PROSAUROLOPHUS MAXIMUS

Prosaurolophus maximus, Ottawa Museum of Nature

Prosaurolophus was a large-headed duckbill dinosaur. The most complete described specimen has a skull around 0.9 metres (3.0 ft) long on a skeleton about 8.5 metres (28 ft) long. It had a small, stout, triangular crest in front of the eyes; the sides of this crest were concave, forming depressions.

This crest grew isometrically (without changing in proportion) throughout the lifetime of the individual, leading to speculation that the species may have had a soft tissue display structure, such inflatable nasal sacs.

When originally described by Brown, Prosaurolophus maximus was known from a skull and jaw. Half of the skull was badly weathered at the time of examination, and the level of the parietal was distortedly crushed upwards to the side.

The different bones of the skull are easily defined with the exception of the parietals and nasal bones. Brown found that the skull of the already described genus Saurolophus is very similar overall but also smaller than the skull of P. maximus. The unique feature of a shortened frontal in lambeosaurines is also found in Prosaurolophus, and the other horned hadrosaurines Brachylophosaurus, Maiasaura, and Saurolophus. Although they lack a shorter frontal, the genera Edmontosaurus and Shantungosaurus share an elongated dentary structure.

Patches of preserved skin are known from two juvenile specimens, TMP 1998.50.1 and TMP 2016.37.1; these pertain to the ventral extremity of the ninth through fourteenth dorsal ribs, the caudal margin of the scapular blade, and the pelvic region. Small basement scales (scales which make up the majority of the skin surface), 3–7 millimetres (0.12–0.28 in) in diameter, are preserved on these patches - this is similar to the condition seen in other saurolophine hadrosaurs.

More uniquely, feature scales (larger, less numerous scales which are interspersed within the basement scales) around 5 millimetres (0.20 in) wide and 29 millimetres (1.1 in) long are found interspersed in the smaller scales in the patches from the ribs and scapula (they are absent from the pelvic patches). Similar scales are known from the tail of the related Saurolophus angustirostris (on which they have been speculated to indicate pattern), and it is considered likely adult Prosaurolophus would've retained the feature scales on their flanks like the juveniles.

HOPLOSCAPHITES NEBRASCENSIS

This sweet beauty with lovely colouring is a Hoploscaphites nebrascensis (Owen, 1852) macroconch. This is the female form of the ammonite that has a larger shell than the male, or microconch.

Hoploscaphites nebrascensis is an upper Maastrichtian species and index fossil. It marks the top of ammonite zonation for the Western Interior. This species has been recorded from Fox Hills Formation in North and South Dakota as well as the Pierre Shale in southeastern South Dakota and northeastern Nebraska.

It is unknown from Montana, Wyoming, and Colorado due to the deposition of coeval terrestrial units. It has possibly been recorded in glacial deposits in Saskatchewan and northern North Dakota, but that is hearsay. Outside the Western Interior, this species has been found in Maryland and possibly Texas in the Discoscaphites Conrad zone. This lovely one is in the collection of the deeply awesome (and enviable) José Juárez Ruiz. A big thank you to Joshua DrSlattmaster J Slattery for his insights on this species.

CRETACEOUS HADROSAUR TOOTH

A rare and very beautifully preserved Cretaceous Hadrosaur Tooth. This lovely specimen is from one of our beloved herbivorous "Duck-Billed" dinosaurs from 68 million-year-old outcrops near Drumheller, Alberta, Canada — and is likely from an Edmontosaurus.

When you scour the badlands of southern Alberta, most of the dinosaur material you'll find are from hadrosaurs. These lovely tree-less valleys make for excellent-searching grounds and have led us to know more about hadrosaur anatomy, evolution, and paleobiology than for most other dinosaurs. 

We have oodles of very tasty specimens and data to work with. We've got great skin impressions and scale patterns from at least ten species and interesting pathological specimens that provide valuable insights into hadrosaur behaviour. These herbivorous beauties are also found in Europe, South America, Mexico, Mongolia, China and Russian. Together, this abundance of specimens has provided great insight into their evolution, dining habits and social preferences. We know they liked to live in herds. They were terrestrial but also water babies — paddling around in freshwater pools to snack on the tasty greenery that lined its sides. They had adapted webbing in their feet to be as nimble on land as they were in the water. 

There are papers on all aspects of hadrosaurian life and not surprisingly — given the ideal collecting grounds — many of those papers focus on our Canadian finds. Hadrosaurs had teeth arranged in stacks designed for grinding and crushing, similar to how you might picture a cow munching away on the grass in a field. These complex rows of "dental batteries" contained up to 300 individual teeth in each jaw ramus. But even with this great number, we rarely see them as individual specimens.

They didn't appear to shed them all that often. Older teeth that are normally shed in our general understanding of vertebrate dentition, were resorped, meaning that their wee osteoclasts broke down the tooth tissue and reabsorbed the yummy minerals and calcium.

As the deeply awesome Mike Boyd notes, "this is an especially lucky find as hadrosaurs did not normally shed so much as a tooth, except as the result of an accident when feeding or after death. Typically, these fascinating dinosaurs ground away their teeth... almost to nothing."

In hadrosaurs, the root of the tooth formed part of the grinding surface as opposed to a crown covering over the core of the tooth. And curiously, they developed this dental arrangement from their embryonic state, through to hatchling then full adult.

There's some great research being done by Aaron LeBlanc, Robert R. Reisz, David C. Evans and Alida M. Bailleul. They published in BMC Evolutionary Biology on work that looks at the histology of hadrosaurid teeth analyzing them through cross-sections. Jon Tennant did a nice summary of their research. I've included both a link to the original journal article and Jon Tennant's blog below.

LeBlanc et al. are one of the first teams to look at the development of the tissues making up hadrosaur teeth, analyzing the tissue and growth series (like rings of a tree) to see just how these complex tooth batteries formed.

They undertook the first comprehensive, tissue-level study of dental ontogeny in hadrosaurids using several intact maxillary and dentary batteries and compared them to sections of other archosaurs and mammals. They used these comparisons to pinpoint shifts in the ancestral reptilian pattern of tooth ontogeny that allowed hadrosaurids to form complex dental batteries.

References:

LeBlanc et al. (2016) Ontogeny reveals function and evolution of the hadrosaurid dinosaur dental battery, BMC Evolutionary Biology. 16:152, DOI 10.1186/s12862-016-0721-1 (OA link)

To read more from Jon Tennant, visit: https://blogs.plos.org/paleocomm/2016/09/14/all-the-better-to-chew-you-with-my-dear/

Photo credit: Derrick Kersey. For more awesome fossil photos like this from Derrick, visit his page: https://www.facebook.com/prehistoricexpedition/

MARINE REPTILES OF THE HUMBOLDTS

A very well preserved ichthyosaur block with three distinct vertebrae and some ribs just peeking out. You can see the edges of the ribs nicely outlined against the matrix.

Ichthyosaurs are an extinct order of marine reptiles from the Mesozoic era. They evolved from land-dwelling, lung-breathing reptiles, they returned to our ancient seas and evolved into the fish-shaped creatures we find in the fossil record today.

They were visibly dolphin-like in appearance but seem to share some other qualities as well. These lovelies were warm-blooded and used their coloration as camouflage. The smaller of their lineage to avoid being eaten and the larger to avoid being seen by prey. Ichthyosaurs also had insulating blubber, a lovely adaptation to keep them warm in cold seas.

Over time, their limbs fully transformed into flippers, sometimes containing a very large number of digits and phalanges. Their flippers tell us they were entirely aquatic as they were not well-designed for use on land. And it was their flippers that first gave us the clue that they gave birth to live young; a hypothesis later confirmed by fossil embryo and wee baby ichy specimens.

We find their fossil remains in outcrops spanning from the mid-Cretaceous to the earliest Triassic. As we look through the fossils, we see a slow evolution in body design moving towards that enjoyed by dolphins and tuna by the Upper Triassic, albeit with a narrower, more pointed snout. During the early Triassic period, ichthyosaurs evolved from a group of unidentified land reptiles that returned to the sea. They were particularly abundant in the later Triassic and early Jurassic periods before being replaced as a premier aquatic predator by another marine reptilian group, the Plesiosauria, in the later Jurassic and Cretaceous periods.

The block you see here is from Middle Triassic (Anisian/Ladinian) outcrops in the West Humboldt Mountains, Nevada.

SUNRISE FORMATION, NEVADA

At the entrance to the Pliensbachian-Toarcian localities at Joker Peak and Mina Peak Members of the Sunrise Formation, Nevada, USA.

The ammonites of this section were first studied by Dr. Paul Smith, past Chair of Earth and Ocean Sciences, University of British Columbia and more recently by Andrew Caruthers et al.

Caruthers and his team also took a goodly look at the Early Jurassic coral fauna. Caruthers is an interesting cat. He uses a combination of invertebrate paleontology and isotope geochemistry to ponder the effects of paleoclimate change and mass extinction. He's turned his eye in recent years to the Paleozoic of the Michigan Basin AND he's based in Kalamazoo, MI. Yep, Kalamazoo.

Others have taken up the mantle of discovery from these sites. Pengfei Hou did his 2014 Masters thesis comparing the Sinemurian (Early Jurassic) stratigraphic sections of Last Creek, British Columbia and Five Card Draw, Nevada including a detailed taxonomic study from the Involutum Zone to the lower part of the Harbledownense Zone of the Sinemurian.

DUBIOUS DAONELLA DUBIA

Triassic ammonoids, West Humboldt Mountains, Nevada, USA. This was the site of the 1905 Expedition of the University of California’s Department of Geology in Berkeley funded by the beautiful and bold, Annie Alexander, the women to whom the UCMP owes both its collection and existence.

Paleontologist J.P. Smith joined that expedition and published on the marine fauna in the early 1900s.

They formed the basis for his monograph on North American Middle Triassic marine invertebrate fauna published in 1914. N. J. Siberling from the US Geological Survey published on these outcrops in 1962. His work included nearly a dozen successive ammonite faunas, many of which were variants on previously described species.

Evidently, his collections consisted mainly of weathered material and were made without stratigraphic control because he believed that most, if not all, of these species, were coexistent. The fossiliferous beds found here, as well as localities in north-western Nevada, were designated the 'Daonella dubia' zone. Dubious would be closer to the truth. We've since mapped them out from stratigraphic sections to place them in the correct order of their occurrence.

LATE HETTANGIAN TO EARLY SINEMURIAN FAUNA

Hiking the hills of Nevada looking for David Taylor's faunal succession based on ammonoids established for the Late Hettangian to Early Sinemurian interval in the Western Cordillera.

It was a tremendous experience to walk through time and compare the fossil assemblages here with our own in the Canadian Rockies.

Here the faunal sequence consists of one zone and four informal biochronologic units or assemblages and was outlined by Taylor as follows: Paracaloceras morganense assemblage, Badouxia oregonensis assemblage, Canadensis Zone, Metophioceras trigonatum assemblage and Coroniceras involutum. They matched up to specimens we collected over three field seasons to similar faunal outcrops of Late Hettangian to Early Sinemurian of the Last Creek and Tyaughton area of the Canadian Rockies.

The succession also correlates with the interval delineated by the Northwest European Angulata Zone through the Lyra Subzone. Two new genera (Guexiceras and Tipperoceras) are described along with 23 new species. The phylogenetic relationships of the earliest Jurassic ammonite superfamilies indicate that it is useful to include under the Psiloceratida, the Psilocerataceae and their derivatives including the Lytocerataceae. The Arietitaceae were derived from Hettangian lytocerataceans.

MIDDLE TRIASSIC OF NEVADA

Searching for bedrock in outcrops of the West Humboldt Mountains, Nevada. Perhaps the most famous and important locality for the Middle Triassic (Anisian/Ladinian) of North America. These beautiful hills are home to Triassic ammonoid outcrops and plentiful ichthyosaur fossils.

J.P. Smith published on the marine fauna in the early 1900's. They formed the basis for his monograph on North American Middle Triassic marine invertebrate fauna published in 1914. N. J. Siberling from the US Geological Survey published on these outcrops in 1962. His work included nearly a dozen successive ammonite faunas, many of which were variants on previously described species.

Evidently, his collections consisted mainly of weathered material and were made without stratigraphic control because he believed that most, if not all, of these species were coexistent. The fossiliferous beds found here, as well as localities in north-western Nevada, were designated the 'Daonella dubia' zone. Dubious would be closer to the truth. Smith joined the 1905 Expedition of the University of California’s Department of Geology in Berkeley funded by the beautiful and bold, Annie Alexander, the women to whom the UCMP owes both its collection and existence.

GRASPING HOOKLETS AND CALAMARI

This well-preserved partial ichthyosaur was found in the Blue Lias shales by Lewis Winchester-Ellis in 2018. The vertebrae you see are from the tail section of this marine reptile.

The find includes stomach contents that tell us a little about how this particular fellow liked to dine.

As with most of his brethren, he enjoyed fish and cephalopods. Lewis found fishbone and squid tentacle hooklets in his belly. 

Oh yes, these ancient cephies had grasping hooklets on their tentacles. I'm picturing them wiggling all ominously. The hooklets were the only hard parts of the animal preserved in this case as the softer parts of this ancient calamari were fully or partially digested before this ichthyosaur met his end.

Ichthyosaurus was an extinct marine reptile first described from fossil fragments found in 1699 in Wales. Shortly thereafter, fossil vertebrae were published in 1708 from the Lower Jurassic and the first member of the order Ichthyosauria to be discovered.

To give that a bit of historical significance, this was the age of James Stuart, Jacobite hopeful to the British throne. While scientific journals of the day were publishing the first vertebrae ichthyosaur finds, he was avoiding the French fleet in the Firth of Forth off Scotland. This wasn’t Bonnie Prince Charlie, this was his Dad. Yes, that far back.

The first complete skeleton was discovered in the early 19th century by Mary Anning & her brother Joseph along the Dorset Jurassic Coast. Joseph had mistakenly, but quite reasonably, taken the find for an ancient crocodile. Mary excavated the specimen a year later and it was this and others that she found that would supply the research base others would soon publish on.

Mary's find was described by a British surgeon, Sir Everard Home, an elected Fellow of the Royal Society, in 1814. The specimen is now on display at the Natural History Museum in London bearing the name Temnodontosaurus platyodon, or “cutting-tooth lizard.”

In 1821, William Conybeare and Henry De La Beche, a friend of Mary's, published a paper describing three new species of unknown marine reptiles based on Anning's finds.

The Rev. William Buckland would go on to describe two small ichthyosaurs from the Lias of Lyme Regis, Ichthyosaurus communis and Ichthyosaurus intermedius, in 1837.

Remarkable, you'll recall that he was a theologian, geologist, palaeontologist AND Dean of Westminster. It was Buckland who published the first full account of a dinosaur in 1824, coining the name, "Megalosaurus."

The Age of Dinosaurs and Era of the Mighty Marine Reptile had begun.

Ichthyosaurs have been collected in the Blue Lias near Lyme Regis and the Black Ven Marls. More recently, specimens have been collected from the higher succession near Seatown. Paddy Howe, Lyme Regis Museum geologist, found a rather nice Ichthyosaurus breviceps skull a few years back. A landslip in 2008 unveiled some ribs poking out of the Church cliffs and a bit of digging revealed the ninth fossil skull ever found of a breviceps, with teeth and paddles to boot.

Specimens have since been found in Europe in Belgium, England, Germany, Switzerland and in Indonesia. Many tremendously well-preserved specimens come from the limestone quarries in Holzmaden, southern Germany.

Ichthyosaurs ranged from quite small, just a foot or two, to well over twenty-six metres in length and resembled both modern fish and dolphins.

Dean Lomax and Sven Sachs, both active (and delightful) vertebrate palaeontologists, have described a colossal beast, Shonisaurus sikanniensis from the Upper Triassic (Norian) Pardonet Formation of northeastern British Columbia, Canada, measuring 3-3.5 meters in length. The specimen is now on display in the Royal Tyrrell Museum of Palaeontology in Alberta, Canada. It was this discovery that tipped the balance in the vote, making it British Columbia's Official Fossil. Ichthyosaurs have been found at other sites in British Columbia, on Vancouver Island and Haida Gwaii, but Shoni tipped the ballot.

The first specimens of Shonisaurus were found in the 1990s by Peter Langham at Doniford Bay on the Somerset coast of England.

Dr. Betsy Nicholls, Rolex Laureate Vertebrate Palaeontologist from the Royal Tyrrell Museum, excavated the type specimen of Shonisaurus sikanniensis over three field sessions in one of the most ambitious fossil excavations ever ventured. Her efforts from 1999 through 2001, both in the field and lobbying back at home, paid off. Betsy published on this new species in 2004, the culmination of her life’s work and her last paper as we lost her to cancer in autumn of that year.

Charmingly, Betsy had a mail correspondence with Roy Chapman Andrews, former director of the American Museum of Natural History, going back to the late 1950s as she explored her potential career in palaeontology. Do you recall the AMNH’s sexy paleo photos of expeditions to the Gobi Desert in southern Mongolia in China in the early 20th century? You’d remember if you’d seen them. Roy Chapman Andrews was the lead on that trip. His photos are what fueled the flames of my own interest in paleo.

We've found at least 37 specimens of Shonisaurus in Triassic outcrops of the Luning Formation in the Shoshone Mountains in northwestern Nye County of Nevada, USA. The finds go back to the 1920s. The specimens that may it to publication were collected by Margaret Wheat of Fallon and Dr. C. L. Camp, UCMP, in the 1950s.  The aptly named Shonisaurus popularis became the Nevada State Fossil in 1977. Our Shoni got around. Isolated remains have been found in a section of sandstone in Belluno, in the Eastern Dolomites, Veneto region of northeastern Italy. The specimens were published by Vecchia et al. in 2002.

For a time, Shonisaurus was the largest ichthyosaurus known.

Move over, Shoni, as a new marine reptile find competes with the Green Anaconda (Eunectes murinus) and the Blue Whale (Balaenoptera musculus) for size at a whopping twenty-six (26) metres.

The find is the prize of fossil collector turned co-author, Paul de la Salle, who (you guessed it) found it in the lower part of the intertidal area that outcrops strata from the latest Triassic Westbury Mudstone Formation of Lilstock on the Somerset coast. He contacted Dean Lomax and Judy Massare who became co-authors on the paper.

The find and conclusions from their paper put "dinosaur" bones from the historic Westbury Mudstone Formation of Aust Cliff, Gloucestershire, UK site into full reinterpretation.

And remember that ichthyosaur the good Reverend Buckland described back in 1837, the Ichthyosaurus communis? Dean Lomax was the first to describe a wee baby. A wee baby ichthyosaur! Awe. I know, right? He and paleontologist Nigel Larkin published this adorable first in the journal of Historical Biology in 2017.

They had teamed up previously on another first back in 2014 when they completed the reconstruction of an entire large marine reptile skull and mandible in 3-D, then graciously making it available to fellow researchers and the public. The skull and braincase in question were from an Early Jurassic, and relatively rare, Protoichthyosaurus prostaxalis. 

The specimen had been unearthed in Warwickshire back in the 1950's. Unlike most ichthyosaur finds of this age, it was not compressed and allowed the team to look at a 3-D specimen through the lens of computerized tomography (CT) scanning. Another superb 3-D ichthyosaur skull was found near Lyme Regis by fossil hunter-turned-entrepreneur-local David Sole and prepped by the late David Costain. I'm rather hoping it went into a museum collection as it would be wonderful to see the specimen studied, imaged, scanned and 3-D printed for all to share. Here's hoping.

Lomax and Sven Sachs also published on an embryo from one of the largest ichthyosaurs known, a new species named Ichthyosaurus somersetensis. Their paper in the ACTA Palaeontologica Polonica from 2017, describes the third embryo known for Ichthyosaurus and the first to be positively identified to species level. The specimen was collected from the Lower Jurassic strata (lower Hettangian, Blue Lias Formation) of Doniford Bay, Somerset, UK and is housed in the collection of the Niedersächsisches Landesmuseum (Lower Saxony State Museum) in Hannover, Germany.

We've learned a lot about them in the time we've been studying them. We now have thousands of specimens, some whole, some as bits and pieces. Many specimens that have been collected are only just now being studied and the tools we are using to study them are getting better and better.

While they resembled fish and dolphins, Ichthyosaurs were large marine reptiles belonging to the order known as Ichthyosauria or Ichthyopterygia. In 2018, Benjamin Kear and his team were able to study ichthyosaur remains at the molecular level, Their findings suggest ichthyosaurs had skin and blubber quite similar to our modern dolphins.

While ichthyosaurs evolved from land-dwelling, lung-breathing reptiles, they returned to our ancient seas and evolved into the fish-shaped creatures we find in the fossil record today.

Their limbs fully transformed into flippers, sometimes containing a very large number of digits and phalanges. Their flippers tell us they were entirely aquatic as they were not well-designed for use on land. And it was their flippers that first gave us the clue that they gave birth to live young; a find later confirmed by fossil embryo and wee baby ichy finds.

They thrived during much of the Mesozoic era; based on fossil evidence, they first appeared around 250 million years ago (Ma) and at least one species survived until about 90 million years ago into the Late Cretaceous.

During the early Triassic period, ichthyosaurs evolved from a group of unidentified land reptiles that returned to the sea. They were particularly abundant in the later Triassic and early Jurassic periods before being replaced as a premier aquatic predator by another marine reptilian group, the Plesiosauria, in the later Jurassic and Cretaceous periods.

In the Late Cretaceous, ichthyosaurs were hard hit by the Cenomanian-Turonian anoxic event. As the deepest benthos layers of the seas became anoxic, poisoned by hydrogen sulphide, deep water marine life died off. This caused a cascade that wreaked havoc all the way up the food chain. At the end of that chain were our mighty predaceous marine reptiles. Bounty turned to scarcity and a race for survival began. The ichthyosaurs lost that race as the last lineage became extinct. It may have been their conservative evolution as a genus when faced with a need for adaptation to the world in which they found themselves and/or being outcompeted by early mosasaurs.

There are promising discoveries coming out of strata from the Cretaceous epeiric seas of Texas, USA from Nathan E. Van Vranken. His published paper from 2017, "An overview of ichthyosaurian remains from the Cretaceous of Texas, USA," looks at ichthyosaurian taxa from the mid-Cretaceous (Albian–Cenomanian) time interval in North America with an eye to ichthyosaurian distribution and demise.

The find and photos are all credited to Lewis Winchester-Ellis. Thank you for sharing your tremendous specimen with us. Lewis did much of the preparation of the specimen, removing the majority of the matrix. The spectacular final prep is credited to Lizzie Hingley, Stonebarrow Fossils, Oxfordshire. Her skill with an air scribe is unparalleled.

Link to Lomax Paper: https://journals.plos.org/plosone/article…

Link to Nathan's Paper: https://www.tandfonline.com/…/10.1080/03115518.2018.1523462…

Nicholls Paper: E. L. Nicholls and M. Manabe. 2004. Giant ichthyosaurs of the Triassic - a new species of Shonisaurus from the Pardonet Formation (Norian: Late Triassic) of British Columbia. Journal of Vertebrate Paleontology 24(4):838-849 [M. Carrano/H. Street]

ICHTHYOSAUR QUARRIES OF THE HUMBOLDT

Looking out over the Middle Triassic exposures of the Humboldt Mountain Range.

These hills were the site of the 1905 Expedition of the University of California’s Department of Geology in Berkeley funded by the beautiful and bold, Annie Alexander, the women to whom the UCMP owes both its collection and existence. Annie brought together a paleontological crew to explore these localities and kept an expedition journal of their trip which is now on display at the University of California Museum of Paleontology at Berkeley.

Annie's interest was the ichthyosaurs and she was well pleased with the results. They dodged rattlesnakes and tarantulas, finding many new specimens as they opened up new quarries in the hills of the Humboldt Range of Nevada.

Ichthyosaurs range from quite small, just a foot or two, to well over twenty-six metres in length and resembled both modern fish and dolphins. The specimens from Nevada are especially large and well-preserved. They hail from a time, some 217 million years ago, when Nevada, and parts of the western USA, was covered by an ancient ocean that would one day become our Pacific Ocean. Many ichthyosaur specimens have come out of Nevada. So many, in fact, that they named it their State Fossil back in 1977.

Fossil fragments and complete specimens of these marine reptiles have been collected in the Blue Lias near Lyme Regis and the Black Ven Marls. More recently, specimens have been collected from the higher succession near Seatown. Paddy Howe, Lyme Regis Museum geologist, found a rather nice Ichthyosaurus breviceps skull a few years back. A landslip in 2008 unveiled some ribs poking out of the Church cliffs and a bit of digging revealed the ninth fossil skull ever found of a breviceps, with teeth and paddles to boot.

Specimens have since been found in Europe in Belgium, England, Germany, Switzerland and in Indonesia. Many tremendously well-preserved specimens come from the limestone quarries in Holzmaden, southern Germany.

KASKAPAU FORMATION: DINOSAUR BONE

Dinosaur bone / Kaskapau Formation

Bones from a variety of dinosaurs have been found in the Tumbler Ridge area of British Columbia.

Here plaster is used to protect a valuable dinosaur bone collected from Flatbed Creek near Tumbler Ridge. The bone is from the Kaskapau Formation (Turonian) and was found a few metres away from a Tetrapodosaurus, "four-footed lizard," trackway.

Both Rich McCrea and Lisa Buckley have published extensively on the fossil material from this area. Additional Papers: Arbour et al. (2008ish) wrote up a paper in the Canadian Journal of Earth Science on dinosaur material collected in the 60s from BC; Rylaarsdam et al. contributed to the same journal two years earlier on the association of dinosaur footprints and skeletal material in the Kaskapau Formation.

MIDDLE TRIASSIC AMMONOIDS OF NEVADA

A beautiful block of marine fossils found in bedrock outcrops of the West Humboldt Mountains, Nevada. Perhaps the most famous and important locality for the Middle Triassic (Anisian/Ladinian) of North America. These beautiful hills are home to Triassic ammonoid outcrops and plentiful ichthyosaur fossils.

J.P. Smith published on the marine fauna in the early 1900s. They formed the basis for his monograph on North American Middle Triassic marine invertebrate fauna published in 1914. N. J. Siberling from the US Geological Survey published on these outcrops in 1962. His work included nearly a dozen successive ammonite faunas, many of which were variants on previously described species.

Smith was a surface collector and it showed in his research. His collections consisted mainly of weathered material and were made without stratigraphic control because he believed that most, if not all, of these species, were coexistent.

The fossiliferous beds found here, as well as localities in north-western Nevada, were designated the 'Daonella dubia' zone. Dubious would be closer to the truth. Smith joined the 1905 Expedition of the University of California’s Department of Geology in Berkeley funded by the beautiful and bold, Annie Alexander, the women to whom the UCMP owes both its collection and existence.

SHOSHONE MOUNTAIN RANGE

Berlin-Ichthyosaur State Park encompasses 1,540 acres. The elevation ranges from 6,840 feet to a high point of 7,880 feet. The hillsides in the park are covered with big sagebrush, while pinyon pine and Utah juniper dominate the upper elevations.

Common animal inhabitants include mule deer, black-tailed jackrabbits, cottontails, western bluebirds, pinyon jays, chukar partridge, whiptail lizards, western fence lizards, gophers and snakes.

​​Nestled at 7,000 feet on the western slope of central Nevada’s Shoshone mountain range, the park provides an array of stimulating recreational opportunities. The forested slopes provide shade, and breezes help to moderate the summer temperatures which seldom exceed 90˚F. Late spring and early fall can often be the nicest times of the year. Winter visits to the park are possible, but call for weather and road conditions before visiting.

MIDDLE TRIASSIC AMMONOID & BELEMNITE

A beautiful plate with a lovely example of the Middle Triassic ammonoid and belemnite from Fossil Hill in the Humboldt Mountains of Nevada. This locality is perhaps the most famous and important locality for the Middle Triassic (Anisian/Ladinian) of North America.

This was the site of the 1905 Expedition of the University of California’s Department of Geology in Berkeley funded by the beautiful and bold, Annie Alexander, the women to whom the UCMP owes both its collection and existence. J. P. Smith joined the expedition, though he was interested in the invertebrate fauna, not the mighty marine reptiles that helped get the project funded.

J.P. Smith published on the marine fauna in the early 1900s. They formed the basis for his monograph on North American Middle Triassic marine invertebrate fauna published in 1914.

N. J. Siberling from the US Geological Survey published on these outcrops in 1962. His work included nearly a dozen successive ammonite faunas, many of which were variants on previously described species. Evidently, his collections consisted mainly of weathered material and were made without stratigraphic control because he believed that most, if not all, of these species, were coexistent. The fossiliferous beds found here, as well as localities in north-western Nevada, were designated the 'Daonella dubia' zone. Dubious would be closer to the truth. Smith joined the 1905 Expedition of the University of California’s Department of Geology in Berkeley funded by the beautiful and bold, Annie Alexander, the women to whom the UCMP owes both its collection and existence.

FERGUSON HILL, NEVADA

Ferguson Hill contains the most complete macrofossil record spanning the Triassic-Jurassic boundary in North America. The ammonoids from the uppermost Triassic can be traced to the boundary and the earliest ammonites (Psiloceratids) can be seen right at the base of the Jurassic (Hettangian).

It was in contention for the Global Stratotype Section and Point (GSSP) defining the base of the Jurassic System, Lower Jurassic Series and Hettangian Stage. However, in 2008, the Kuhjoch (Karwendel Mountains, Northern Calcareous Alps, Tyrol, Austria) was chosen over Ferguson Hill mainly because the beds containing the oldest Psiloceras (P. spelae spelae) were better preserved and contained Hettangian microfossils.

NOTOCHORDS: CHORDATES TO CALAMARI

You and I are vertebrates, we have backbones. Having a backbone or spinal column is what sets apart you, me and almost 70,000 species on this big blue planet.

So which lucky ducks evolved one? Well, ducks for one. Warm-blooded birds and mammals cheerfully claim those bragging rights. They're joined by our cold-blooded, ectothermic friends, the fish, amphibians and reptiles. All these diverse lovelies share this characteristic.

And whether they now live at sea or on land, all of these lineages evolved from a marine organism somewhere down the line, then went on to develop a notochord and spinal column. Notochords are flexible rods that run down the length of chordates and vertebrates. They are handy adaptations for muscle attachment, helping with signalling and coordinating the development of the embryonic stage. The cells from the notochord play a key role in the development of the central nervous system and the formation of motor neurons and sensory cells. Alas, we often take our evolution for granted.

Let's take a moment to appreciate just how marvellous this evolutionary gift is and what it allows us to do. Your backbone gives your body structure, holds up that heavy skull of yours and connects your tasty brain to your body and organs. Eating, walking, fishing, hunting, your morning yoga class, are all made possible because of this adaptation. Pick pretty near anything you love to do and it is only possible because of your blessed spine.

And it sets us apart from our invertebrate friends.

While seventy thousand may seem like a large number, it represents less than three to five per cent of all described animal species. The rest is made up of the whopping 97%'ers, our dear invertebrates who include the arthropods (insects, arachnids, crustaceans, and myriapods), molluscs (our dear chitons, snails, bivalves, squid, and octopus), annelids (the often misunderstood earthworms and leeches), and cnidarians (our beautiful hydras, jellyfish, sea anemones, and corals). You'll notice that many of our invertebrate friends occur as tasty snacks. Having a backbone provides a supreme advantage to your placement in the food chain. Not always, as you may include fish and game on your menu. But generally, having a backbone means you're more likely to be holding the menu versus being listed as an appetizer. So, enjoy your Sunday 'downward dog' and thank your backbone for the magical gift it is.

DOUVELLICERAS SPINIFERUM

Douvelliceras spiniferum, Cretaceous Haida Formation

Haida Gwaii or the Queen Charlotte Islands lay at the western edge of the continental shelf due west of the central coast of British Columbia. They form Wrangellia, an exotic tectonostratigraphic terrane that includes Vancouver Island, parts western British Columbia and Alaska.

The Geological Survey of Canada sponsored many expeditions to these remote islands and has produced numerous reference papers on this magnificent terrain, exploring both the geology and paleontology of the area.

Joseph Whiteaves, the GSC 's chief palaeontologist in Ottawa, published a paper in 1876 describing the Jurassic and Cretaceous faunas of Skidegate Inlet, furthering his reputation globally as both a geologist and paleontologist.

The praise was well-earned and foreshadowed his significant contributions to come. Sixteen years later, he wrote up and published his observations on a strange Mount Stephen fossil that resembled a kind of headless shrimp with poorly preserved appendages. Because of the unusual pointed shape of the supposed ventral appendages and the position of the spines near the posterior of the animal, Whiteaves named it Anomalocaris canadensis. The genus name "Anomalocaris" meant "unlike other shrimps" and the species name "canadensis" referred to the country of origin.

Whiteaves work on the paleontology of the Queen Charlotte Island provided us with excellent reference tools, particularly his work on the Cretaceous exposures and fauna that can be found there.

One of our fossil field trips was to the ruggedly beautiful Cretaceous exposures of Lina Island. We’d planned this trip as part of our “trips of a lifetime.” Both John Fam and Dan Bowen can be congratulated for their efforts in researching the area and ably coordinating a warm welcome by the First Nations community and organizing fossil field trips to some of the most amazing fossil localities in the Pacific Northwest. With great sandstone beach exposures, the fossil-rich (Albian to Cenomanian) Haida formation provided ample specimens, some directly in the bedding planes and many in concretion. Many of the concretions contained multiple specimens of typical Haida Formation fauna, providing a window into this Cretaceous landscape.

It is always interesting to see who was making a living and co-existing in our ancient oceans at the time these fossils were laid down. We found multiple beautifully preserved specimens of the spiny ammonite, Douvelleiceras spiniferum along with Brewericeras hulenense, Cleoniceras perezianum and many cycads in concretion.

Pictured above is Douvilleiceras spiniferum with his naturally occurring black, shiny appearance. Danna Straaf had recently asked me what my favourite cephalopod is, and I have to say that it is a hard choice but this fellow is in the top three. He is 6 inches long and 5 inches deep, and a beautiful example of the species.

MCABEE FOSSIL BEDS

Eocene Fossil Feather / McAbee Fossil Beds

The McAbee fossil beds are known for their incredible abundance, diversity and quality of fossils including lovely plant, insect and fish species that lived in an old lake bed setting 52 million years ago.

I was sharing with some friends, Lawrence and Shivinder (hello you two!) about the site earlier this evening. It is one of the best local sites in the province to experience a fossil dig first-hand. 

It is an easy 4-hour drive from Vancouver and easily done as a day trip. The site was designated a Provincial Heritage Site under British Columbia's Heritage Conservation Act in July of 2012, then promptly closed to the public.

It has recently been reopened to public collecting (as of June 21, 2019), with plans to build out a visitor's centre and educational programs. The Province is committed to providing access to the site to scientists and the lay public. The direction on what happens next at McAbee is being driven by the Heritage Branch in consultation with members of the Shuswap Nation and Bonaparte Band. Bonaparte traditional territory is located within the Shuswap Nation and includes the area known as McAbee.

Local members of the Bonaparte Band are Secwepemc. They want to share the spiritual significance of the area from a First Nations perspective and see McAbee as an indigenous tourism destination. So it looks like it will be palaeontology, archaeology with a cultural focus to add spice. In any case, the collection of fossils will continue with oversight to ensure significant fossil finds make their way to science.

While the area is referred to as the Okanagan, the term is used in a slightly misleading fashion to describe an arc of Eocene lakebed sites that extend from Smithers in the north, down to the fossil site of Republic Washington, in the south. The grouping includes the fossil sites of Driftwood Canyon, Quilchena, Allenby, Tranquille, McAbee, Princeton and Republic.

Fossils from the Okanagan Highlands, an area centred in the Interior of British Columbia, provide important clues to our ancient climate. The fossils range in age from the Early to Middle Eocene. McAbee had a more temperate climate, slightly cooler and wetter than other Eocene sites to the south at Princeton, British Columbia, Republic in north-central Washington, in the Swauk Formation near Skykomish and the Chuckanut Formation of northern Washington state.

The McAbee fossil beds consist of 30 metres of fossiliferous shale in the Eocene Kamloops Group.
The fossils are preserved here as impressions and carbonaceous films. We see gymnosperm (16 species); a variety of conifers (14 species to my knowledge); two species of ginkgo, a large variety of angiosperm (67 species); a variety of insects and fish remains, the rare feather and a boatload of mashed deciduous material. Nuts and cupules are also found from the dicotyledonous Fagus and Ulmus and members of the Betulaceae, including Betula and Alnus.

We see many species that look very similar to those growing in the Pacific Northwest today. Specifically, cypress, dawn redwood, fir, spruce, pine, larch, hemlock, alder, birch, dogwood, beech, sassafras, cottonwood, maple, elm and grape. If we look at the pollen data, we see over a hundred highly probable species from the site. Though rare, McAbee has also produced spiders, birds (and lovely individual feathers) along with multiple specimens of the freshwater crayfish, Aenigmastacus crandalli.

For insects, we see dragonflies, damselflies, cockroaches, termites, earwigs, aphids, leaf hoppers, spittlebugs, lacewings, a variety of beetles, gnats, ants, hornets, stick insects, water striders, weevils, wasps and March flies. The insects are particularly well-preserved. Missing are the tropical Sabal (palm), seen at Princeton and the impressive Ensete (banana) and Zamiaceae (cycad) found at Eocene sites in Republic and Chuckanut, Washington.

My first trips up there were as a teenager, dragging my mother, sister and pretty near anyone else I could convince to hike up. This was in 1986-87, years before Dave Langevin and John Leahy, mineral rights/lease-holder and resident curator, respectively, began working at the site. I think Dave put in his mineral claim in 1991ish. 

Once they did a whole new world opened up with their efforts. Much of the overburden was removed and new exposures were revealed. John also used to leave a jeep at the base of the hill with a bit of gas in it that we'd hot wire and use to avoid the hike heading up and pack down fossils heading back. Good man, John. He was an avid collector and meticulous in his curation. Both of those gents have now passed and are sorely missed. Most of their personal collection is now in the Royal BC Museum in Victoria, British Columbia, and much of Dave is still at the site as his ashes were sprinkled there.

McAbee is located just east of Cache Creek, just north of and visible from Highway 1/97. 14.5 km to be exact and exactly the distance you need to drink one large coffee and then need a washroom. You'll be pleased to know they have installed one at the site. McAbee is a site for hiking boots, hand, head and eye protection. Keep yourself safe and well-hydrated.

As you drive up, you'll see telltale hoodoos on the ridge to let you know you've reached the right spot. If you have a GPS, pop in these coordinates and you're on your way. 50°47.831′N 121°8.469′W.

HYPODICRANOTUS STRIATULUS OF ONTARIO

Hypodicranotus striatulus

Take a gander at this unusual trilobite, Hypodicranotus striatulus (Walcott, 1875), with his gloriously bulbous head shield. Missing from this specimen is the wonderful forked hypostome from the dorsal exoskeleton that marks him as H. striatulus.

He’s from outcrops in the Verulam Formation, Bowmanville, Ontario, Canada. He lived in a deep subtidal environment as a nektobenthic deposit feeder some 460.9 to 449.5 million years ago.

These extinct pelagic trilobites are in the order Asaphida in the family Remopleuridae. Specimens have been found in Middle Ordovician marine outcrops from Ontario, Canada (this fellow is from here), the Northwest Territories, Quebec and in New York State, United States. Some of his sister taxa also in Remopleurididae (Hawle and Corda, 1847)  have been found in the Northwest Territories, Quebec, the UK and in Iowa, Wisconsin and Nevada. Collection of the awesome Marc R. Hänsel

SKØKKENMØDDINGER

Many First Nations sites were inhabited continually for centuries. These sites were both home, providing continuity and community and also formed a spiritual connection to the landscape.

The day to day activities of each of these communities would much like our own. Babies were born, meals were served and life followed a natural cycle.

As coastal societies lived their lives they also left their mark. Sometimes through totems and carvings but almost always through discarded shells and scraps of bone from their food. These refuse heaps contain a wealth of information about how that community lived, what they ate and what environmental conditions looked like over time. This physical history provides a wonderful resource for archaeologists in search of botanical material, artifacts, broken cooking implements and my personal favourite, mollusc shells.

These wonderfully informative heaps of the local gastronomic record provide a wealth of information. Especially those formed from enormous mounds of bivalves and clams. We call these middens. Left over time, these unwanted dinner scraps transform through a process of preservation.

The Danish term køkkenmøddinger (plural) was first used by Japetus Steenstrup, a Danish zoologist and biologist, to describe shell heaps and continues to be used by some researchers. I still prefer middens, but to each his own. Time and pressure leach the calcium carbonate, CaCO3, from the surrounding marine shells and help “embalm” bone and antler artifacts that would otherwise decay. Useful this, as antler makes for a fine sewing tool when worked into a needle. A bone that has had some time to harden through this natural embalming process makes for a fine needle indeed. Much of what we know around the modification of natural objects into tools comes from this preservation.

Calcium carbonate is a chemical compound that shares the typical properties of other carbonates. In prepping fossil specimens embedded in limestone, it is useful to know that limestone, itself a carbonate sedimentary rock, reacts with stronger acids, releasing carbon dioxide: CaCO3(s) + 2HCl(aq) → CaCl2(aq) + CO2(g) + H2O(l).

Calcium carbonate reacts with water saturated with carbon dioxide to form the soluble calcium bicarbonate. Bone already contains calcium carbonate, as well as calcium phosphate, Ca2, but it is also made of protein, cells and living tissue.

Decaying bone acts as a sort of natural sponge that wicks in the calcium carbonate displaced from the shells. As protein decays inside the bone, it is replaced by the incoming calcium carbonate, making the bone harder and more durable.

The shells, beautiful in their own right, make the surrounding soil more alkaline, helping to preserve the bone and turning the dinner scraps into exquisite scientific specimens for future generations.