JOGGINS FOSSIL CLIFFS: NOVA SCOTIA

Hylonomus lyelli, Ancestor of all dinosaurs

The fossil cliffs at Joggins are one of Canada's gems, now a UNESCO World Heritage Site, you can visit to see our ancient world frozen in time. 

Preserved in situ is a snapshot of an entire food chain of a terrestrial Pennsylvanian Coal Age wetland.

The outcrop holds fossil plant life — including impressive standing lycopsid trees that formed the framework of these wetlands — decomposing detritivores in the invertebrates and tetrapods, the predatory carnivores of the day.

The Coal Age trees were fossilized where they stood 300-million-years ago with the remains of the earliest reptiles entombed within. The preservation is quite marvellous with the footprints of creatures who once lived in these wetlands are frozen where they once walked and the dens of amphibians are preserved with remnants of their last meal. 

Nowhere is a record of plant, invertebrate and vertebrate life within now fossilized forests rendered more evocatively. The fossil record at Joggins contains 195+ species of plants, invertebrates and vertebrates. The fossil plant life became the vast coal deposits for which this period of Earth's history is named. 

Recorded in the rock are vertebrate and invertebrate fauna both aquatic and terrestrial. This broad mix of specimens gives us a view into life back in the Pennsylvanian and sets us up to understand their ecological context.

Pennsylvanian Coal Age Ecosystem, 300-Million-Years-Old

The fossil record includes species first defined at Joggins, some of which are found nowhere else on Earth. 

It was here that Sir Charles Lyell, with Sir William Dawson, founder of modern geology, discovered tetrapods — amphibians and reptiles — entombed in the upright fossil trees. 

Later work by Dawson would reveal the first true reptile, Hylonomus lyelli, ancestor of all dinosaurs that would rule the Earth 100 million years later. 

This tiny reptile serves as the reference point where animals finally broke free of the water to live on land. This evolutionary milestone recorded at Joggins remains pivotal to understanding the origins of all vertebrate life on land, including our own species. 

Sir Charles Lyell, author of Principles of Geology, first noted the exceptional natural heritage value of the Joggins Fossil Cliffs, calling them “...the finest example in the world of a natural exposure in a continuous section ten miles long, occurs in the sea cliffs bordering a branch of the Bay of Fundy in Nova Scotia.” Indeed, the world-famous Bay of Fundy with its impressive tides, the highest in the world, and stormy nature exposed much of this outcrop. 

Geological accounts of the celebrated coastal section at Joggins first appear in the published literature in 1828–1829, by Americans C.T. Jackson and F. Alger, and by R. Brown and R. Smith, managers for the General Mining Association in the Sydney and Pictou coal fields. Brown and Smith’s account is the first to document the standing fossil trees.

Joggins Fossil Cliffs Map (Click to Enlarge)

Plan Your Joggins Fossil Cliffs Staycation

Joggins Fossil Cliffs is a Canadian gem — and they welcome visitors. They offer hands-on learning and discovery microscope activities in their Fossil Lab.

You can explore interpretive displays in the Joggins Fossil Centre before heading out to the beach and cliffs with an interpreter.

Their guided tours of the fossil site include an educational component that tells you about the geology, ecology, palaeontology and conservation of this very special site. 

Joggins / Chegoggin / Mi'kmaq L'nu

We know this area as Joggins today. In Mi'kmaw, the language spoken in Mi'kma'ki, the territory of the Mi'kmaq L'nu, the area bears another name, Chegoggin, place of fishing weirs.

Booking Your Class Field Trip

If you are a teacher and would like to book a class field trip, contact the Director of Operations via the contact information listed below. They will walk you through Covid safety and discuss how to make your visit educational, memorable and fun.

Know Before You Go

The Bay of Fundy has the highest tides in the world. Beach walks are scheduled according to the tides and run regardless of the weather. Good low tides but raining, the beach walk goes on. Lovely and sunny but with a high tide, the beach walk must wait. So, you will want to dress for it as they will not be cancelled in the event of rain. Should severe weather be a factor, bookings may need to be rescheduled at the discretion of the Joggins staff.

Any questions about booking your school field trip? Feel free to email:  operations@jogginsfossilcliffs.net or call: 1 (902) 251-2727 EXT 222.

References & further reading:

Joggins Fossil Cliffs: https://jogginsfossilcliffs.net/cliffs/history/

Image: Hylonomus lyelli, Una ricostruzione di ilonomo by Matteo De Stefano/MUSEThis file was uploaded by MUSE - Science Museum of Trento in cooperation with Wikimedia Italia., CC BY-SA 3.0, https://commons.wikimedia.org/w/index.php?curid=48143186

Image: Arthropleura: Par Tim Bertelink — Travail personnel, CC BY-SA 4.0, https://commons.wikimedia.org/w/index.php?curid=48915156

Joggins Map: Joggins Fossil Cliffs: https://jogginsfossilcliffs.net/cliffs/history/

KWAGULTH GUDZI UMAGALIS

The door to the Long House in Tsaxis, Fort Rupert. This was the Opening Ceremony for the Long House & Hunt Family Potlatch in 1995 hosted by Ross Hunt Sr. 

I went to the potlatch with my Uncle Doug Henderson and Grandmother Betty Henderson (nee Hunt). Both danced her Chilkat blanket. It was at this potlatch that Calvin Hunt received his Chief's name, Tlasutiwalis, from his wife's side of the family.

Look at this beautiful carving and wonderful use of colour. The building is truly something to be proud of. 

There have been upgrades to the building since then. To those that have contributed, I say a warm thank you. Gilakas'la. You know who you are. You know the love & skill you have contributed to the community. 

Tsaxis ''Kwagulth Gudzi Umagalis'' Fort Rupert, BC, British Columbia. Kwakwaka'wakw Big House '' House of the Chiefs'' Kanada, Turtle Island. 

Master carver Tony Hunt Sr headed up the Gudzi Community Big House known as gigukwdzi in Kwak'wala. 

The late Henry Hunt Sr shared his skill and oversaw the carving of the three Sisiutl Heads and Frogs on Tongues. Tony Hunt Jr, Tommy Hunt Jr., Steven Hunt, George Jr Hunt carved the Double-Headed Serpent and Frogs on Tongues with Uncle Henry Hunt Sr.

The Kwakiutl Chiefs were discussing the creation of their ancestors while waiting for the second course at a feast given by one of the chiefs at Tsaxis. At first, no one spoke for a while. Then Malid spoke, saying, “It is the Sun, our chief, who created our ancestors of all the tribes. And when the others asked him how this was possible, for the Sun never made even one man, the chief was silent. Others said, It is Mink, Tlisalagi’lakw, who make our first ancestors. 

Then spoke Great-Inviter, saying, “Listen Kwakiutl, and let me speak a really true word.  I see it altogether mistaken what the others say, for it was the Seagull who first became man by taking off his mask and turning into a man. 

This was the beginning of one of the groups of our tribe. And the others were caused when the Sun, and Grizzly Bear, and Thunderbird also took off their masks. That is the reason that we Kwakiutl are many groups, for each group had its own original ancestor.”

A chief visiting from Nawitti disagreed, and the Kwakiutl of all four groups became angry. The Nawitti believe that the Transformer (or Creator) went about creating the first ancestors of all the tribes from people who already existed. 

But the chiefs of the Kwakiutl scoffed at this, saying, “Do not say that the Transformer was the creator of all the tribes. Indeed, he just came and did mischief to men, when he made him into a raccoon, and land otter, and deer, for he only transformed them into animals. We of the Kwakiutl know that our ancestors were the Seagull, Sun, Grizzly Bear, and Thunderbird.

OLENELLUS OF THE EAGER FORMATION

The partial specimen you see here is an Olenellus trilobite from the Eager Formation near Cranbrook, British Columbia.

It was exciting to crack open the rock and find a specimen, many specimens, more than half a billion years old. It is something we so rarely do but the opportunity is all around us in the many sedimentary rocks that outcrop near the surface around the globe. 

Near Cranbrook, the Eager Formation outcrops at several locations just outside of town. This particular lovely is from the Rifle Range locality and is in my collection at 98-CR-EF042 — meaning it was collected in 1998 and the 42nd find of the day. This is a prolific site and with diligent collecting, you can find many wonderful specimens of scientific and aesthetic value in the course of a day.

The Rifle Range locality sits on the Silhouette Rifle Range — which is literally on a rifle range where folks go to shoot at things.

The fossils we find here are just a shade older than those found at the Burgess Shale. Burgess is Middle Cambrian and the species match the Eager fauna one for one but the Eager fauna are much less varied. 

Olenellus is an extinct genus of redlichiid trilobites, early arthropods, that litter this glorious Cambrian site. Olenellus is the only genus currently recognised in the subfamily Olenellinae. The sister group called the Mesonacinae consists of the genera Mesonacis and Mesolenellus.

Olenellus range in size but are about 5 centimetres or 2.0 inches long on average. They lived during the Botomian and Toyonian stages, Olenellus-zone, 522 to 510 million years ago, in what is currently North America in what was part of the paleocontinent of Laurentia.

Olenellus are both common in and restricted to Early Cambrian rocks — 542 million to 521 million years old — and thus a useful Index Fossil for the Early Cambrian. 

Olenellus had a well-developed head, large and crescentic eyes, and a poorly developed, small tail. The fellow you see had a bit of his tail crushed as he turned to stone.

Trilobites were amongst the earliest fossils with hard skeletons. While they are extinct today, they were the dominant life form at the beginning of the Cambrian and it is what we find as the primary fossil in the fauna of the Eager Formation. 

A slightly crushed lingulida brachiopod

The Eager Formation has produced many beautifully preserved Wanneria, abundant Olellenus and a handful of rare and treasured Tuzoia and Anomalocaris claws. The shale matrix lends itself to amazing preservation. 

The specimens of Wanneria from here are impressively large. Some are up to thirteen centimetres long and ten centimetres wide. You find a mixture of complete specimens and head impressions from years of perfectly preserved moults.

From July 21 to 31, 2015, the Royal Ontario Museum (ROM), under the direction of Dr. Jean-Bernard Caron carried out a palaeontological dig at an exposure of the Eager Formation that outcrops between Cranbrook and  Fort Steele in the East Kootenay Region of British Columbia. 

The team included David George (APS), Dr. Bob Gaines (Pomona College), Dr. Jean-Bernard Caron (ROM), Dr. Gabriela Mangano (University of Saskatchewan), Maryam Akrami (ROM), Darrell Nordby (APS), Joe Moysiuk (University of Toronto), and local, Guy Santucci (APS and project field co-ordinator), and Dr. Mark Webster (University of Chicago).

Dr. Caron was interested in the fauna of the Eager Formation as there is an overlap with the Burgess Shale species — the Eager is a window into time 513 million years ago — 8 million years earlier than the Burgess. 

Lower Cambrian Brachiopod

They found the usual suspects, including multiple specimens of Wanneria dunnae and Olenellus ricei along with the rarer genus Mesonacis, in addition to specimens of the elusive Tuzoia. 

They also found a block with at least 112 individual trilobites (mostly moulted cephalons) of Olenellus ricei and Wanneria dunnae. 

The most exciting of their finds included a sponge, Anomalocaris, Morania (a cyanobacterial growth), and a hyolithid similar to the Burgess Haplophrentis.

They also found many trace fossils. There was a particularly fetching 30 cm trace fossil, likely from a large Wanneria, that I hope Dr. Mángano or one of her graduate students lend their gaze — Gabriela is a particularly good writer. 

She is co-editor of Palaios, in addition to being a member of the editorial board of a number of journals, including Journal of Paleontology, Paleontologia Electronica, Palaeogeography Palaeoclimatology Palaeoecology, Ameghiniana, and Revista Brasilera de Paleontologia. Gabriela is a member of the Scientific Board of the UNESCO International Geoscience Program (IGCP), member of the SEPM Board, and Treasurer of the International Ichnological Association. Add all that to extensive fieldwork and supervising over fifteen graduate students and postdoctoral fellows — she's an amazing woman.

Their excavation of the site was thorough — reducing all of the potentially fossil-bearing strata to pieces the size or smaller than a dinner plate. The 2015 team used a backhoe to take off the weathered top layer and get down to the bedrock below.

It has been six years since their visit and we will hopefully see some worthy publications from their efforts. There had been talk of multiple publications stemming from the spectrum of species, a comparison to the Burgess fauna and papers on the trace fossils. I checked in with Joe Moysuik from the University of Toronto who had been on the dig in 2015. To his knowledge, no new papers have yet to be published — though, Caron has been a busy bee on a sexy new nektobenthic suspension feeder from the Burgess material. I am rather hoping their team sorts out the naming of some of the species and gets them to publication so we can finally put them to bed.

Days after my correspondence with Moysuik, Chris Jenkins, a Cranbrook local and huge contributor to our knowledge of Upper Cambrian trilobites, shared an exciting find. 

He and Don Askew had ventured out together for their first fossil field trip of 2021 — and made a rather auspicious start to the year. 

The two had met some 10 years previous when Don, an avid outdoorsman and Jenkins' neighbour, had wandered over to see what all the rocks were about in Jenkins' yard. 

Tales of trilobites and a lifelong friendship ensued. It was also the beginnings of shared fieldwork. This time, it was to outcrops of the Eager Formation just outside of Cranbrook. Together, the two unearthed a three-foot-wide band of Eager Formation bedrock. Not unusual in and of itself — but instead of the usual trilobites — this rock revealed several varieties of Lower Cambrian brachiopods. 

Jenkins shared photos of at least three different types of brachiopods — potentially new fauna for the Eager. Although they superficially resemble the molluscs that make modern seashells, they are not related. Brachiopods were the most abundant and diverse fossil invertebrates of the Paleozoic — over 4500 genera known; the number of species is far greater. So, naturally, we had expected to find brachiopods in the Eager Formation as they were abundant in Lower Cambrian seas — but so far they had eluded us.

And, interestingly, the rock containing the brachiopods is devoid of any trilobite specimens — not a one. Have they found a wee slip of the Eager Formation that records a nearshore environment or have they stumbled across a segment that records another time period altogether?

The brachiopods you see in the photos above are roughly 1/4 inch to 3/4 inches. Should Caron and team return to the site, these new brachiopods will be of great interest as they look to rewrite the geology and palaeontology of the site and region. 

FRANZ BOAS WITH THE HUNT FAMILY 1894

This photograph was taken in 1894 of Franz Boas with the Hunt family.

Back row, left to right: Sam Hunt, George Hunt, Mary Ebbets Hunt (Anislaga/Anéin)

(George’s mother), Franz Boas; standing left to right: Lalaxs’a (wife of David Hunt, not pictured), Jonathan Hunt; seated left to right:

Emily Hunt (holding Marion Hunt), Lucy Hunt; kneeling left to right:

Mary Hunt, George Hunt Jr. Photographed by Oregon C. Hastings.

Courtesy of the American Philosophical Society, APS 466.

FOSSIL HUNTRESS — PALAEO SOMMELIÈR

Geeky Palaeo Goodness from the Fossil Huntress — Palaeo Sommelièr — musings in natural history meant to captivate, educate & inspire! Listen to the Fossil Huntress Podcast on Spotify, Apple iTunes & Anchor — Dead Sexy Science for your ears!

Why Palaeo Sommelièr? 

For many years, I used the title Explorer-in-Residence until one day, I noticed that many National Geographic colleagues had adopted it. So, Palaeo Sommelièr was born.

A sommelier (/ˈsɒməljeɪ/ or /sʌməlˈjeɪ/ or /sɒˈmɛlijeɪ/; French pronunciation: ​[sɔməlje]), or wine steward, is a trained and knowledgeable wine professional, normally working in fine restaurants, who specializes in all aspects of wine service as well as wine and food pairing. 

A sommelièr is a fossil steward — a trained and knowledgeable fossil professional, normally working in science communication with a specialization in palaeontology. As I live in British Columbia, and by law, everyone here who finds a fossil is a steward of the fossil, it is the perfect pairing. 

CARNOTAURUS: FLESH-EATING BULL

Carnotaurus sastrei, a genus of large theropod dinosaurs that roamed the southern tip of Argentina, South America during the Late Cretaceous, 72 to 69.9 million years ago. His name means "flesh-eating bull,' and he lives up to it.

This fellow — or at least his robust skull with the short, knobby eyebrow horns and fierce-looking teeth — is on display at the Natural History Museum in Madrid, Spain. For now, he is the only known genus of this species of bipedal predator.

The first specimen of Carnotaurus sastrei was found in Chubut on vast plains between the Andes Mountains and the Atlantic Ocean. A physician, Dr. A'ngel Tailor noticed a large concretion showing some bone fragments. A team led by José F. Bonaparte excavated the find in 1984 as part of a paleontological expedition funded by the Argentine Museum of Natural Sciences.

Sadly, Bonaparte — the Maestro del Mesozoico — passed away the 18th February 20220 at the age of 91. He spent the majority of his career as head of the Vertebrate Palaeontology Division of the Museo Argentino de Ciencias Naturales “Bernardino Rivadavia,” in Buenos Aires. Bonaparte opened up the vertebrate finds of Argentina to the world. He was instrumental in the finding, excavating and naming many iconic dinosaurs — Carnotaurus, Amargasaurus, Abelisaurus, Argentinosaurus, Noasaurus along with the finding of the first fossilised remains of Mesozoic South American mammals. He mentored many palaeontologists who will miss his keen eye and tremendous work ethic — Luis Chiappe, Rodolfo Coria, Agustín Martinelli, Fernando Novas, Jaime Powell, Guillermo Rougier, Leonardo Salgado, Sebastián Apesteguía and many others.

His excavation of Carnotaurus was the first of its kind and he recognized that the skull is quite unusual. Initially, it has a very marine reptile feel — but make no mistake this guy is clearly a terrestrial theropod. He had smallish, underdeveloped arms — teeny by theropod standards. Once you look closer you see his bull-like horns from whence he gets his name — horns that imply battle between rivals for the best meal, sexual partner and to be the one who leads the herd. 

He was covered in leathery skin lined with rows of cone-shaped nodules or bumps. These get larger as they move towards his spine. He had forward-facing eyes, similar to tyrannosaurs like T-rex and smaller theropods like Velociraptor and Troodon — who had better vision even that T-rex — which would have given him the advantage of binocular vision and depth perception. Forward-facing eyes are also quite helpful with nocturnal hunting — think owls and cats — as they take in more light and help with nighttime predation. So perhaps this flesh-eating bull fancied a late-night snack on his menu from time to time.

Species like squirrels, pigeons and crocodiles have eyes on the sides of their heads. They lack the important competitive feature of well-developed depth perception — being able to easily and estimate distance — but perhaps make up for it with a panorama that offers a wider field of view.   

BLACK-SPOTTED YELLOW SEAHORSE

The glorious black-spotted lemon yellow lovely you see here is a seahorse. They are one of the fairytale creatures that I was pleased to see truly exist. These marine lovelies are any of 46 species of small marine fish in the genus Hippocampus. 

Hippocampus comes from the Ancient Greek hippókampos (ἱππόκαμπος) — a cobbling together of híppos (ἵππος) meaning horse and kámpos (κάμπος) meaning sea monster.  

A delightful albeit tad sinister name for these charming wee sea monster horses from our world's shallow tropical and temperate seas. 

Having a head and neck suggestive of a horse, seahorses also feature segmented bony armour, wee tiny, spiny plates from tip to tail. They bob about in seagrasses, gripping with their curled prehensile tails when they want to stay in one place or using their dorsal or back fin to help them move up and down to swim about. Along with the pipefishes and seadragons — Phycodurus and Phyllopteryx — they form the fused jaw family Syngnathidae. The fellow you see here looks like he's nestled in a holiday display. He's not, of course, but it is not such a hard thing to image considering I once believed them to be fictional.

CTENOPHORES: COMB JELLIES

Cannibalistic Comb Jellies

This festive lantern looking lovely belongs to a group of invertebrates known as comb jellies.

Comb jellies are named for their unique plates of giant fused cilia, or combs, which run in eight rows up and down the length of their bodies. They are armed with sticky cells or colloblasts, that do not sting but display wonderful bioluminescent colouring as they move through the sea.

Ctenophores or comb jellies are one of the phylogenetically most important and controversial metazoan groups. They are not jellyfish and are not closely related, though they do share some characteristics with the gelatinous members of the subphylum Medusozoa. 

Comb jellies are not picky eaters. Their tastes range to what is at hand, including cannibalizing other comb jellies. They will feast on their kin along with tasty plankton, zooplankton, crustaceans and wee fish.

Interest in their fossil record has been catalysed by spectacularly preserved soft-bodied specimens from Cambrian Lagerstätten of the 518-million-years-old Chengjiang Biota, the 505-million-years-old Burgess Shale and other Burgess Shale-like deposits. 

We find them in the Late Devonian Escuminac Formation at Miguasha National Park, Quebec, Canada — a UNESCO world heritage site famous for its abundance of well-preserved vertebrate fossils including most major evolutionary groups of Devonian lower vertebrates from jawless fish to stem-tetrapods.

Based on morphological similarities of this Canadian fossil with stem-ctenophore fossils from the Cambrian Lagerstätte of the Chinese locality Chengjiang, they have been assessed for their affinity to stem-group ctenophores (dinomischids, Siphusauctum, scleroctenophorans) and early crown-group ctenophores. Modern ctenophores and many fossil forms lack mineralized hard parts, which renders the rare fossils that have been extracted from several Lagerstätten quite remarkable. 

Like the soft bodies of jellyfish and the polyps of hydrozoans and anthozoans, the probability for such soft bodies (or body regions) to become fossilized is extremely low. In spite of this low preservation potential, remains of stem-ctenophores have become known from several Cambrian and younger conservation deposits, and with even older candidate ctenophores in the Ediacaran. 

While Cambrian Lagerstätten have yielded several genera, ctenophore remains are much rarer in the Devonian; in particular, two studies, describing material from the German Hunsrück Slate. 

Bioluminescent Comb Jellies

This Early Devonian material, however, appears to belong to crown ctenophores morphologically similar to living forms such as Pleurobrachia, unlike the stem Cambrian taxa and the new Devonian stem taxon described here.

The most basal stem ctenophores are the dinomischids: sessile benthic petaloid invertebrates, many of which are equipped with a stalk. This group first was described from the Middle Cambrian Burgess Shale. Based on the genus Dinomischus, these early stalked forms were commonly called ‘dinomischids’. 

Zhao et al. shared that dinomischids "form a grade in the lower part of the ctenophore stem group” and include taxa such as Xianguangia, Daihua, and Dinomischus that have hexaradiate-based symmetry (e.g., sixfold, 18-fold). 

Some later, skeletonised stem-ctenophores were termed ‘Scleroctenophora’; ‘scleroctenophorans’ have a shorter stalk, lack the ‘petals’ and have no bracts and might be monophyletic. 

To date, all known dinomischids and scleroctenophorans are Cambrian. Remarkably, analysis of the material described here suggests it is a very late-surviving member of this part of the ctenophore tree, occurring in strata over a hundred million years younger with no intervening known record, thus making it a Lazarus taxon with an extensive ghost lineage. 

Palaeozoic sediments yield a growing number of fossil invertebrates with radial symmetries, some being quite enigmatic with body plans differing radically from those of extant organisms.

The morphological similarities to Cambrian forms and the mix of characters regarding overall shape and symmetries render this discovery important. The aims of this study are to describe the only known specimen of this Devonian ctenophore, discuss its phylogenetic and systematic position, and the impact of fossil data for ctenophore affinities, and assess its palaeoecological role.

FRAGILE BEAUTY: FOSSILIZED SCLERACTINIAN CORAL

Scleractinian Fossil Coral, Florida

The delicate wintery beauty you see here is a Scleractinian coral we find first in the fossil record in the Mesozoic. 

Corals first appeared in the Cambrian about 535 million years ago. Fossils are extremely rare until the Ordovician period, 100 million years later, when rugose and tabulate corals became widespread. 

Palaeozoic corals seem to make friends wherever they live and often contain numerous endobiotic symbionts.

Tabulate corals occur in limestones and calcareous shales of the Ordovician and Silurian periods, and often form low cushions or branching masses of calcite alongside rugose corals. 

Their numbers began to decline during the middle of the Silurian period, and they became extinct at the end of the Permian period, 250 million years ago.

Rugose or horn corals became dominant by the middle of the Silurian and became extinct early in the Triassic period. The rugose corals existed in solitary and colonial forms and were also composed of calcite.

The famous Great Barrier Reef is thought to have been laid down about two million years ago. If you have had the pleasure of scuba diving near it to take in its modern wonders, perhaps you will be interested to learn how it was formed. Over long expanses of time, the corals here have broken up, fragmented and died. Sand and rubble accumulate between the corals, and the shells of clams and other molluscs decay to form a gradually evolving calcium carbonate structure to what you view today. 

Coral reefs are extremely diverse marine ecosystems hosting over 4,000 species of fish, massive numbers of cnidarians, molluscs, crustaceans, and many other animals.

JURASSIC SEA URCHIN: AM'DA'MA

This lovely little biscuit is a Holectypus sea urchin from 120 million-year-old deposits from the Lagniro Formation of Madagascar.

The specimen you see here is in the collections of my beautiful friend Ileana. She and I were blessed to meet in China many years ago and formed an unbreakable bond that happens so few times in one's life. 

Holectypus are a genus of extinct echinoids related to modern sea urchins and sand dollars. They were abundant from the Jurassic to the Cretaceous (between 200 million and 65.5 million years ago).

This specimen is typical of Holectypus with his delicate five-star pattern adorning a slightly rounded test and flattened bottom. The specimen has been polished and was harvested both for its scientific and aesthetic value. 

I have many wonderful memories of collecting their modern cousins that live on the north end of Vancouver Island and along the beaches of Balaklava Island. In the Kwak̓wala language of the Kwakiutl or Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, sea urchins are known as a̱m'da̱'ma and it is this name that I hear in my head when I think of them.

In echinoids, the skeleton is almost always made up of tightly interlocking plates that form a rigid structure or test — in contrast with the more flexible skeletal arrangements of starfish, brittle stars, and sea cucumbers. Test shapes range from nearly globular, as in some sea urchins, to highly flattened, as in sand dollars. 

Sea Urchin Detail

Living echinoids are covered with spines, which are movable and anchored in sockets in the test. These spines may be long and prominent, as in typical sea urchins and most have lovely raised patterns on their surface. 

In sand dollars and heart urchins, however, the spines are very short and form an almost felt-like covering. The mouth of most echinoids is provided with five hard teeth arranged in a circle, forming an apparatus known as Aristotle’s lantern.

Echinoids are classified by the symmetry of the test, the number and arrangement of plate rows making up the test, and the number and arrangement of respiratory pore rows called petals. Echinoids are divided into two subgroups: regular echinoids, with nearly perfect pentameral (five-part) symmetry; and irregular echinoids with altered symmetry.

Because most echinoids have rigid tests, their ability to fossilize is greater than that of more delicate echinoderms such as starfish, and they are common fossils in many deposits. The oldest echinoids belong to an extinct regular taxon called the Echinocystitoidea. 

They first appeared in the fossil record in the Late Ordovician. Cidaroids or pencil urchins appear in the Mississippian (Early Carboniferous) and were the only echinoids to survive the mass extinction at the Permo-Triassic boundary. Echinoids did not become particularly diverse until well after the Permo-Triassic mass extinction event, evolving the diverse forms we find them in today. 

True sea urchins first appear in the Late Triassic, cassiduloids in the Jurassic, and spatangoids or heart urchins in the Cretaceous. Sand dollars, a common and diverse group today, do not make an appearance in the fossil record until the Paleocene. They remain one of my favourite echinoderms and stand tall amongst the most pleasing of the invertebrates.

LEMURS OF MADAGASCAR

These lovelies are lemurs. They are very social and like to live in groups of half a dozen to up to 30 individuals. A female leads the group, or troop, and will likely have mated with the stinkiest — think sexiest — male. 

Once they breed, mamma will give birth to one to a half dozen pups after a gestation period of about 100 to 107 days. The wee pups cling to mamma's belly for the first few weeks of life then crawl up to live on her back for the next few months. By three to six months, the wee ones are weaned and a year to three years later, this pup will be mature and ready to mate. If all goes well, some species can live up to 30 years — rather a long life in the wild.

Lemurs are mammals of the order Primates, divided into 8 families and consisting of 15 genera and around 100 highly diverse species — 105 to be exact. They are native only to the island of Madagascar.

Most lemurs are relatively small, have a pointed snout, large eyes, and a long tail. They are arboreal, living primarily in trees and nocturnal, preferring to be active at night, snacking on leaves, shoots, fruit, flowers, tree bark, nectar and sap.

Phylogenetic, genetic, and anatomical evidence all suggest that lemurs split from other primates on Africa around 62 million years ago and that the ancestral lemur lineage had dispersed to Madagascar by around 54 million years ago. They must have come over to the island clinging to floating vegetation.

Once on the island, the lemur lineage diversified. Now there are at least 105 species of lemur, all endemic to Madagascar. They range in size from just an ounce and just 9 to 11 cm in the case of the Madame Berthe's mouse lemurs to up to 15 to 22 lbs or 7 to 10 kilograms, in the case of the Indri.

The evolutionary and biogeographic processes experienced by the lemurs are not unusual. Madagascar is home to many groups of endemic organisms with close within-group relationships. The simplest — or most parsimonious — explanation for this pattern is that, like the lemurs, the groups first arrived on the island by dispersal as a single lineage and then rapidly diversified. This diversification was likely spurred on by other geologic and geographic characteristics of Madagascar.

The east coast of the island is lined with a mountain range — and this causes different parts of the island to get drastically different amounts of rain. Hence, the island is made of many different habitat types — from deserts to rainforests — that have shifted and changed over the past 88 million years. This likely provided many opportunities for subpopulations to become isolated and evolve traits for specializing in different niches. And that likely encouraged lineages to diversify.

Today, Madagascar is one of the most diverse places on Earth. Understanding where that diversity comes from requires understanding, not just the living world, but the geologic, geographic, and climactic histories that have shaped the evolution of lineages on the island. Now, human history in the making threatens to undo tens of millions of years of evolution in just a few years of political turmoil — unless safeguards can be put in place to protect Madagascar's unique biota from the instabilities of human institutions.

Cooper, A., Lalueza-Fox, C., Anderson, S., Rambaut, A., Austin, J., and Ward, R. (2001). Complete mitochondrial genome sequences of two extinct moas clarify ratite evolution. Nature 409:704-707.

Goodman, S. M., and Benstead, J. P. (2005). Updated estimates of biotic diversity and endemism for Madagascar. Oryx 39(1):73-77.

Evolution Berkeley: https://evolution.berkeley.edu/evolibrary/news/091001_madagascar

Vences, M., Wollenberg, K. C., Vieites, D. R., and Lees, D. C. (2009). Madagascar as a model region of species diversification. Trends in Ecology and Evolution 24(8):456-465.

HO HO, OH! IS THAT A DINOSAUR TRACK?

This is a theropod track left by an Allosaurus walking through a muddy bank that turned to stone. These big beasties were large carnosaurian theropod dinosaurs that lived 155 to 145 million years ago.

This track and many others are at Dinosaur Ridge, a segment of the Dakota Hogback in the Morrison Fossil Area National Natural Landmark located in Jefferson County, Colorado, near the town of Morrison and just west of Denver.

The Dinosaur Ridge area is one of the world's most famous dinosaur fossil localities. Along with the dinosaur bones, you find plant fossils, crocodile bones and a variety of reptile tracks. 

In 1877, fossil excavation began at Dinosaur Ridge under the direction of palaeontologist Othniel Charles Marsh. Some of the best-known dinosaurs were found here, including Stegosaurus, Apatosaurus, Diplodocus, and Allosaurus. 

In 1973, the area was recognized for its uniqueness as well as its historical and scientific significance when it was designated the Morrison Fossil Area National Natural Landmark by the National Park Service. In 1989, the Friends of Dinosaur Ridge was formed to address increasing concerns regarding the preservation of the site and to offer educational programs on the area's resources.

Visit Dinosaur Ridge Morrison Fossil Area National Natural Landmark

Fancy a trip? You can visit these wonderful tracks, the Dinosaur Ridge Exhibit Hall and walk the trails through the tracks. They have put up helpful interpretive signs that explain the local geology, a volcanic ash bed, trace fossils, paleo-ecology, economic development of coal, oil and clay, and many other geologic and paleontological features.

There are two trails and a visitor centre at Dinosaur Ridge. The visitor centre features information on trails and a small gift shop. If you do not pack a lunch, you'll want to visit the Stegosaurus Snack Shack located outside the visitor centre offering coffee, cold drinks, burritos, pretzels and more. It is open from 10am-4pm Mon-Sat and 11am-4pm on Sun from June through August. 

Check updates on their website concerning Covid-19 restrictions. This is a vast site with easy trails so it makes for a great family trip when museums and other indoor facilities are closed.

Know Before You Go

Uncover Colorado: https://www.uncovercolorado.com/landmarks/morrison-golden-fossil-areas-dinosaur-ridge/

AWKWARD AND AWESOME: DIMORPHODON

This remarkable fellow is Dimorphodon — a genus of medium-sized pterosaur from the Early Jurassic. He is another favourite of mine for his charming awkwardness.

You can see this fellow's interesting teeth within his big, bulky skull. Dimorphodon had two distinct types of teeth in their jaws — an oddity amongst reptiles — and also proportionally short wings for their overall size. 

Just look at him. What an amazing beast. We understand their anatomy quite well today, but can you imagine being the first to study their fossils and try to make sense of them. 

The first fossil remains now attributed to Dimorphodon were found in England by fossil collector Mary Anning, at Lyme Regis in Dorset, the United Kingdom in December 1828. While she faced many challenges in her life, she was blessed to live in one of the richest areas in Britain for finding fossils. 

She walked the beaches way back in the early 1800s of what would become the Jurassic Coast UNESCO World Heritage Site. The Jurassic Coast holds some of the most interesting fossils ever found — particularly within the strata of the Blue Lias which date back to the Hettangian-Sinemurian. It is one of the world’s most famous fossil sites. Millions come to explore the eroding coastline looking for treasures that provide delight and inspiration to young and old.

These fossil treasures provide us with tremendous insights into our world 185 million years ago when amazing animals like Dimorphodon ruled the skies. 

Mary's specimen was acquired by William Buckland and reported in a meeting of the Geological Society on 5 February 1829. Six years later, in 1835, William Clift and William John Broderip built upon the work by Buckland to publish in the Transactions of the Geological Society, describing and naming the fossil as a new species. 

As was the case with most early pterosaur finds, Buckland classified the remains in the genus Pterodactylus, coining the new species Pterodactylus macronyx. The specific name is derived from Greek makros, "large" and onyx, "claw", in reference to the large claws of the hand. The specimen, presently NHMUK PV R 1034, consisted of a partial and disarticulated skeleton on a slab — notably lacking the skull. Buckland in 1835 also assigned a piece of the jaw from the collection of Elizabeth Philpot to P. macronyx. 

Later, the many putative species assigned to Pterodactylus had become so anatomically diverse that they began to be broken into separate genera.

In 1858, Richard Owen reported finding two new specimens, NHMUK PV OR 41212 and NHMUK PV R 1035, again partial skeletons but this time including the skulls. Having found the skull to be very different from that of Pterodactylus, Owen assigned Pterodactylus macronyx its own genus, which he named Dimorphodon. 

His first report contained no description and the name remained a nomen nudum. In 1859, however, a subsequent publication by Owen provided a description. After several studies highlighting aspects of Dimorphodon's anatomy, Owen finally made NHMUK PV R 1034 the holotype in 1874  — 185 million years after cruising our skies the Dimorphodon had finally fully arrived.

GOD JUL: TRILACINOCERAS NORVEGICUM

Trilacinoceras norvegicum

A lovely example of Trilacinoceras norvegicum (Sweet, 1958), a nektonic carnivorous cephalopod from Ordovician outcrops on Helgö Island, Hovindsholm, Helgøya, Lake Mjosa, Norway.

This has been a site of human habitation for more than 5,000 years. Vikings, kings, traders, farmers —  and geologists have walked these fields.

To give that timeframe a bit of context, that's about the age of Skara Brae, the Neolithic settlement in Orkney, Scotland — and older than Stonehenge which clocks in at 3000 BC to 2000 BC and the Great Pyramids — built around 2560 BC.

For my friend, Gale Bishop, that's about 469 km west or a good 7-hour drive from your ancestral home in Ask, just north of Bergen and just south of Knarvik where many of my relatives live — Hei du!

The fossils found here are part of the Engervik Member, Elnes Formation, Aseri, and date back to the Middle Ordovician, 463.5 - 460.9 million years ago. W. C. Sweet did fossil fieldwork here in the 1950s and published a paper on the Middle Ordovician of the Oslo Region, Norway 10. Nautiloid Cephalopods. Norsk Geologisk Tidsskrift 38:1-178.

Deservedly, Sweetoceras boreale is named for him and is one of the most delightful species names of all time. In the 1960s, Yochelson picked up where Sweet left off, continuing the survey of the Middle Ordovician of the Oslo region. I chose this Trilacinoceras for a holiday post because their curly tops remind me of a wee Norwegian gnome, or Nisse from the Norse niðsi, a dear little relative. My Swedish relatives call them Tomte, a throwback to Saint Birgitta of Sweden in the 1300s.

Helgøya is an island in Mjøsa located in the Ringsaker municipality of Hedmark county, Norway. It was formerly a part of the Nes municipality. 

Long before that, it was the ruling centre for the Kings in Hedmark, where bold men and women held great blót celebrations to Odin and planned raids and expansion into Europe and Russia — roughly A.D. 793 — the beginning of the Viking Age.

Today, it is lush and green and easy to explore — or fish. Mjøsa is Norway's largest lake, as well as one of the deepest lakes in Norway and in Europe. 

Battles have been fought on its waters and its depths hold interesting archaeological and paleontological secrets. They also hold a goodly amount of large and tasty trout, pike, perch, burbot and graylings.

Helgøya is the largest freshwater island in Norway at 18.3 km². The island is delightful to explore and home to 32 farms. One of the most beautiful of these is the Hovinsholm manor. You can visit the farm in both summer and winter — both equally beautiful — and enjoy a café, workshop or their Christmas market. They have lush gardens and some very friendly horses you can pet — or spoil with apples, as you do. The property is massive at 2012 acres, divided into grain, potatoes and forest. It has been home to kings and court. It was a monastery in the Middle Ages from the 5th to the 15th century. Today, Tolle Hoel Slotnæs and his wife, Charlotte Holberg Sveinsen own and run the manor with their three daughters.

Hovinsholm, Helgøya, Lake Mjosa, Norway

Helgøya means holy island, in Norwegian. There is a lovely double meaning here and such layered history. The manor, in its various iterations, has been on this site since the 1500s. They had their own Christian manor church until 1612.

On the southern tip of the island, there is an old pagan temple to the Norse Gods, Thor, Frigg, Loki, Hod, Heimdall, Tyr, and Baldur.

Here, farmers of the area would gather at four blót sacrifices a year that followed the seasons — one for each of the winter solstice, spring equinox, summer solstice and autumn equinox. Animals would be sacrificed, their blood splattered on altars, walls and folk around them. Toasts were made. The first was in honour of Thor or Odin, “to the king and victory.” 

Odin, although nominally chief of the gods, was more the god of aristocrats. If a king were toasting, particularly a Danish King, it would be for Odin. If you look at place names in Scandinavia, you'll see him conspicuously absent in favour of Thor, the god of the common man.

When the farmers at Helgøya were shouting Skål, it was likely for Thor. The toasting and drinking continued with cups emptied for Njörd and Freyr and Freyja in the hope of securing a prosperous future. 

Finally, personal pledges (and beer-soaked boasts) would be made to undertake great exploits, Valknut — to die well in battle — and finally to kinsmen laid to rest now drinking with the gods in Valhalla. Weapons, jewellery and tools were thrown into the lake as offerings.

If they were gathering for Jol (Old Norse), Jul (Norwegian) or the Yule blót, they'd also make a large sun wheel (picture a circle with a cross in the middle), carve it up with runes, set it on fire and roll it down a hill. 

It was quite a celebration with the festivities going on for three days and nights. With the formalities over, people did as people do  — drink, sing, boast, play games and find someone to bed down with — Gods be good.

Thor and Odin are still going strong nearly 1,000 years after the end of the Viking Age. You'd think that the old Nordic religion — the belief in the Norse gods — disappeared with the introduction of Christianity. That is not the case. There are still folk in Denmark (Odin-lovers) and Norway (Thor's their guy) who follow the old Norse religion and worship its ancient gods — right down to the splatter.

If you visit Norway at Christmas, Jul (Yule), you'll find much more of the pagan than the Christian in the festivities. King Haakon, old Haakon the Good, Hákon Góði or Håkon den Gode,  moved the Winter Solstice or Yule, Jul, Jol blót over to match up with the Christian holiday on December 25th in his attempts to introduce Christianity in the 10th century. Both traditions are still celebrated but without an overtly religious tone.

Old traditions run deep, animals are still sacrificed (but without all the splatter), bread is baked, houses cleaned, beer is abundant and fires warmth the hearth.

After all the drinking, toasting and feasting at the Jul blót, leftover food was not cleaned up but left overnight for the little relatives. Though shy, Nisse like a good feast and failing to offer them their tithe brings ill-fortune.

But we started this journey together admiring a lovely (and oddly festive) Ordovician cephalopod. Go on, picture him in red and white with a little beard. If you fancy a visit to the Ordovician outcrops, you can find them at Nes-Hamar, Norway. 60.0° N, 11.2° E: paleo-coordinates 33.7° S, 10.3° W. Look for gastropods (five known species) and cephalopods (at least 15 species).

If you'd like to visit the burial mound of Haakon the Good, you'll want to head to Seim, Hordaland, about 10 km north of Knarvik. Good 'ol Haakon may have tried to bring Christianity to Norway but he died full Viking — taking an arrow at the Battle of Fitjar. Many of my rellies live in Knarvik. 

We have enjoyed many a sunny afternoon feasting at the Håkonarspelet summer festivals and exploring Haakon's burial mound at Håkonhaugen in Seim.

If you're more of the manor type, you can stop by Hovinsholm gård, Helgøyvegen 850, 2350 Nes på Hedmarken, Norway. 

If you're curious and want to see the farmstead, head on over to: https://www.skafferiet.no/about. 

If you need to square things up with Odin, you're on your own.

E. L. Yochelson. 1963. The Middle Ordovician of the Oslo Region, Norway. 15. Monoplacophora and Gastropoda. Norsk Geologisk Tidsskrift 43 (2):133-213.

ICE, SNOW, REINDEER AND ICHTHYOSAURS: SVALBARD

Reindeer, Rangifer tarandus 

Ice, Snow, Reindeer & Ichthyosaurs — Svalbard is just what I imagine my version of Valhalla to be like, without all the mead, murder and mayhem. 

This Norwegian archipelago sits between mainland Norway and the North Pole. 

One of the world’s northernmost inhabited areas, it is known for its rugged, remote terrain of glaciers and frozen tundra sheltering polar bears, reindeer and Arctic fox. 

It is also known for reindeer. The lovelies you see here are all females as the males lose their antlers in the winter. So Rudolf and the rest of Santa's crew who pull his sleigh for him would have all been females as they are pictured with antlers. They are also shown flying across the sky, so the science gets a bit creative.

The Northern Lights or Nordlys are visible during winter, and summer brings the Midnight Sun — sunlight 24 hours a day. Norway or Norge is one of the very few locations where sunset merges into the sunrise, with no darkness in between, creating a soft, captivating twilight in which to view the world. 

The Botneheia Formation is made up of dark grey, laminated shales coarsening upwards to laminated siltstones and sandstones. South of the type area, the formation shows four coarsening-upward units. 

The formation is named for Botneheia Mountain, a mountain in Nordenskiöld Land at Spitsbergen, Svalbard. It has a height of 522 m.a.s.l., and is located south of Sassenfjorden, east of the valley of De Geerdalen. 

Svalbard, Norway

I was asked recently if folk head out in the torrential rain or ice and snow to fossil collect. I would generally say yes for those where the potential prize always outweighs the weather. For Svalbard, it is a resounding yes. 

You have to remove the snow cover — or ice if you are impatient or unlucky — to get to the outcrops here. It is well worth the effort. Beneath the icy cover, you find lovely ammonoids and bivalves. 

Tastier still, ichthyosaur remains are found here. The first Triassic ichthyosaurs from Svalbard were found in the early 20th century. Now there are quite a few Triassic and Jurassic ichthyosaur species from this archipelago.

Two ichthyosaur specimens have been recovered that are of particular interest. They comprise part of the trunk and the caudal vertebral column respectively. 

Some features, such as the very high and narrow caudal and posterior thoracic neural spines, the relatively elongate posterior thoracic vertebrae and the long and slender haemapophyses indicate that they probably represent a member of the family Toretocnemidae. 

Ichthyosaur Bones

Numerous ichthyosaur finds are known from the underlying Lower Triassic Vikinghøgda Formation and the overlying Middle to Upper Triassic Tschermakfjellet Formation, the new specimens help to close a huge gap in the fossil record of the Triassic ichthyosaurs from Svalbard. 

There is a resident research group working on the Triassic ichthyosaur fauna, the Spitsbergen Mesozoic Research Group. 

Lucky for them, they often find the fossil remains fully articulated — the bones having retained their spatial relationship to one another. 

Most of their finds are of the tail sections of primitive Triassic ichthyosaurs. In later ichthyosaurs, the tail vertebrae bend steeply downwards and have more of a fish-like look. 

In these primitive ancestors, the tail looks more eel-like — bending slightly so that the spines on the vertebrae form more of the tail. 

Maisch, Michael W. and Blomeier, Dierk published on these finds back in 2009: Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen Band 254 Heft 3 (2009), p. 379 - 384. Nov 1, 2009.

Svalbard, Norway (Norge)

Svalbard was so remote that there were no Inuit or First Nation settlements. It is certainly possible an earlier people came through these islands, but they did not leave any trace of their travels. 

The first documented travellers to explore Spitsbergen arrived in 1795 as part of a hunting expedition. They included people from the arctic town of Hammerfest in Norway's far north. They were an excellent choice as they were used to barren, inhospitable lands and sailed to discover more. 

We know them as the Coast Sámi — a hearty, rugged people probably best known in history for their chieftain, Ottar. He left Hammerfest in the 9th century to visit then join King Alfred the Great's court in a newly forming England. 

Expeditions to the remote islands of Svalbard continued into the early 1800s and finally, a settlement was eked out of the cold landscape and slowly expanded to the rest of the islands. While today the islands are called Svalbard, I would have named them for the Norwegian word for remote — fjernkontroll.

Aristoptychites euglyphus and Daonella sp.

This marvellous block is filled with Aristoptychites (syn = Arctoptychites) euglyphus (Mojsisovics, 1886) and Daonella sp., oyster-like clams or bivalves from the Middle Triassic, Ladinian, rugged windswept outcrops at the top of the Daonella Shales, Botneheia Formation, Spitzbergen, Edgeøya and Barentsøya, eastern Svalbard, Norway. 

Daonella and Monotis are important species for our understanding of biostratigraphy in the Triassic and are useful as Index fossils. 

Index fossils are fossils used to define and identify geologic periods or faunal stages. To be truly useful, they need to have a short vertical range, wide geographic distribution and rapid evolutionary development.

Daonellids preferred soft, soupy substrates and we tend to find them in massive shell beds. Generally, if you find one, you find a whole bunch cemented together in coquina. The lovely block you see here is in the collections of the deeply awesome John Fam. 

Learning Languages

The Sámi languages (/ˈsɑːmi/ SAH-mee), Sami or Saami, are a group of Uralic languages spoken by the Sámi people in Northern Europe in parts of northern Finland, Norway, Sweden, and extreme northwestern Russia. Of the world's languages, I find them the most difficult for my mind and tongue to wrap around. The Uralic languages will be familiar to you as Hungarian (Magyar nyelv), Finnish and Estonian. 

Since my Sámi is terrible, I will share a few words of Norwegian that may come in handy if you visit Svalbard and have a hankering for their tasty fossils or fossiler. To say, ice, snow, reindeer and ichthyosaurs in Norwegian, you would say: is, snø, reinsdyr og ikthyosaurer. 

To say, "hello, where can I find fossils?" Use, "Hei, hvor kan jeg finne fossiler?" An expression you may not need but circumstances being what they are, "That is a big polar bear," is "Det er en stor isbjørn." A solid follow-up would be, "nice bear, run..." as "Fin bjørn, løp..." Good luck with that.

Wishing you and yours the very best of the holidays however you celebrate. 

DINOFLAGELLATES: TEENSY OCEAN STARS

This showy Christmas Cracker is a Dinoflagellate

The showy royal blue Christmas cracker looking fellow you see here is a dinoflagellate. 

Bioluminescent dinoflagellates are a type of plankton — teensy marine organisms that make the seaways shimmer as you swim through them or the tide crashes them against the shore. 

The first modern dinoflagellate was described by Baker in 1753, the first species was formally named by Muller in 1773. 

The first fossil forms were described by Ehrenberg in the 1830s from Cretaceous outcrops. More dinoflagellates have lived, died and gone extinct than there are living today. We know them mainly from fossil dinocysts dating back to the Triassic. They are one of the most primitive of the eukaryotic group with a fossil record that may extend into the Precambrian. They combine primitive characteristics of prokaryotes and advanced eukaryotic features.

The luciferase found in dinoflagellates is related to the green chemical chlorophyll found in plants. Their twinkling lights are brief, each containing about 100 million photons that shine for only a tenth of a second. While each individual flicker is here and gone in the wink of an eye, en masse they are breathtaking. I have spent several wondrous evenings scuba diving amongst these glittering denizens off our shores. What you know about light above the surface does not hold true for the light you see as bioluminescence. Its energy and luminosity come from a chemical reaction. 

In a luminescent reaction, two types of chemicals — luciferin and luciferase — combine together. Together, they produce cold light — light that generates less than 20% thermal radiation or heat. 

The light you see is produced by a compound called Luciferin. It is the shiny, showy bit in this chemical show. Luciferase acts as an enzyme, the substance that acts as a catalyst controlling the rate of chemical reactions, allowing the luciferin to release energy as it is oxidized. 

The colour of the light depends on the chemical structures of the chemicals. There are more than a dozen known chemical luminescent systems, indicating that bioluminescence evolved independently in different groups of organisms.

Coelenterazine is the type of luciferin we find in shrimp, fish and jellyfish. Dinoflagellates and krill share another class of unique luciferins, while ostracods or firefleas and some fish have a completely different luciferin — but all produce lights of various colours to great effect.  

AIOLOCERAS OF MADAGASCAR

Aioloceras besairiei (Collingnon, 1949)

A stunning example of the internal suturing with calcite infill in this sliced Aioloceras besairiei (Collingnon, 1949) ammonite from the Upper Cretaceous (Lower Albian) Boeny region of Madagascar. 

This island country is 400 kilometres off the coast of East Africa in the Indian Ocean and a wonderful place to explore off the beaten track.

Madagascar has some of the most spectacular of all the fossil specimens I have ever seen. This beauty is no exception. The shell has a generally small umbilicus, arched to acute venter, and typically at some growth stage, falcoid ribs that spring in pairs from umbilical tubercles, usually disappearing on the outer whorls. I had originally had this specimen marked as a Cleoniceras besairiei, except Cleoniceras and Grycia are not present in Madagascar. 

This lovely, seen in cross-section, is now far from home and in the collection of a wonderful friend. It is an especially lovely example of the ammonite, Aioloceras besairiei, making it a beudanticeratinae. Cleoniceras and Grycia are the boreal genera. If you'd like to see (or argue) the rationale on the name, consider reading Riccardi and Medina's riveting work from back in 2002, or Collingnon from 1949.

The beauty you see here measures in at a whopping 22 cm, so quite a handful. This specimen is from the youngest or uppermost subdivision of the Lower Cretaceous. I'd originally thought this locality was older, but dating reveals it to be from the Lower Albian, so approximately 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma.

Aioloceras are found in the Cretaceous of Madagascar at geo coordinates 16.5° S, 45.9° E: paleo-coordinates 40.5° S, 29.3° E.; and in four localities in South Africa: at locality 36, near the Mzinene River at 28.0° S, 32.3° E: paleo-coordinates 48.6° S, 7.6° E. 

We find them near the Mziene River, at a second locality north of Hluhluwe where the Mzinene Formation overlies the Aptian-Albian Makatini Formation at 28.0° S, 32.3° E: paleo-coordinates 48.6° S, 7.6° E; and at Haughton Z18, on the Pongola River in the Albian III, Tegoceras mosense beds at 27.3° S, 32.2° E: paleo-coordinates 48.0° S, 7.8° E.

If you happen to be trekking to Madagascar, know that it's big. It’s 592,800 square kilometres (or  226,917 square miles), making it the fourth-largest island on the planet — bigger than Spain, Thailand, Sweden and Germany. The island has an interesting geologic history.

Although there has been a geological survey, which was active extending back well into French colonial times, in the non-French-speaking world our geological understanding of the island is still a bit of a mystery. 

Plate tectonic theory had its beginnings in 1915 when Alfred Wegener proposed his theory of "continental drift." 

Wegener proposed that the continents ploughed through the crust of ocean basins, which would explain why the outlines of many coastlines (like South America and Africa) look like they fit together like a puzzle. Half a century after Wegener there is still no agreement as to whether in continental reconstructions Madagascar should be placed adjacent to the Tanzanian coast to the north (e.g., McElhinny and Embleton,1976), against the Mozambique-Natal coast (Flores 1970), or basically left where it is (Kent 1974, Nairn 1978).

There have been few attempts apart from McKinley’s (1960) comparison of the Karoo succession of southwestern Tanzania with that of Madagascar to follow the famous geological precept of “going to sea.” One critical reason is that although there may be a bibliography of several thousand items dealing with Madagascan geology as Besairie (1971) claims, they are items not generally available to the general public. The vital information gained of the geology of the offshore area by post-World War II petroleum exploration has remained largely proprietary. 

Without this data to draw upon, our understanding remains incomplete. I don't actually mind a bit of a mystery here. It is interesting to speculate on how these geologic puzzle pieces fit together and wait for the big reveal. Still, we have good old Besairie from his 1971, Geologie de Madagascar, and a later précis (Besairie, 1973).

We do know that Madagascar was carved off from the African-South American landmass early on. The prehistoric breakup of the supercontinent Gondwana separated the Madagascar–Antarctica–India landmass from the Africa–South America landmass around 135 million years ago. Madagascar later split from India about 88 million years ago, during the Late Cretaceous, so the native plants and animals on the island evolved in relative isolation. 

It is a green and lush island country with more than its fair share of excellent fossil exposures. Along the length of the eastern coast runs a narrow and steep escarpment containing much of the island's remaining tropical lowland forest. If you could look beneath this lush canopy, you'd see rocks of the Precambrian age stretching from the east coast all the way to the centre of the island. The western edge is made up of sedimentary rock from the Carboniferous to the Quaternary.

Red-Tailed Lemurs, Waiwai & Hedgehog

Madagascar is a biodiversity hotspot. Just as Darwin's finches on the Galápagos were isolated, evolving into distinct species (hello, adaptive radiation), over 90% of the wildlife from Madagascar is found nowhere else. 

The island's diverse ecosystems, like so many on this planet, are threatened by Earth's most deadly species, homo sapien sapiens. 

We arrived back in 490 CE and have been chopping down trees and eating our way through the island's tastier populations ever since. Still, they have cuties like this Red-Tailed Lemur. Awe, right?

Today, beautiful outcrops of wonderfully preserved fossil marine fauna hold appeal for me. The material you see from Madagascar is distinctive — and prolific.

Culturally, you'll see a French influence permeating the language, architecture and legal process. There is a part of me that pictures these lovely Lemurs chatting away in French. "Ah, la vache! Regarde le beau fossile, Hérissonne!"

We see the French influence because good 'ol France invaded sleepy Madagascar back in 1883, during the first Franco-Hova War. Malagasy (the local Madagascarian residents) were enlisted as troops, fighting for France in World War I.  During the Second World War, the island was the site of the Battle of Madagascar between the Vichy government and the British. By then, the Malagasy had had quite enough of colonization and after many hiccuping attempts, reached full independence in 1960. Colonization had ended but the tourist barrage had just begun. You can't stop progress.

If you're interested in learning more about this species, check out the Treatise on Invertebrate Paleontology, Part L (Ammonoidea). R.C. Moore (ed). Geological Soc of America and Univ. Kansas Press (1957), p L394. Or head over to look at the 2002 paper from Riccardi and Medina. 2002. Riccardi, A., C. & Medina, F., A. The Beudanticeratinae and Cleoniceratinae (Ammonitina) from the Lower Albian of Patagonia in Revue de Paléobiologie - 21(1) - Muséum d’Histoire Naturelle de la ville de Genève, p 313-314 (=Aioloceras besairiei (COLLIGNON, 1949). You have Bertrand Matrion to thank for the naming correction. Good to have friends in geeky places!

Collignon, M., 1933, Fossiles cenomaniens d’Antmahavelona (Province d’ Analalave, Madagascar), Ann. Geol. Serv. Min. Madagascar, III, 1934 Les Cephalopods du Trias inferieur de Madagascar, Ann. Paleont. XXII 3 and 4, XXII 1.

Besairie, H., 1971, Geologie de Madagascar, 1. Les terrains sedimentaires, Ann. Geol. Madagascar, 35, p. 463.

J. Boast A. and E. M. Nairn collaborated on a chapter in An Outline of the Geology of Madagascar, that is very readable and cites most of the available geologic research papers. It is an excellent place to begin a paleo exploration of the island.

If you happen to parle français, check out: Madagascar ammonites: http://www.ammonites.fr/Geo/Madagascar.htm

CRINOIDS: BEAUTIES OF ECHINODERMATA

Uintacrinus socialis from Utah, USA

Crinoids are one of my favourite echinoderms. It is magical when all the elements come together to preserve a particularly lovely specimen in such glorious detail. 

If you look closely at the detail here you can see a stunning example of Upper Cretaceous, Santonian age, Uintacrinus socialis — named by O.C. Marsh for the Uinta Mountains of Utah nearly 150 years ago.  

These lovelies are best known from the Smoky Hills Niobrara Formation of central Kansas.

Crinoids are unusually beautiful and graceful members of the phylum Echinodermata. They resemble an underwater flower swaying in an ocean current. 

But make no mistake they are marine animals. Picture a flower with a mouth on the top surface that is surrounded by feeding arms. Awkwardly, add an anus right beside that mouth. 

Crinoids with root-like anchors are called sea lilies. They have graceful stalks that grip the ocean floor. Those in deeper water have longish stalks up to 3.3 ft or a meter in length. Then there are other varieties that are free-swimming with only vestigial stalks. They make up the majority of this group and are commonly known as feather stars or comatulids. 

Unlike the sea lilies, the feather stars can move about on tiny hook-like structures called cirri. It is these same cirri that allow crinoids to latch to surfaces on the seafloor. Like other echinoderms, crinoids have pentaradial symmetry. The aboral surface of the body is studded with plates of calcium carbonate, forming an endoskeleton similar to that in starfish and sea urchins.

These make the calyx somewhat cup-shaped, and there are few, if any, ossicles in the oral (upper) surface, an area we call the tegmen. It is divided into five ambulacral areas, including a deep groove from which the tube feet project, and five interambulacral areas between them. 

Crinoids are alive and well today. They are also some of the oldest fossils on the planet. We have lovely fossil specimens dating back to the Ordovician — if one ignores the enigmatic Echmatocrinus of the Burgess Shale. And they can be quite plentiful. Crinoid fossils, and in particular disarticulated crinoid columnals, can be so abundant that they at times serve as the primary supporting clasts in sedimentary rocks.