TRACKING THROUGH THE TRIASSIC

Grambergia sp. Middle Triassic Ammonoid of  BC, Canada

In the early 1980s, Tim Tozer, Geological Survey of Canada was looking at the spread of marine invertebrate fauna in the Triassic of North America. 

In the western terranes of the Cordillera, marine faunas from southern Alaska and Yukon to Mexico are known from the parts that are obviously allochthonous with regard to the North American plates.

Lower and upper Triassic faunas of these areas, as well as some that are today up to 63 ° North, have the characteristics of the lower palaeo latitudes. 

In the western Cordillera, these faunas of the lower paleo latitudes can be found up to 3,000 km north of their counterparts on the American plate. This indicates a tectonic shift of significant magnitude. There are marine triads on the North American plate over 46 latitudes from California to Ellesmere Island. 

For some periods, two to three different faunal provinces can be distinguished. The differences in faunal species are linked, not surprisingly, to their palaeolatitude. They are called LPL, MPL, HPL (lower, middle, higher palaeolatitude).

Nevada provides the diagnostic features of the lower (LPL); northeastern British Columbia that of the middle (MPL) and Sverdrup Basin, the large igneous province on Axel Heiberg Island and Ellesmere Island, Nunavut, Canada near the rifted margin of the Arctic Ocean, that of the higher palaeolatitude (HPL).

A distinction between the provinces of the middle and the higher palaeo-situations can not be made for the lower Triassic and lower Middle Triassic (anise). However, all three provinces can be seen in the deposits of Ladin, Kam and Nor.

In the early 2000s, as part of a series of joint UBC, VIPS and VanPS fossil field trips (and then Chair of the VanPS), I explored much of the lower faunal outcrops of northeastern British Columbia. It was my first time seeing many of British Columbia's Triassic outcrops. Years later, and fueled by seeing paper after paper correlating the faunal assemblages of BC to those of Nevada, I had the very great pleasure of walking through the Nevada strata with John Fam (VanPS, Vice-Chair), Dan Bowen (VIPS, Chair) and Betty Franklin (VIPS, Goddess of Everything and BCPA, Treasurer) — and witnessing first-hand the correlation between the Nevada fauna and those from the Triassic of British Columbia, Canada.

Triassic ammonoids, West Humboldt Mountains, Nevada, USA

The Nevada faunal assemblages are a lovely match. The quality of preservation at localities like Fossil Hill in the Humboldt Mountains of Nevada, perhaps the most famous and important locality for the Middle Triassic (Anisian/Ladinian) of North America, is truly outstanding.

Aside from sheer beauty and spectacular preservation, the ammonoids and belemnites were tucked in cozily with very well preserved ichthyosaur remains.

Tozer's interest in our marine invert friends was their distribution. How and when did certain species migrate, cluster, evolve — and for those that were prolific, how could their occurrence — and therefore significance — aide in an assessment of plate and terrane movements that would help us to determine paleolatitudinal significance. 

I share a similar interest but not exclusive to our cephalopod fauna. The faunal collection of all of the invertebrates holds appeal.

Middle Triassic (Anisian/Ladinian) Fauna

This broader group held an interest for J.P. Smith who published on the marine fauna in the early 1900s based on his collecting in scree and outcrops of the West Humboldt Mountains, Nevada. He published his first monograph on North American Middle Triassic marine invertebrate fauna in 1914.

N. J. Siberling from the US Geological Survey published on these same Nevada outcrops in 1962. His work included nearly a dozen successive ammonite faunas, many of which were variants on previously described species. Both their works would inform what would become a lifelong piecing together of the Triassic puzzle for Tozer.

If one looks at the fauna and the type of sediment, the paleogeography of the Triassic can be interpreted as follows: a tectonically calm west coast of the North American plate that bordered on an open sea; in the area far from the coast, a series of volcanic archipelagos delivered sediment to the adjacent basins. 

Some were lined or temporarily covered with coral wadding and carbonate banks. Deeper pools were in between. The islands were probably within 30 degrees of the triadic equator. They moved away from the coast up to about 5000 km from the forerunner of the East Pacific Ridge. The geographical situation west of the back was probably similar.

Jurassic and later generations of the crust from near the back have brought some of the islands to the North American plate; some likely to South America; others have drifted west, to Asia. There are indications that New Guinea, New Caledonia and New Zealand were at a northern latitude of 30 ° or more during the Triassic period.

The terranes that now form the western Cordillera were probably welded together and reached the North American plate before the end of the Jurassic period.

Marine Triassic occurs on the North American Plate over a latitudinal spread of 46 degrees, from California to Ellesmere Island. At some intervals of time faunas on the Plate permit the discrimination of two or three provinces with distinctively different coeval faunas. 

The faunal differences are evidently related to paleolatitude and the provinces are designated LPL, MPL, HPL (low, mid, high paleolatitude). Nevada provides the diagnostic characters of the LPL province; northeastern British Columbia the MPL; the Sverdrup Basin the HPL. In the Lower Triassic and early Middle Triassic (Anisian), the distinction between the MPL and HPL provinces cannot be made. All three provinces are recognized in the Ladinian, Carnian and Norian deposits.

Juvavites sp. Geological Survey of Canada. Photo: John Fam

In the western tracts of the Cordillera, the part formed of suspect terranes, apparently allochthonous with respect to the North American Plate, marine faunas are known all the way from southern Alaska and Yukon to Mexico.

Lower and Upper Triassic faunas from these terranes, including some which today are at 63 degrees north, have the characters of the LPL province.

Middle Triassic faunas from the terranes, as presently known, do not contribute significant data. In the terranes of the Western Cordillera, LPL faunas were now up to 3,000 km north of their counterparts on the American Plate. Through the fossil fauna assemblages, we can see this level of tectonic displacement.

Taking into account the faunas and the nature of the rocks, the Triassic paleogeography is interpreted as a tectonically quiet west shore for the North American Plate, bordered by an open sea or ocean; then, well off-shore, a series of volcanic archipelagos shedding sediment into adjacent basins. Some were fringed or intermittently covered by coralline shoals and carbonate banks. Deeper basins were in between. The islands probably were within 30 degrees of the Triassic equator and extended offshore for about 5000 km, to the spreading ridge directly ancestral to the East Pacific Rise. The geography west of the spreading ridge was probably comparable.

Jurassic and later generation of crust at the ridge had driven some of the islands into the North American Plate; some probably to South America; others have gone west to Asia. Evidence is given that northern New Guinea, New Caledonia and New Zealand may have been at a north latitude of 30 degrees or more in the Triassic. The terranes now forming the Western Cordillera had probably amalgamated, and reached the North American Plate, before the end of the Jurassic.

At the end of the Rhaetian (part of the Triassic period), most of the ammonites had died out. These are the lovely coiled molluscs you often see in museums and gift shops that sell fossils. They are a particular favourite of mine and they are both beautiful and useful to tell us much about deep time. The Hettangian, a rather poorly understood 3 million year time interval followed the Triassic-Jurassic mass extinction event.

During the Hettangian, the new or  Neoammonites developed quite quickly. Within a million years, a fairly large, diverse selection of genera and species had risen to fill the void. The gap created by the Triassic-Jurassic extinction event was re-filled and our ability to "read the rocks' to understand their continued movement through tectonic plate shifting recommenced.

Alsatites proaries, Hettangian Ammonite

It is during the Hettangian that the smooth shelled ammonite genus Psiloceras first appears. They span the time between 201.3 ± 0.2 Ma and 199.3 ± 0.3 Ma (million years ago). For my European friends, the Hettangian is the time span in which the marine limestone, shales and clay Lias of western Europe were deposited.

This Hettangian ammonite, Alsatites proaries, is a lovely example of the cephalopods cruising our ancient oceans at that time. Alsatites is an extinct genus of cephalopod belonging to the Ammonite subclass. They lived during the Early Jurassic, Hettangian till the Sinemurian and are generally extremely evolute, many whorled with a broad keel. Or, as described by one of my very young friends, he looks like a coiled snake you make in pottery class.

The Hettangian is an interesting little period of our history. It spans the time between 201.3 ± 0.2 Ma and 199.3 ± 0.3 Ma (million years ago). For my European friends, the Hettangian is the time in which the marine limestone, shales and clay Lias of western Europe were deposited. In British Columbia, Canada, we see the most diverse middle and late Hettangian (Early Jurassic) ammonite assemblages in the Queen Charlotte Islands (Haida Gwaii), an archipelago about 50 km off British Columbia's northern Pacific coast. In total, 53 ammonite taxa are described of which Paradasyceras carteri, Franziceras kennecottense, Pleuroacanthites charlottensis, Ectocentrites pacificus and Curviceras haidae are new.

In general, North American Early Jurassic ammonites are of Tethyan affinity or endemic to the eastern Pacific. For this reason, a separate zonation for the Hettangian and Sinemurian of the Western Cordillera of North America was established. Taylor et al. (2001), wrote up and published on much of this early research though, at the time, very little Canadian information was included.

Longridge, L. M., et al. “Three New Species of the Hettangian (Early Jurassic) Ammonite Sunrisites from British Columbia, Canada.” Journal of Paleontology, vol. 82, no. 1, 2008, pp. 128–139. JSTOR, www.jstor.org/stable/20144175. Accessed 27 Jan. 2020.

Tozer, ET (Tim): Marine Triassic faunas of North America: Their significance for assessing plate and terrane movements. Geol Rundsch 71, 1077-1104 (1982). https://doi.org/10.1007/BF01821119

Danner, W. (Ted): Limestone resources of southwestern British Columbia. Montana Bur. Mines & Geol., Special publ. 74: 171-185, 1976.

Davis, G., Monger, JWH & Burchfiel, BC: Mesozoic construction of the Cordilleran “collage”, central British Columbia to central California. Pacific Coast Paleography symposium 2, Soc. Economic Paleontologists and Mineralogists, Los Angeles: 1-32, 1978.

Gibson, DW: Triassic rocks of the Rocky Mountain foothills and front ranges of northeastern British Columbia and west-central Alberta. Geol. Surv. Canada Bull. 247, 1975.

Photo of the large belemnite (Atractites sp?) and ammonites (Sunrisites & Badouxia) from the Lower Jurassic (Late Hettangian), Last Creek Formation (Castle Pass member), Taseko Lakes area, British Columbia, Canada in the collection of the deeply awesome John Fam.

Photo: A drawer of Juvavites sp. in the collections of the Geological Survey of Canada. These rarely seen Upper Triassic (Carnian to Norian) ammonoids were collected over many decades by geologists of the Geological Survey of Canada from Northeastern British Columbia. Photo care of the deeply awesome John Fam.

Photo: Grambergia sp. from the Early Anisian (Middle Triassic) ammonoid biostratigraphy of northeastern British Columbia, Canada. Collection of Fossil Huntress.

Photo: Alsatites proaries, Coll. Reiter, Neoammoniten, 30 July 2011, 19:26:10

OH, SHINY. FOSSILS PRESERVED IN PYRITE

We sometimes find fossils preserved by pyrite. They are prized as much for their pleasing gold colouring as they are for their scientific value as windows into the past. 

Sometimes folk add a coating of brass to increase the aesthetic appeal — a practice is frowned upon in paleontological communities.

Pyrite, sometimes called Fool's Gold, is a brass-yellow mineral with a bright metallic lustre. I popped a photo of some pyrite below so you can see the characteristic shape of its cubic crystal system.

Fool's Gold has a chemical composition of iron sulfide (FeS2) and is the most common sulfide mineral. It forms at high and low temperatures usually in small quantities, in igneous, metamorphic, and sedimentary rocks. If these sulfide minerals are close at hand when a fossil is forming, they can infuse specimens, replacing their mineral content to beautiful effect.

When we find a fossil preserved with pyrite, it tells us a lot about the conditions on the seabed where the organism died. Pyrite forms when there is a lot of organic carbon and not much oxygen in the vicinity. 

The reason for this is that bacteria in sediment usually respire aerobically (using oxygen), however, when there is no oxygen, they respire without oxygen (anaerobic) typically using sulphate. 

Sulphate is a polyatomic anion with the empirical formula SO2−4. It is generally highly soluble in water. Sulfate-reducing bacteria, some anaerobic microorganisms, such as those living in sediment or near deep-sea thermal vents, use the reduction of sulfates coupled with the oxidation of organic compounds or hydrogen as an energy source for chemosynthesis.

The sulfide mineral Pyrite, FeS2

High quantities of organic carbon in the sediment form a barrier to oxygen in the water. This also works to encourage anaerobic respiration. Anaerobic respiration using sulphate releases hydrogen sulphide, which is one of the major components in pyrite. 

So, when we find a fossil preserved in pyrite, we know that it died and was buried in sediment with low quantities of oxygen and high quantities of organic carbon. 

If you have pyrite specimens and want to stop them from decaying, you can give them a 'quick' soak in water (hour max) then wash them off, dry thoroughly in a warm oven. 

Cool, then soak in pure acetone for a couple of days. Then soak in paraloid, a thermoplastic resin surface coating or acetone for a couple of days. Keep in a sealed container with a desiccant pack afterwards to keep them dry — or leave them out on display to enjoy knowing that the decay will come in time. We do this with cut flowers so why not fossils sometimes.

I have a friend who gives her pyrite fossils on display a quick thumb wipe with vasoline or petroleum jelly. I'm not sure if the hydrocarbons there will play nice over time but it will act as a protective barrier.  

RAINBOW AMMONITE: THE UNLIKELY LANDLORD

What you are seeing here is a protuberance extruding from the venter of Quenstedtoceras cf. leachi (Sowerby). It is a pathology in the shell from hosting immature bivalves that shared the seas with these Middle Jurassic, Upper Callovian, Lamberti zone fauna from the Volga River basin. The collecting site is the now inactive Dubki commercial clay quarry and brickyard near Saratov, Russia. 

The site has produced thousands of ammonite specimens. A good 1,100 of those ended up at the Black Hills Institute of Geological Research in Hill City, South Dakota. 

Roughly 1,000 of those are Quenstedtoceras (Lamberticeras) lamberti and the other 100 are a mix of other species found in the same zone. These included Eboraciceras, Peltoceras, Kosmoceras, Grossouvria, Proriceras, Cadoceras and Rursiceras. 

What is especially interesting is the volume of specimens — 167 Quenstedtoceras (Lamberticeras) lamberti and 89 other species in the Black Hills collection — with healed predation injuries. It seems Quenstedtoceras (Lamberticeras) lamberti are the most common specimens found here and so not surprisingly the most common species found injured. Of the 1,000, 655 of the Quenstedtoceras (Lamberticeras) lamberti displayed some sort of deformation or growth on the shell or had grown in a tilted manner. 

Again, some of the Q. lamberti had small depressions in the centre likely due to a healed bite and hosting infestations of the immature bivalve Placunopsis and some Ostrea. 

The bivalves thrived on their accommodating hosts and the ammonites carried on, growing their shells right up and over their bivalve guests. This relationship led to some weird and deformities of their shells. They grow in, around, up and over nearly every surface of the shell and seem to have lived out their lives there. It must have gotten a bit unworkable for the ammonites, their shells becoming warped and unevenly weighted. Over time, both the flourishing bivalves and the ammonite shells growing up and over them produced some of the most interesting pathology specimens I have ever seen.    

In the photo here from Emil Black, you can see some of the distorted shapes of Quenstedtoceras sp. Look closely and you see a trochospiral or flattened appearance on one side while they are rounded on the other. 

All of these beauties hail from the Dubki Quarry near Saratov, Russia. The ammonites were collected in marl or clay used in brick making. The clay particles suggest a calm, deep marine environment. One of the lovely features of the preservation here is the amount of pyrite filling and replacement. It looks like these ammonites were buried in an oxygen-deficient environment. 

The ammonites were likely living higher in the water column, well above the oxygen-poor bottom. An isotopic study would be interesting to prove this hypothesis. There's certainly enough of these ammonites that have been recovered to make that possible. It's estimated that over a thousand specimens have been recovered from the site but that number is likely much higher. But these are not complete specimens. We mostly find the phragmocones and partial body chambers. Given the numbers, this may be a site documenting a mass spawning death over several years or generations.

If you fancy a read on all things cephie, consider picking up a copy of Cephalopods Present and Past: New Insights and Fresh Perspectives edited by Neil Landman and Richard Davis. Figure 16.2 is from page 348 of that publication and shows the hosting predation quite well. 

Photos: Courtesy of the deeply awesome Emil Black. These are in his personal collection that I hope to see in person one day. 

It was his sharing of the top photo and the strange anomaly that had me explore more about the fossils from Dubki and the weird and wonderful hosting relationship between ammonites and bivalves. Thank you, my friend!

DRIFTWOOD CANYON FOSSIL BEDS / KUNGAX

White Eared Puffbird, Nystalus chacuru

Driftwood Canyon Provincial Park 

Driftwood Canyon Provincial Park covers 23 hectares of the Bulkley River Valley, on the east side of Driftwood Creek, a tributary of the Bulkley River, 10 km northeast of the town of Smithers in northern British Columbia. 

Wet'suwet'en First Nation

The parklands are part of the asserted traditional territory of the Wet'suwet'en First Nation which includes lands around the Bulkley River, Burns Lake, Broman Lake, and François Lake in the northwestern Central Interior of British Columbia. 

The Wetʼsuwetʼen are part of the Dakelh or Carrier First Nation, and in combination with the Babine First Nation are referred to as the Western Carrier. They speak Witsuwitʼen, a dialect of the Babine-Witsuwitʼen language which, like its sister language Carrier, is a member of the Athabaskan family.

Their oral history or kungax recounts a time when their ancestral village, Dizkle or Dzilke, once stood upstream from the Bulkley Canyon. This cluster of cedar houses on both sides of the river was said to be abandoned because of an omen of impending disaster. The exact location of the village has been lost but their stories live on. 

The neighbouring Gitxsan people of the Hazelton area have a similar tale, though the village in their version is referred to as Dimlahamid or Temlahan. Their house groups include the Gilseyhu or Big Frog Clan, the Laksilyu or Small Frog Clan, the Tsayu or Beaver Clan, the Gitdumden or Wolf and Bear Clan and the Laksamshu or Fireweed and Owl Clan.

Driftwood Canyon Fossil Beds

Driftwood Canyon's Fossil Beds record life in the earlier portion of the Eocene when British Columbia — and indeed our world — was much warmer than it is today. This site is recognized as one of the world’s most significant fossil beds. It provides us with a fascinating opportunity to understand the area’s evolutionary processes of both geology and biology. The park was created in 1967 by the donation of the land by the late Gordon Harvey (1913–1976) to protect fossil beds on the east side of Driftwood Creek. The beds were discovered around the beginning of the 20th century. 

Metasequoia, the Dawn Redwood

We have found plant, fish and insect fossil here including Metasequoia, the Dawn Redwood, alder, fossil salmon, wasps, water striders and vertebrate material. Bird feathers are infrequently collected from the shales; however, two bird body fossils have been found here.

In 1968, a bird body fossil was collected in the Eocene shales of the Ootsa Lake Group in Driftwood Canyon Provincial Park by Pat Petley of Kamloops. Pat Petley donated the specimen in 2000 to the Thompson Rivers University (TRU) palaeontology collections. This fossil bird specimen is tentatively identified as the puffbird, Piciformes Bucconidae, of the genus Primobucco.

Primobucco is an extinct genus of bird placed in its own family, Primobucconidae. The type species, Primobucco mcgrewi, lived during the Lower Eocene of North America. It was initially described by American paleo-ornithologist Pierce Brodkorb in 1970, from a fossil right-wing, and thought to be an early puffbird. However, the discovery of a further 12 fossils in 2010 indicate that it is instead an early type of roller.

Related fossils from the European Messel deposits have been assigned to the two species P. perneri and P. frugilegus. Two specimens of P. frugilegus have been found with seeds in the area of their digestive tract, which suggests that these birds were more omnivorous than the exclusively predaceous modern rollers. The Driftwood specimen has never been thoroughly studied. If there is a grad student out there looking for a worthy thesis, head on down to the Thompson Rivers University where you'll find the specimen on display.

Another fossil bird, complete with feathers, was collected at Driftwood Canyon in 1970, This one was found by Margret and Albrecht Klöckner who were travelling from Germany. Theirs is a well-travelled specimen, having visited many sites in BC as they toured around, then to Germany and finally back to British Columbia when it was repatriated and donated to the Royal British Columbia Museum in Victoria. I'm not sure if it is still on display or back in collections, but it was lovingly displayed back in 2008. There is a new grad student, Alexis, looking at Eocene bird feathers down at the RBCM, so perhaps it is once again doing the rounds. 

This second bird fossil is of a long-legged water bird and has been tentatively identified by Dr. Gareth Dyke of the University of Southampton as possibly from the order Charadriiformes, a diverse order of small to medium-ish water birds that include 350 species of gulls, plovers, sandpipers, terns, snipes, and waders. Hopefully, we'll hear more on this find in the future.

Tapirs and Tiny Hedgehogs

The outcrops at Driftwood Canyon are also special because they record a record of some of the first fossil mammals ever to be found in British Columbia at this pivotal point in time. Wee proto-hedgehogs smaller than your thumb lived in the undergrowth of that fossil flora. They shared the forest floor with an extinct tapir-like herbivore in the genus Heptodon that looked remarkably similar to his modern, extant cousins but lacked their pronounced snout (proboscis). I'm guessing that omission made him the more fetching of his lineage.

In both cases, it was a fossilized jaw bone that was recovered from the mud, silt and volcanic ash outcrops in this ancient lakebed site. And these two cuties are significant— they are the very first fossil mammals we've ever found from the early Eocene south of the Arctic.

How can we be sure of the timing? The fossil outcrops here are found within an ancient lakebed. Volcanic eruptions 51 million years ago put loads of fine dust into the air that settled then sank to the bottom of the lake, preserving the specimens that found their way here — leaves, insects, birds, mammals.

 As well as turning the lake into a fossil making machine (water, ash, loads of steady sediment to cover specimens and stave off predation...) the volcanic ash contains the very chemically inert (resistant to mechanical weathering) mineral zircon which we can date with uranium/lead (U/Pb). 

The U/Pb isotopic dating technique is wonderfully accurate and mighty helpful in dating geologic events from volcanic eruptions, continental movements to mass extinctions. This means we know exactly when these lovelies were fossilized and, in turn, their significance.

What To Know Before You Go

If you fancy a visit to Driftwood Canyon Park, the park is accessible from Driftwood Road from Provincial Highway 16. You are welcome to view and photograph the fossils found here but collecting is strictly forbidden. 

Driftwood Canyon is recognized as one of the world’s most significant fossil beds. It provides park users with a fascinating opportunity to understand the area’s evolutionary processes of both geology and biology. The day-use area is open from May 15 to September 2. There is a short, wheelchair-accessible interpretative trail that leads from the parking are to the fossil beds. Pets are welcome on leash. Signs along the trail provide information on fossils and local history. 

Below a cliff face at the end of the trail is a viewing area that has interpretive information and viewing area overlooking Driftwood Creek.

This park proudly operated by Mark and Anais Drydyk

Email: kermodeparks@gmail.com / Tel: 1 250 877-1482 or 1 250 877-1782

Driftwood Canyon Provincial Park Brochure: 

https://bcparks.ca/explore/parkpgs/driftwood_cyn/driftwood-canyon-brochure.pdf?v=1638723136455

A TASTE FOR STUDIES

Chelonia. Schildkröten by Ernst Haeckel, 1904

Care for some tarantula with that walrus? No? how about some Woolly mammoth?

While eating study specimens is not de rigueur today, it was once common practice for researchers in the 1700-1880s. 

The English naturalist, Charles Darwin belonged to an elite men's club dedicated to tasting exotic meats. In his first book, Darwin wrote almost three times as much about dishes like armadillo and tortoise urine as he did on the biogeography of his Galapagos finches. 

From his great love of gastronomy, I am surprised any of his tasty specimens made it back from his historic voyage on the HMS Beagle — particularly the turtles.

One of the most famous scientific meals occurred one Saturday evening on the 13th of January, 1951. This was at the 47th Explorers Club Annual Dinner (ECAD) when members purportedly dined on a frozen woolly mammoth. 

Commander Wendell Phillips Dodge was the promotor of the banquet. He sent out press notices proclaiming the event's signature dish would be a selection of prehistoric meat. Whether Dodge did this simply to gain attendees or play a joke remains a mystery. 

The prehistoric meat was supposedly found at Woolly Cove on Akutan in the Aleutians Islands of Alaska, USA, by the eminent polar explorers' Father Bernard Rosecrans Hubbard, American geologist, explorer sometimes called the Glacier Priest, and polar explorer Captain George Francis Kosco of the United States Navy.

Fried Tarantula & Goat Eyeballs

This much-publicized meal captured the public’s imagination and became an enduring legend and source of pride for the Club, popularizing an annual menu of exotics that continues today. The Club is well-known for its notorious hors d’oeuvres like fried tarantulas and goat eyeballs as it is for its veritable whose who of notable members — Teddy Roosevelt, Neil Armstrong, Buzz Aldrin, Roy Chapman Andrews, Thor Heyerdahl, James Cameron.

The Yale Peabody Museum holds a sample of meat preserved from the 1951 meal, interestingly labelled as a South American Giant Ground Sloth, Megatherium, not Mammoth. The specimen of meat from that famous meal was originally designated BRCM 16925 before a transfer in 2001 from the Bruce Museum to the Yale Peabody Museum of Natural History (New Haven, CT, USA) where it gained the number YPM MAM 14399.

The specimen is now permanently deposited in the Yale Peabody Museum with the designation YPM HERR 19475 and is accessible to outside researchers. The meat was never fixed in formalin and was initially stored in isopropyl alcohol before being transferred to ethanol when it arrived at the Peabody Museum. DNA extraction occurred at Yale University in a clean room with equipment reserved exclusively for aDNA analyses.

In 2016, Jessica Glass and her colleagues sequenced a fragment of the mitochondrial cytochrome-b gene and studied archival material to verify its identity, which if genuine, would extend the range of Megatherium over 600% and alter views on ground sloth evolution. 

Mammoth, Megatherium — Green Sea Turtle

Their results showed that the meat was not Mammoth or Megatherium, but a bit of Green Sea Turtle, Chelonia mydas. So much for elaborate legends. The prehistoric dinner was likely meant as a publicity stunt. 

Glass's study emphasizes the value of museums collecting and curating voucher specimens, particularly those used for evidence of extraordinary claims. Not so long before Glass et al. did their experiment, a friend's mother (and my kayaking partners) served up a venison steak from her freezer to dinner guests in Castlegar that hailed from 1978. Tough? Inedible? I have it on good report that the meat was surprisingly divine.

Reference: Glass, J. R., Davis, M., Walsh, T. J., Sargis, E. J., & Caccone, A. (2016). Was Frozen Mammoth or Giant Ground Sloth Served for Dinner at The Explorers Club?. PloS one, 11(2), e0146825. https://doi.org/10.1371/journal.pone.0146825

Image: Chelonia. Schildkröten by Ernst Haeckel, 1904, Prints & Photographs Division, Library of Congress, LC-DIG-ds-07619.

Join the Explorer's Club

Fancy yourself an explorer who should join the club? Here is a link to their membership application. The monied days of old are still inherent, but you will be well pleased to learn you can now join for as little as $50 US.

Link: https://www.explorers.org/wp-content/uploads/Membership-Application_2021-11-19.pdf

VANCOUVER ISLAND'S HASLAM FORMATION

Steller's Jay, Cyanocitta stelleri

One of the classic Vancouver Island fossil localities is the Santonian-Maastrichtian, Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake, Nanaimo, British Columbia, Canada.

The quarry is no longer active as such though there is a busy little gravel quarry a little way down the road closer to Ammonite falls near Benson Creek Falls.

Today it is an active motocross site and remains one of the classic localities of the Nanaimo Group. We find well-preserved nautiloids and ammonites — Canadoceras, Pseudoschloenbachia, Epigoniceras — the bivalves — Inoceramus, Sphenoceramus— gastropods, and classic Nanaimo Group decapods — Hoploparia, Linuparus. We also find fossil fruit and seeds which tell the story of the terrestrial history of Vancouver Island.

Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake

It was John Fam, Vice-Chair, Vancouver Island Paleontological Society (VanPS), who originally told me about the locality. John is one of the most delightful and knowledgeable people you'd be well-blessed to meet.

While he lived on Vancouver Island, he was an active member of the VanPS back when I was Chair. Several of the best joint VIPS/VanPS paleontological expeditions were planned with or instigated by his passion for fossils. I tip my hat to him for his passion and shared love of all things paleo.

John grew up 15 minutes from the motocross locality and used to collect there a few times a week with his father. John has wonderful parents and since marrying his childhood sweetheart, the amazing Grace, those excellent genetics, curiosity and love of fossils are now being passed to a new generation. It's lovely to see John and Grace continuing tradition with two boys of their own.

I met John way back then and did an overnight at his parent's house the Friday before a weekend field trip to Jurassic Point. It was a joy to have him walk me through his collections and tell his stories from earlier years. After learning about the site from John, I headed up to the Motocross Pit with my Uncle Doug. He was a delightful man who grew up on the coast and had explored much of it but not the fossil site just 10-minutes from his home. It was wonderful to walk through time with him so many years ago and then again solo this past year with sadness in my belly that one of the best I've ever known has left this Earth.

Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake

There were some no trespassing signs up but no people around, so I walked the periphery looking for the bedrock of the Haslam.

The rocks we find here were laid down south of the equator as small, tropical islands. They rode across the Pacific heading north and slightly east over the past 80 million years to where we find them today.

Jim Haggart and Peter Ward have done much to increase our understanding of the molluscan fauna of the Nanaimo Group. Personally, both personify the charming Indiana Jones school of rugged manly palaeontologists you picture in popular film. Professionally, their singular contributions and collaborative efforts have helped shape our understanding of the correlation of Nanaimo Group fauna to those we find in the Gulf Islands of British Columbia and down in the San Juan Islands of Washington State.

Their work builds on the work of Usher (1952), Matsumoto (1959a, 1959b) and Mallory (1977). A healthy nod goes out to the work of Muller and Jeletzky (1970) for untangling the lithostratigraphic and biostratigraphic foundation for our knowledge of the Nanaimo Group.

Candoceras yokoyama, Photo: John Fam, VanPS

As I walked along the bedrock of the Haslam, a Steller's Jay, Cyanocitta stelleri, followed me from tree to tree making his guttural shook, shook, shook call. Instructive, he seemed to be encouraging me, timing his hoots to the beat of my hammer. Vancouver Island truly has glorious flora and fauna.

Fancy some additional reading? Check out a paper published in the Journal of Paleontology back in 1989 by Haggard and Ward on new Nanaimo Group Ammonites from British Columbia and Washington State.

In it, they look at the ammonite species Puzosia (Mesopuzosia) densicostata Matsumoto, Kitchinites (Neopuzosia) japonicus Spath, Anapachydiscus cf. A. nelchinensis Jones, Menuites cf. M. menu (Forbes), Submortoniceras chicoense (Trask), and Baculites cf. B. boulei Collignon are described from Santonian--Campanian strata of western Canada and northwestern United States.

Stratigraphic occurrences and ranges of the species are summarized and those taxa important for correlation with other areas in the north Pacific region and Late Cretaceous ammonite fauna of the Indo-Pacific region. Here's the link: https://www.jstor.org/stable/1305358?seq=1

Peter Ward is a prolific author, both of scientific papers and more popularized works. I highly recommend his book Gorgon: Paleontology, Obsession, and the Greatest Catastrophe in Earth's History. It is an engaging romp through a decade's research in South Africa's Karoo Desert.

Photo: Candoceras yokoyamai from Upper Cretaceous Haslam formation (Lower Campanian) near Nanaimo, British Columbia. One of the earliest fossils collected by John Fam (1993). Prepared using only a cold chisel and hammer. Photo & collection of John Fam, VIPS.

CRANBROOK EAGER FORMATION: TUZOIA

This specimen of the arthropod Tuzoia sp. is from the Lower Cambrian Eager Formation of British Columbia. 

Tuzoia is an extinct genus of large bivalved arthropod known from Early to Middle Cambrian marine environments in what is now the Burgess Shale, Emu Bay Shale, Kaili, the Rockslide Formation, the Spence Shale, Wheeler Formation, and Marjum Formation, and the Kinzers Formation. 

As you can see here, they are also found in the Lower Cambrian Eager Formation near Cranbrook British Columbia. 

This particular specimen is from the Rifle Range outcrop near Cranbrook where you can also find numerous fragments and complete specimens of the olenellid trilobites Ollenellus sp. and the larger, more robust Wanneria dunnae, along with Mesonacis eagerensis (Best, 1952).

The site outcrops at a few locations as you head east out of Cranbrook towards Fort Steele. The first trilobites were discovered with the building of the Kootenay Highway connecting Cranbrook to Fort Steele and beyond. Several other localities, including the outcrops at the Silhouette Rife Range — which is literally on a Rifle Range where folks go to shoot at things — is a shade older than the Middle Cambrian Burgess Shale but the fauna here is much less varied. 

The site has been known and collected since the 1920s. Back in the day, fossil collecting was a family affair with folks heading out in their lightly coloured finery to picnic and surface collect the eroding exposures. Cranbrook local, Clement Hungerford Pollen was an engineer and avocational palaeontologist. He promoted collecting the exposures of the Eager Formation around 1921. As a pedigreed Englishman of considerable means, he had invested in the Kootenay Central Railway, revitalizing the town by opening up railway access within the region.

FOSSIL COLLECTING AT THE MYSTERIOUS CREEK FORMATION

Located three hours east of Vancouver, most folks head to Harrison Lake to enjoy its crisp waters, soak in the hot springs, camp or four-wheel-drive immersed in rugged scenery, or look for the elusive Sasquatch reported to live in the area. 

But some come to Harrison Lake and miss the town entirely. Instead, they favour the upper west side of the lake and the fossiliferous bounty found here.

Indeed, this is the perfect location for local citizen scientists to strut their stuff. Harrison is a perfect family day trip, where you can discover wonderful marine fossil specimens as complete or partially crushed fossilized shells embedded in rock. 

Amazingly, we can find them at all. These beauties range in age from Jurassic to Cretaceous, with most being Lower Callovian, meaning the ammonites here swam our ancient oceans more than 160 million years ago. 

The area around Harrison Lake has been home to the Sts’ailes, a sovereign Coast Salish First Nation for thousands of years. Sts’ailes’ means, “the beating heart,” and it sums up this glorious wilderness perfectly. They describe their ancient home as Xa’xa Temexw or Sacred Earth. 

With the settling of Canada, Geologists began exploring the area in the 1880s, calling upon the Sts’ailes to help them look for coal and a route for the Canadian Pacific Railway. Coal was the aim, but happily, they also found fossils. Sacred Earth, indeed.  

Belemnite Fossils

In my favourite outcrops, you can find large, smooth inflated Jurassic ammonites along with their small grey and brown cousins. 

Further up the road, you will see Cretaceous cigar-shaped squid-like cephalopods called Belemnites, and the bivalve (clam) Buchia — gifts deposited by glaciers. Here are the most common.

Ammonites

Almost all of the ammonite specimens found near Harrison Lake are the toonie sized Cadoceras (Paracadoceras) tonniense with well-preserved outer whorls but flattened inner whorls. We find semi-squished elliptical specimens here, too. If you see a large, smooth, inflated grapefruit-sized ammonite, you are holding a rare prize — a Cadoceras comma ammonite, the macroconch or female of the species.  

Ammonites were predatory, squid-like creatures that lived inside coil-shaped shells. Like other cephalopods, ammonites had sharp, beak-like jaws inside a ring of squid-like tentacles that extended from their shells. They used these tentacles to snare prey — plankton, vegetation, fish and crustaceans — similar to the way a squid or octopus hunts today.

Within their shells, ammonites had a number of chambers called septa filled with gas or fluid, and they were interconnected through a wee air tube. By pushing air in or out, they were able to control their buoyancy. 

These small but mighty marine predators lived in the last chamber of their shell and continuously built new shell material as they grew. As they added each new chamber, they would move their squid-like body down to occupy the final outside chamber.

Interestingly, ammonites from Harrison Lake are quite similar to the ones found within the lower part of the Chinitna Formation near Cook Inlet, Alaska, and Jurassic Point, Kyuquot, on the west coast of Vancouver Island — some of the most beautiful places on Earth. 

Buchia (bivalve) Clams

The bivalve or clam Buchia are commonly found at Harrison Lake. You will see them cemented together en masse. . They populated Upper Jurassic–Lower Cretaceous waters like a team sport. When they thrived, they really thrived, building up large coquinas of material. Large boulders of Buchia cemented together en masse hitched a ride with the glaciers and were deposited around Harrison Lake. Some kept going and we find similar erratics or glacier-deposited boulders as far south as Washington state. 

Buchia is used as Index Fossils. Index fossils help us to figure out the age of the rock we are looking at because they are abundant, populate an area en masse, and then die out quickly. In other words, they make it easy to identify a geologic time span.

So what does this mean to you? Now, when you are out and about with friends and discover rocks with Buchia, or made entirely of Buchia, you can say, “Oh, this looks to be Upper Jurassic or Lower Cretaceous. Come take a look! We're likely the first to lay eyes on this little clam since dinosaurs roamed the Earth.” 

Fossil Collecting at Harrison Lake Fossil Field Trip — Getting there

This Harrison Lake site is a great day trip from Vancouver or the Fraser Valley. You will need a vehicle with good tires for travel on gravel roads. Search out the route ahead of time and share your trip plan with someone you trust. If you can pre-load the Google Earth map of the area, you will thank yourself. 

Heading east on from Vancouver, it will take you 1.5-2 hours to reach Harrison Mills. 

Access Forestry Road #17 at the northeast end of the parking lot from the Sasquatch Inn at 46001 Lougheed Hwy, Harrison Mills. From there, it will take about an hour to get to the site. Look for signs for the Chehalis River Fish Hatchery to get you started. 

Drive 30 km up Forestry Road #1, and stop just past Hale Creek at 49.5° N, 121.9° W (paleo-coordinates 42.5° N, 63.4° W) on the west side of Harrison Lake. You will see Long Island to your right. 

The first of the yummy fossil exposures are just north of Hale Creek on the west side of the road. Keep in mind that this is an active logging road, so watch your kids and pets carefully. Everyone should be wearing something bright so they can be easily spotted.

How to Spot the Fossils

The fossils here are easily collected—look in the bedrock and in the loose material that gathers in the ditches. Specimens will show up as either dark grey, grey-brown or black. Look for the large, dark-grey boulders the size of smart cars packed with Buchia. 

And while you are at it, be on the lookout for anything that looks like bone. This site is also ripe for marine reptiles—think plesiosaur, mosasaur and elasmosaur. As a citizen scientist and budding palaeontologist, you might just find something new!

What to Know Before You Go

Fill your gas tank and pack a tasty lunch. As with all trips into British Columbia's wild places, dress for the weather. You will need hiking boots, rain gear, gloves, eye protection, and a good geologic hammer and rock (cold) chisel. 

Wear bright clothing and keep your head covered. Slides are common, and you may start a few if you hike the cliffs. If you are with a group, those collecting below may want to consider hardhats in case of rockfall — chunks of rock the size of your fist up to the size of a grapefruit. They pack a punch. 

Bring a colourful towel or something to put your keepers on. Once you set rock down, it can be hard to find again given the terrain. I take the extra precaution of spraying the ends of my hammers and chisels with yellow fluorescent paint, as I have lost too many in the field. You will also want to bring a camera for the blocks of Buchia that are too big to carry home. 

Identifying Your Treasures

When you have finished for the day, compare your treasures to see which ones you would like to keep. In British Columbia, you are a steward of the fossil, which means they belong to the province, but you can keep them safe. You are not allowed to sell or ship them outside British Columbia without a permit. 

Once you get home, wash and identify your finds. Harrison Lake does not have a large variety of fossil fauna, so this should not be difficult. If your find is coiled and round, it is an ammonite. If it is long and straight, it is a belemnite. And if it looks like a wee fat baby oyster, it is Buchia. This is not always true, but mostly true.

What about collecting fossils in all seasons?. Everyone has a preference. I prefer not to collect in the snow, but I have done it. While sunny days are lovely, it can also be easier to see the specimens when the rock is wet. So, do we do this in the rain? Heck, yeah! 

In torrential rain? 

Yes — once you are hooked, but for your casual friends or the kiddos, the answer is likely no. Choose your battles. They may come with you, but a cold day getting soaked is no fun. 

In time, you will find your inner fossil geek — probably with your first find. And that's just the tip of the iceberg. First, it will be you, then your kids, your friends and then your neighbour. Once you start, it is easy to get hooked. Fossil addiction is real, and the only cure is to get out there and do it some more. You've got this!

References and further information:

A. J. Arthur, P. L. Smith, J. W. H. Monger and H. W. Tipper. 1993. Mesozoic stratigraphy and Jurassic palaeontology west of Harrison Lake, southwestern British Columbia. Geological Survey of Canada Bulletin 441:1-62

R. W. Imlay. 1953. Callovian (Jurassic) ammonites from the United States and Alaska Part 2. The Alaska Peninsula and Cook Inlet regions. United States Geological Survey Professional Paper 249-B:41-108

An overview of the tectonic history of the southern Coast Mountains, British Columbia; Monger, J W H; in, Field trips to Harrison Lake and Vancouver Island, British Columbia; Haggart, J W (ed.); Smith, P L (ed.). Canadian Paleontology Conference, Field Trip Guidebook 16, 2011 p. 1-11 (ESS Cont.# 20110248).

FOSSIL FLORA AND FAUNA OF MADAGASCAR

Aioloceras besairiei (Collingnon, 1949)

A stunning example of the internal suturing with calcite infill in this sliced Aioloceras besairiei (Collingnon, 1949) ammonite from the Upper Cretaceous (Lower Albian) Boeny region of Madagascar. 

This island country is 400 kilometres off the coast of East Africa in the Indian Ocean and a wonderful place to explore off the beaten track.

Madagascar has some of the most spectacular of all the fossil specimens I have ever seen. This beauty is no exception. The shell has a generally small umbilicus, arched to acute venter, and typically at some growth stage, falcoid ribs that spring in pairs from umbilical tubercles, usually disappearing on the outer whorls. I had originally had this specimen marked as a Cleoniceras besairiei, except Cleoniceras and Grycia are not present in Madagascar. 

This lovely, seen in cross-section, is now far from home and in the collection of a wonderful friend. It is an especially lovely example of the ammonite, Aioloceras besairiei, making it a beudanticeratinae. Cleoniceras and Grycia are the boreal genera. If you'd like to see (or argue) the rationale on the name, consider reading Riccardi and Medina's riveting work from back in 2002, or Collingnon from 1949.

The beauty you see here measures in at a whopping 22 cm, so quite a handful. This specimen is from the youngest or uppermost subdivision of the Lower Cretaceous. I'd originally thought this locality was older, but dating reveals it to be from the Lower Albian, so approximately 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma.

Aioloceras are found in the Cretaceous of Madagascar at geo coordinates 16.5° S, 45.9° E: paleo-coordinates 40.5° S, 29.3° E.; and in four localities in South Africa: at locality 36, near the Mzinene River at 28.0° S, 32.3° E: paleo-coordinates 48.6° S, 7.6° E. 

We find them near the Mziene River, at a second locality north of Hluhluwe where the Mzinene Formation overlies the Aptian-Albian Makatini Formation at 28.0° S, 32.3° E: paleo-coordinates 48.6° S, 7.6° E; and at Haughton Z18, on the Pongola River in the Albian III, Tegoceras mosense beds at 27.3° S, 32.2° E: paleo-coordinates 48.0° S, 7.8° E.

If you happen to be trekking to Madagascar, know that it's big. It’s 592,800 square kilometres (or  226,917 square miles), making it the fourth-largest island on the planet — bigger than Spain, Thailand, Sweden and Germany. The island has an interesting geologic history.

Although there has been a geological survey, which was active extending back well into French colonial times, in the non-French-speaking world our geological understanding of the island is still a bit of a mystery. 

Plate tectonic theory had its beginnings in 1915 when Alfred Wegener proposed his theory of "continental drift." 

Wegener proposed that the continents ploughed through the crust of ocean basins, which would explain why the outlines of many coastlines (like South America and Africa) look like they fit together like a puzzle. Half a century after Wegener there is still no agreement as to whether in continental reconstructions Madagascar should be placed adjacent to the Tanzanian coast to the north (e.g., McElhinny and Embleton,1976), against the Mozambique-Natal coast (Flores 1970), or basically left where it is (Kent 1974, Nairn 1978).

There have been few attempts apart from McKinley’s (1960) comparison of the Karoo succession of southwestern Tanzania with that of Madagascar to follow the famous geological precept of “going to sea.” One critical reason is that although there may be a bibliography of several thousand items dealing with Madagascan geology as Besairie (1971) claims, they are items not generally available to the general public. The vital information gained of the geology of the offshore area by post-World War II petroleum exploration has remained largely proprietary. 

Without this data to draw upon, our understanding remains incomplete. I don't actually mind a bit of a mystery here. It is interesting to speculate on how these geologic puzzle pieces fit together and wait for the big reveal. Still, we have good old Besairie from his 1971, Geologie de Madagascar, and a later précis (Besairie, 1973).

We do know that Madagascar was carved off from the African-South American landmass early on. The prehistoric breakup of the supercontinent Gondwana separated the Madagascar–Antarctica–India landmass from the Africa–South America landmass around 135 million years ago. Madagascar later split from India about 88 million years ago, during the Late Cretaceous, so the native plants and animals on the island evolved in relative isolation. 

It is a green and lush island country with more than its fair share of excellent fossil exposures. Along the length of the eastern coast runs a narrow and steep escarpment containing much of the island's remaining tropical lowland forest. If you could look beneath this lush canopy, you'd see rocks of the Precambrian age stretching from the east coast all the way to the centre of the island. The western edge is made up of sedimentary rock from the Carboniferous to the Quaternary.

Red-Tailed Lemurs, Waiwai & Hedgehog

Madagascar is a biodiversity hotspot. Just as Darwin's finches on the Galápagos were isolated, evolving into distinct species (hello, adaptive radiation), over 90% of the wildlife from Madagascar is found nowhere else. 

The island's diverse ecosystems, like so many on this planet, are threatened by Earth's most deadly species, homo sapien sapiens. 

We arrived back in 490 CE and have been chopping down trees and eating our way through the island's tastier populations ever since. Still, they have cuties like this Red-Tailed Lemur. Awe, right?

Today, beautiful outcrops of wonderfully preserved fossil marine fauna hold appeal for me. The material you see from Madagascar is distinctive — and prolific.

Culturally, you'll see a French influence permeating the language, architecture and legal process. There is a part of me that pictures these lovely Lemurs chatting away in French. "Ah, la vache! Regarde le beau fossile, Hérissonne!"

We see the French influence because good 'ol France invaded sleepy Madagascar back in 1883, during the first Franco-Hova War. Malagasy (the local Madagascarian residents) were enlisted as troops, fighting for France in World War I.  During the Second World War, the island was the site of the Battle of Madagascar between the Vichy government and the British. By then, the Malagasy had had quite enough of colonization and after many hiccuping attempts, reached full independence in 1960. Colonization had ended but the tourist barrage had just begun. You can't stop progress.

If you're interested in learning more about this species, check out the Treatise on Invertebrate Paleontology, Part L (Ammonoidea). R.C. Moore (ed). Geological Soc of America and Univ. Kansas Press (1957), p L394. Or head over to look at the 2002 paper from Riccardi and Medina. 2002. Riccardi, A., C. & Medina, F., A. The Beudanticeratinae and Cleoniceratinae (Ammonitina) from the Lower Albian of Patagonia in Revue de Paléobiologie - 21(1) - Muséum d’Histoire Naturelle de la ville de Genève, p 313-314 (=Aioloceras besairiei (COLLIGNON, 1949). You have Bertrand Matrion to thank for the naming correction. Good to have friends in geeky places!

Collignon, M., 1933, Fossiles cenomaniens d’Antmahavelona (Province d’ Analalave, Madagascar), Ann. Geol. Serv. Min. Madagascar, III, 1934 Les Cephalopods du Trias inferieur de Madagascar, Ann. Paleont. XXII 3 and 4, XXII 1.

Besairie, H., 1971, Geologie de Madagascar, 1. Les terrains sedimentaires, Ann. Geol. Madagascar, 35, p. 463.

J. Boast A. and E. M. Nairn collaborated on a chapter in An Outline of the Geology of Madagascar, that is very readable and cites most of the available geologic research papers. It is an excellent place to begin a paleo exploration of the island.

If you happen to parle français, check out: Madagascar ammonites: http://www.ammonites.fr/Geo/Madagascar.htm

TOROSAURUS WALKING THROUGH THE FOG

Torosaurus was a ceratopsian dinosaur that lived during the Cretaceous. These fellows look very similar to their Triceratops cousins but are an entirely different species in their own right. 

In 1891, two years after the naming of Triceratops, a pair of ceratopsian skulls with elongated frills bearing holes were found in southeastern Wyoming, Niobrara County, by John Bell Hatcher. 

Hatcher's employer, palaeontologist Professor Othniel Charles Marsh, coined the genus Torosaurus for them. While an estimated 2,000 Triceratops specimens have been collected from the American West, only seven partial skulls of Torosaurus have been found, so they are pretty rare.

Like Triceratops, they had massive skulls. Torosaurus had one of the largest skulls of any known land animal, with the frilled skull reaching 2.77 metres (9.1 ft) in length. Torosaurus were about the same size as Triceratops. 

They had an elongated frill with large openings (fenestrae), long squamosal bones of the frill with a trough on their upper surface, and the presence of five or more pairs of hornlets (epoccipitals) on the back of the frill. Torosaurus also lacked the long nose horn seen in Triceratops prorsus, and instead resembled the earlier and more basal Triceratops horridus in having a short nose horn. Three species have been named, Torosaurus latus, T. gladius and T. utahensis. T. gladius is no longer considered a valid species.

Wyoming Outcrops

The individuals referred to as Torosaurus are all large, comparable to the largest Triceratops specimens. Due to the elongated frill, especially the skull length is considerable. Hatcher estimated the skull of YPM 1830 at 2.2 metres, and of YPM 1831 at 2.35 metres. 

In 1933, Richard Swann Lull increased this to 2.4 metres and 2.57 metres respectively. Based on this, Torosaurus was thought, albeit briefly, to have the longest skull of any known land animal. 

Sixty-five years later in 1998, Thomas Lehman claimed that a Pentaceratops specimen possessed a partial skull that would have been 2.9 metres long in life. This was again doubted by Nicholas Longrich who in 2011 named this exemplar as a separate genus Titanoceratops and concluded its skull had been reconstructed as too long.

In 2006, Andrew Farke, a palaeontologist at the Alf Museum of Paleontology in South Dakota, pointed out that the new skulls described by him were on average even longer than Hatcher's original two: MOR 1122 has a length of 252 centimetres and MOR 981 of 277 centimetres.

Farke’s research interests focus on exploring the Cretaceous continental ecosystems of North America, particularly the ceratopsian (horned) dinosaurs, with active fieldwork in California and Wyoming.

In 2006, Farke published some diagnostic traits of Torosaurus. The frill is extremely long in comparison to the remainder of the skull. The rear, parietal, and edge of the frill bear ten or more epiparietals, triangular osteoderms. A midline epiparietal is absent; likewise, no osteoderm straddles the parietal-squamosal boundary. The parietal bone is thin. It is pierced by parietal fenestrae in the form of circular or transversely oval openings. The parietal bone is about 20% wider than long. 

Farke also identified a single trait in which T. latus differed from both Triceratops horridus and T. utahensis: its squamosal bore a conspicuous ridge on the edge with the parietal combined with a deep longitudinal trough parallel to it.

Farke pointed out that the known Torosaurus specimens are rather variable. The orbital "brow" horns are sometimes large and curved to the front, as with MOR 981, and sometimes short and straight as shown by MOR 1122 and ANSP 15191. 

Also, the position of these horns differs: often they are located directly on top of the eye socket but with YPM 1831 they originate at the rear edge of the orbit. Likewise, there is a variation in the form of the nose horn. YPM 1831 and to a lesser extent YPM 1830 have a straight upright nasal horn but MOR 981, ANSP 15192 and especially MOR 1122 at most possess a low bump. The frill too differs. ANSP 15192 and YPM 1830 have a shield curving upwards at the rear, but the frill of YPM 1831 is nearly flat, though this could be an artefact of restoration. 

The frill of YPM 1831 is also heart-shaped, with a clear midline notch, whereas the rear edge of the other specimens is straight. The frill proportions are quite variable: with YPM 1831 the length-width ratio is 1.26 but MOR 981 has a shield 2.28 times longer than wide. The number of epiparietals is difficult to assess as most fossils seem to have lost them. MOR 981 and MOR 1122 have ten and twelve epiparietals respectively. YPM 1831 has been restored with a fontanelle in the skull roof, which possibly is authentic. Farke also concluded that the degree of variability did not exceed that shown by related genera.

Farke stressed that, apart from the frill, no systematic differences could be found between Torosaurus and Triceratops. All Torosaurus specimens are similar in that they lack a truly long nasal horn and a horizontal arterial groove at the front base of that horn, but Triceratops fossils with the same combination of traits are not uncommon. 

In 2008, Hunt concluded that T. utahensis, contrary to T. latus but similar to Triceratops, possessed a midline epiparietal.

BIG HEAD LITTLE HOOVES: TRICERATOPS

This cutie is a Triceratops. The name means three-horned face in Greek but they might have been named Little Hooves instead as a nod to their weight-bearing fingers and toes that ended in sweet little hooves. 

Three of their five fingers and all of their toes end in a broad, flat-shaped hoof bone with a horny covering. 

Their hooves helped to protect their toes from wear and tear and support their heavy 5-ton bodies as they plodded along munching on cycads and palm fronds in the Late Cretaceous. These ceratopsid dinosaurs loved their plants. They used their beak-like jaws and slicing teeth to pluck and chew tasty foliage. Picture an animal about the size of an elephant, now lose the trunk, add the big head frill and horns. That's them!

Bearing a large bony frill, three horns on the skull, and a large four-legged body, exhibiting convergent evolution with rhinoceroses and bovines, Triceratops is one of the most recognizable of all dinosaurs and the most well-known ceratopsid. It was also one of the largest, up to 8–9 metres (26–30 ft) long and 5–9 metric tons (5.5–9.9 short tons) in body mass. 

It shared the landscape with and was most likely preyed upon by Tyrannosaurus, though it is less certain that two adults did battle in the fanciful manner often depicted in museum displays and popular images. The functions of the frills and three distinctive facial horns on its head have long inspired debate. Traditionally, these have been viewed as defensive weapons against predators. More recent interpretations find it probable that these features were primarily used in species identification, courtship, and dominance display, much like the antlers and horns of modern ungulates.

Triceratops was traditionally placed within the "short-frilled" ceratopsids, but modern cladistic studies show it to be a member of the Chasmosaurinae which usually have long frills. Two species, T. horridus and T. prorsus, are considered valid today, from the seventeen species that have ever been named. Research published in 2010 concluded that the contemporaneous Torosaurus, a ceratopsid long regarded as a separate genus, represents Triceratops in its mature form. This view has been disputed; further data is needed to settle the debate.

Triceratops has been documented by numerous remains collected since the genus was first described in 1889 by American paleontologist Othniel Charles Marsh. Specimens representing life stages from hatchling to adult have been found. As the archetypal ceratopsid, Triceratops is one of the most popular dinosaurs, and has been featured in film, postal stamps, and many other types of media.

AVES; LIVING DINOSAURS

Cassowary, Casuariiformes

Wherever you are in the world, it is likely that you know your local birds. True, you may call them des Oiseaux, pássaros or uccelli — but you'll know their common names by heart.

You will also likely know their sounds. The tweets, chirps, hoots and caws of the species living in your backyard.

Birds come in all shapes and sizes and their brethren blanket the globe. It is amazing to think that they all sprang from the same lineage given the sheer variety. 

If you picture them, we have such a variety on the planet — parrots, finches, wee hummingbirds, long-legged waterbirds, waddling penguins and showy toucans. 

But whether they are a gull, hawk, cuckoo, hornbill, potoo or albatross, they are all cousins in the warm-blooded vertebrate class Aves. The defining features of the Aves are feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. The best features, their ability to dance, bounce and sing, are not listed, but it is how I see them in the world.

These modern dinosaurs live worldwide and range in size from the 5 cm (2 in) bee hummingbird to the 2.75 m (9 ft) ostrich. 

There are about ten thousand living species, more than half of which are passerine, or "perching" birds. Birds have wings whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds.

Wings evolved from forelimbs giving birds the ability to fly

Wings, which evolved from forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. 

The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimming.

Wee Feathered Theropod Dinosaurs

We now know from fossil and biological evidence that birds are a specialized subgroup of theropod dinosaurs, and more specifically, they are members of Maniraptora, a group of theropods that includes dromaeosaurs and oviraptorids, amongst others. As palaeontologists discover more theropods closely related to birds, the previously clear distinction between non-birds and birds has become a bit muddy.

Recent discoveries in the Liaoning Province of northeast China, which include many small theropod feathered dinosaurs — and some excellent arty reproductions — contribute to this ambiguity. 

Still, other fossil specimens found here shed a light on the evolution of Aves. Confuciusornis sanctus, an Early Cretaceous bird from the Yixian and Jiufotang Formations of China is the oldest known bird to have a beak.

Like modern birds, Confuciusornis had a toothless beak, but close relatives of modern birds such as Hesperornis and Ichthyornis were toothed, telling us that the loss of teeth occurred convergently in Confuciusornis and living birds.

The consensus view in contemporary palaeontology is that the flying theropods, or avialans, are the closest relatives of the deinonychosaurs, which include dromaeosaurids and troodontids.

Together, these form a group called Paraves. Some basal members of this group, such as Microraptor, have features that may have enabled them to glide or fly. 

The most basal deinonychosaurs were wee little things. This raises the possibility that the ancestor of all paravians may have been arboreal, have been able to glide, or both. Unlike Archaeopteryx and the non-avialan feathered dinosaurs, who primarily ate meat, tummy contents from recent avialan studies suggest that the first avialans were omnivores. Even more intriguing...

Avialae, which translates to bird wings, are a clade of flying dinosaurs containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to modern birds — Aves — than to deinonychosaurs, though alternative definitions are occasionally bantered back and forth.

The Earliest Avialan: Archaeopteryx lithographica

Archaeopteryx, bird-like dinosaur from the Late Jurassic

Archaeopteryx lithographica, from the late Jurassic Period Solnhofen Formation of Germany, is the earliest known avialan that may have had the capability of powered flight. 

However, several older avialans are known from the Late Jurassic Tiaojishan Formation of China, dating to about 160 million years ago.

The Late Jurassic Archaeopteryx is well-known as one of the first transitional fossils to be found, and it provided support for the theory of evolution in the late 19th century. 

Archaeopteryx was the first fossil to clearly display both traditional reptilian characteristics — teeth, clawed fingers, and a long, lizard-like tail—as well as wings with flight feathers similar to those of modern birds. It is not considered a direct ancestor of birds, though it is possibly closely related to the true ancestor.

Unlikely yet true, the closest living relatives of birds are the crocodilians. Birds are descendants of the primitive avialans — whose members include Archaeopteryx — which first appeared about 160 million years ago in China.

DNA evidence tells us that modern birds — Neornithes — evolved in the Middle to Late Cretaceous, and diversified dramatically around the time of the Cretaceous–Paleogene extinction event 66 mya, which killed off the pterosaurs and all non-avian dinosaurs.

In birds, the brain, especially the telencephalon, is remarkably developed, both in relative volume and complexity. Unlike most early‐branching sauropsids, the adults of birds and other archosaurs have a well‐ossified neurocranium. In contrast to most of their reptilian relatives, but similar to what we see in mammals, bird brains fit closely to the endocranial cavity so that major external features are reflected in the endocasts. What you see on the inside is what you see on the outside.

This makes birds an excellent group for palaeoneurological investigations. The first observation of the brain in a long‐extinct bird was made in the first quarter of the 19th century. However, it was not until the 2000s and the application of modern imaging technologies that avian palaeoneurology really took off.

Understanding how the mode of life is reflected in the external morphology of the brains of birds is but one of several future directions in which avian palaeoneurological research may extend.

Although the number of fossil specimens suitable for palaeoneurological explorations is considerably smaller in birds than in mammals and will very likely remain so, the coming years will certainly witness a momentous strengthening of this rapidly growing field of research at the overlap between ornithology, palaeontology, evolutionary biology and the neurosciences.

Reference: Cau, Andrea; Brougham, Tom; Naish, Darren (2015). "The phylogenetic affinities of the bizarre Late Cretaceous Romanian theropod Balaur bondoc (Dinosauria, Maniraptora): Dromaeosaurid or flightless bird?". PeerJ. 3: e1032. doi:10.7717/peerj.1032. PMC 4476167. PMID 26157616.

Reference: Ivanov, M., Hrdlickova, S. & Gregorova, R. (2001) The Complete Encyclopedia of Fossils. Rebo Publishers, Netherlands. p. 312