment, Wrangellia, the predominately volcanic terrane that now forms Vancouver Island and Haida Gwaii, had made its way to the northern mid-latitudes.

Within the basal part of the sequence, sedimentary beds are found interbedded with lapilli and crystal tuffs. Here you'll see maroon tuffaceous sandstone, orange-grey sandstone, granule sandstone and conglomerate. Within them we find ammonites nestled in with gastropods and pelecypods. 

While the fossiliferous outcrop is quite small, the Bonanza group is much larger, estimated to be at least 1000 metres thick. The site is quite small and in an active logging area, so the window to collect was limited. The drive up the mountain was thrilling as there had just been heavy rains and the road was washed out and narrowed until it was barely the width of our wheelbase and very, very steep. Closer to the top it narrowed to be just shy of the width of the vehicle — thrilling, to say the least. 

So scary that my passengers all got out as there was a high probability of going head-first over the edge. I was navigating by some handwritten field notes and a wee map on a paper napkin that should have read, "park at the bottom and hike up." 

Did we park at the bottom and hike up? No, we did not. 

The torrential rains of the Pacific Northwest had been working their magic on the hillside and slowly washing out the road until it slowly became more of a trail.

At the base of the hillside all looked well. Giddy for the fossils to come, we ventured off with a truck full of enthusiasm. Within 15 minutes of steep elevation gain, we had a wonderful view of the valley below. We were halfway up the mountain before I realized the error of my ways. The road twisted and turned then slowly narrowed to the width of my tires. Too narrow to turn around, so the only way was up. 

Graham Beard from Qualicum Beach was the fellow who showed me the site and drew the wee map for me. I cannot recall everyone on the trip, but Perry Poon was there — he shot a video of the drive up that he described as thrilling. I have never seen it but would like to one day — and so was Patricia Coutts with her lovely Doberman. 

She and I had just done a trip up to Goldbridge where the cliff we were on had turned into a landslide into a ravine so she was feeling understandably cautious about the power of Mother Nature. 

Picture the angle, the hood of my jeep riding high and hiding what remained of the road beneath and a lovely stick shift that made you roll backwards a wee bit with every move to put it into gear. So, without being able to see the very narrow path beneath, I had to just keep going. 

Both Perry and Patricia helped with filling in the potholes so my tires would have something to grip. 

I bent the frame on the jeep heading up and had some explaining to do when I returned it to the car rental place. 

As I recall, I wasn't in my ordinary vehicle but a rental because my car had been stolen the weekend before when I was away with John Fam and Dan Bowen collecting at Jurassic Point, an epic fossil site accessible only by boat on our wild west coast.

Fortuitous timing really, as they stole my car but I had unloaded my precious fossil collecting gear out of the trunk just days before.  

In the end, we found what we were looking for. Memekay yields a mix of ammonites, gastropods and bivalves. 

Many of them are poorly preserved. It was a hell of a ride but well worth the effort as we found some great fossils and with them more information on the palaeontology and geology of Vancouver Island. Just look at the keel on this beauty.

I would share the site information but it is now covered over with debris and inaccessible. One day, this whole region will be developed and the site will be opened up again. Until then, we'll have to enjoy what has been unearthed.