CTENOPHORES: COMB JELLIES

Cannibalistic Comb Jellies

This festive lantern looking lovely belongs to a group of invertebrates known as comb jellies.

Comb jellies are named for their unique plates of giant fused cilia, or combs, which run in eight rows up and down the length of their bodies. They are armed with sticky cells or colloblasts, that do not sting but display wonderful bioluminescent colouring as they move through the sea.

Ctenophores or comb jellies are one of the phylogenetically most important and controversial metazoan groups. They are not jellyfish and are not closely related, though they do share some characteristics with the gelatinous members of the subphylum Medusozoa. 

Comb jellies are not picky eaters. Their tastes range to what is at hand, including cannibalizing other comb jellies. They will feast on their kin along with tasty plankton, zooplankton, crustaceans and wee fish.

Interest in their fossil record has been catalysed by spectacularly preserved soft-bodied specimens from Cambrian Lagerstätten of the 518-million-years-old Chengjiang Biota, the 505-million-years-old Burgess Shale and other Burgess Shale-like deposits. 

We find them in the Late Devonian Escuminac Formation at Miguasha National Park, Quebec, Canada — a UNESCO world heritage site famous for its abundance of well-preserved vertebrate fossils including most major evolutionary groups of Devonian lower vertebrates from jawless fish to stem-tetrapods.

Based on morphological similarities of this Canadian fossil with stem-ctenophore fossils from the Cambrian Lagerstätte of the Chinese locality Chengjiang, they have been assessed for their affinity to stem-group ctenophores (dinomischids, Siphusauctum, scleroctenophorans) and early crown-group ctenophores. Modern ctenophores and many fossil forms lack mineralized hard parts, which renders the rare fossils that have been extracted from several Lagerstätten quite remarkable. 

Like the soft bodies of jellyfish and the polyps of hydrozoans and anthozoans, the probability for such soft bodies (or body regions) to become fossilized is extremely low. In spite of this low preservation potential, remains of stem-ctenophores have become known from several Cambrian and younger conservation deposits, and with even older candidate ctenophores in the Ediacaran. 

While Cambrian Lagerstätten have yielded several genera, ctenophore remains are much rarer in the Devonian; in particular, two studies, describing material from the German Hunsrück Slate. 

Bioluminescent Comb Jellies

This Early Devonian material, however, appears to belong to crown ctenophores morphologically similar to living forms such as Pleurobrachia, unlike the stem Cambrian taxa and the new Devonian stem taxon described here.

The most basal stem ctenophores are the dinomischids: sessile benthic petaloid invertebrates, many of which are equipped with a stalk. This group first was described from the Middle Cambrian Burgess Shale. Based on the genus Dinomischus, these early stalked forms were commonly called ‘dinomischids’. 

Zhao et al. shared that dinomischids "form a grade in the lower part of the ctenophore stem group” and include taxa such as Xianguangia, Daihua, and Dinomischus that have hexaradiate-based symmetry (e.g., sixfold, 18-fold). 

Some later, skeletonised stem-ctenophores were termed ‘Scleroctenophora’; ‘scleroctenophorans’ have a shorter stalk, lack the ‘petals’ and have no bracts and might be monophyletic. 

To date, all known dinomischids and scleroctenophorans are Cambrian. Remarkably, analysis of the material described here suggests it is a very late-surviving member of this part of the ctenophore tree, occurring in strata over a hundred million years younger with no intervening known record, thus making it a Lazarus taxon with an extensive ghost lineage. 

Palaeozoic sediments yield a growing number of fossil invertebrates with radial symmetries, some being quite enigmatic with body plans differing radically from those of extant organisms.

The morphological similarities to Cambrian forms and the mix of characters regarding overall shape and symmetries render this discovery important. The aims of this study are to describe the only known specimen of this Devonian ctenophore, discuss its phylogenetic and systematic position, and the impact of fossil data for ctenophore affinities, and assess its palaeoecological role.

AWKWARD AND AWESOME: DIMORPHODON

This remarkable fellow is Dimorphodon — a genus of medium-sized pterosaur from the Early Jurassic. He is another favourite of mine for his charming awkwardness.

You can see this fellow's interesting teeth within his big, bulky skull. Dimorphodon had two distinct types of teeth in their jaws — an oddity amongst reptiles — and also proportionally short wings for their overall size. 

Just look at him. What an amazing beast. We understand their anatomy quite well today, but can you imagine being the first to study their fossils and try to make sense of them. 

The first fossil remains now attributed to Dimorphodon were found in England by fossil collector Mary Anning, at Lyme Regis in Dorset, United Kingdom in December 1828. While she faced many challenges in her life, she was blessed to live in one of the richest areas in Britain for finding fossils. 

She walked the beaches way back in the early 1800s of what would become the Jurassic Coast UNESCO World Heritage Site. The Jurassic Coast holds some of the most interesting fossils ever found — particularly within the strata of the Blue Lias which date back to the Hettangian-Sinemurian. It is one of the world’s most famous fossil sites. Millions come to explore the eroding coastline looking for treasures that provide delight and inspiration to young and old.

 

These fossil treasures provide us with tremendous insights into our world 185 million years ago when amazing animals like Dimorphodon ruled the skies. 

Mary's specimen was acquired by William Buckland and reported in a meeting of the Geological Society on 5 February 1829. Six years later, in 1835, William Clift and William John Broderip built upon the work by Buckland to publish in the Transactions of the Geological Society, describing and naming the fossil as a new species. 

As was the case with most early pterosaur finds, Buckland classified the remains in the genus Pterodactylus, coining the new species Pterodactylus macronyx. The specific name is derived from Greek makros, "large" and onyx, "claw", in reference to the large claws of the hand. The specimen, presently NHMUK PV R 1034, consisted of a partial and disarticulated skeleton on a slab — notably lacking the skull. Buckland in 1835 also assigned a piece of the jaw from the collection of Elizabeth Philpot to P. macronyx. 

Later, the many putative species assigned to Pterodactylus had become so anatomically diverse that they began to be broken into separate genera.

In 1858, Richard Owen reported finding two new specimens, NHMUK PV OR 41212 and NHMUK PV R 1035, again partial skeletons but this time including the skulls. Having found the skull to be very different from that of Pterodactylus, Owen assigned Pterodactylus macronyx its own genus, which he named Dimorphodon. 

His first report contained no description and the name remained a nomen nudum. In 1859, however, a subsequent publication by Owen provided a description. After several studies highlighting aspects of Dimorphodon's anatomy, Owen finally made NHMUK PV R 1034 the holotype in 1874  — 185 million years after cruising our skies the Dimorphodon had finally fully arrived.

TEYLERS OF THE NETHERLANDS

Exceptional fossil starfish Helianthaster preserved in minute detail in pyrite from the Devonian of Bundenbach, Germany.

Helianthaster rhenanus was first described in 1862 by Roemer, based on fossils found in the Bundenbach area in Germany, dating back to the lower Devonian. 

Helianthaster was variously attributed to Asteroidea, Ophiuroidea or to another group (Auluroidea ), but only recently this echinoderm and its close relatives (Helianthasteridae ) have been attributed with some certainty to Asteroidea (Blake, 2009). 

Other very similar starfish were the North Americans Arkonaster (Middle Devonian) and Lepidasterella ( Carboniferousmedium), the latter with 24 arms.

This animal, similar to modern starfish, had a diameter that could exceed 15 centimetres with extended arms. Helianthaster had 14 - 16 arms, elongated and thin, with an aboral surface with granular ossifications. The mouth was wide and composed of rather large oral plates; there were thorns on the adambulacrali, while the central disc was composed of small ossicles.

A study of the type specimen was examined with the use of X- rays. The result was images that seem to confirm the presence of large semicircular muscle flanges along the middle of the arms (Südkamp, ​​2011).

The second image you see here is a specimen from the Teylers Museum in Haarlem, the oldest museum in the Netherlands established in 1778. 

We have a cloth merchant turned banker to thank for both the building and this specimen. And, in a way, the beginnings of nomenclature. Pieter Teyler van der Hulst left us this legacy including many of the museum's specimens and the nest egg that would allow its expansion to the glory we enjoy today. 

Pieter lived next to George Clifford III, the financier of Swedish naturalist Carlo Linnaeus (1707-1778). Pieter's funds aided George in funding Linnaneus' work. In a bit of full circle scientific poetry, it was those dollars and this work that gave us the naming system that allowed us to attach a scientific name to this very specimen through Carl's binomial nomenclature. 

In taxonomy, binomial nomenclature ("two-term naming system"), or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, although they can be based on words from other languages. Such a name is called a binomial name (which may be shortened to just "binomial"), a binomen, binominal name or a scientific name; more informally it is also historically called a Latin name. So, for this lovely specimen, Helianthaster rhenanus is this specimen's Latin name.

In his will, Pieter Teyler decided that his collection and part of his fortune should be used to create a foundation for their promotion, the Teylers Stichting (Teyler foundation). 

Teyler's legacy to the city of Haarlem was divided into two societies Teylers Eerste Genootschap (Dutch: Teyler's First Society ) or 'Godgeleerd Genootschap' ( Theological Society ), aimed at the study of religion, and the Teylers Tweede Genootschap ( Second Society ), dedicated to physics, poetry, history, drawing and numismatics.

The executors of Teyler's wishes, the first directors of Teylers Stichting, decided to establish a centre for study and education. Books, scientific instruments, drawings, fossils and minerals, would be housed under one roof. 

The concept was based on a revolutionary ideal derived from the Enlightenment: people could discover the world independently, without coercion from the church or the state. The example that guided the founders in creating the Teyler Museum was the Mouseion of classical antiquity: a "temple for the muses of the arts and sciences" which would also be a meeting place for scholars and host various collections.

This was a time when science and religion were still intermixed but beginning to divide into separate camps. The world was at war, expeditions were undertaken to secure new lands and trade routes—and the slave trade was slowly being abolished. 

In 1778, Russia controlled Alaska and would not sell to the USA, a country two years old in 1778, for another eighty-nine years in 1867. It was also the year that we lost Carl Linnaeus. He left his scientific work and his legacy of more than 1,600 books covering the literature of natural history from the 15th century to his death, a collection that would become the foundation of the Linnaeus Society, established in his name a decade later in 1788—and to which I am an elected fellow.

Here are some of the world events that happened in 1778, the year this museum was founded to give all of this a bit more context:

• January 18 – Third voyage of James Cook: Captain James Cook, with ships HMS Resolution and HMS Discovery, first views Oahu then Kauai in the Hawaiian Islands of the Pacific Ocean, which he names the Sandwich Islands.

• February 5 – South Carolina becomes the first state to ratify the Articles of Confederation. General John Cadwalader shoots and seriously wounds Major General Thomas Conway in a duel after a dispute between the two officers over Conway's continued criticism of General George Washington's leadership of the Continental Army.

• February 6 – American Revolutionary War – In Paris, the Treaty of Alliance and the Treaty of Amity and Commerce are signed by the United States and France, signalling official French recognition of the new republic.

• February 23 – American Revolutionary War – Friedrich Wilhelm von Steuben arrives at Valley Forge, Pennsylvania and begins to train the American troops.

• March 6–October 24 – Captain Cook explores and maps the Pacific Northwest coast of North America, from Cape Foulweather (Oregon) to the Bering Strait.

• March 10 – American Revolutionary War – George Washington approves the dishonourable discharge of Lieutenant Frederick Gotthold Enslin, for "attempting to commit sodomy, with John Monhort a soldier."

• July 10 – Louis XVI of France declares war on the Kingdom of Great Britain.

• July 27 – American Revolutionary War – First Battle of Ushant – British and French fleets fight to a standoff.

• August 3 – The La Scala Opera House opens in Milan, with the première of Antonio Salieri's Europa riconosciuta.

Many more things happened, of course. Folk were born, fell in love, died—and some left legacies that we still enjoy to this day. 

Photo two by Ghedoghedo, CC BY-SA 4.0.

BACK IN THE USSR: KEPPLERITES

This glorious chocolate block contains the creamy grey ammonite Kepplerites gowerianus (Sowerby 1827) with a few invertebrate friends, including two brachiopods: Ivanoviella sp., Zeilleria sp. and the deep brown gastropod Bathrotomaria sp. There is also a wee bit of petrified wood on the backside.

These beauties hail from Jurassic, Lower Callovian outcrops in the Quarry of Kursk Magnetic Anomaly (51.25361,37.66944), Kursk region, Russia. Diameter ammonite 70мм. 

In the mid-1980s, during the expansion and development of one of the quarries, an unusual geological formation was found. This area had been part of the seafloor around an ancient island surrounded by Jurassic Seas. 

The outcrops of this geological formation turned out to be very rich in marine fossil fauna. This ammonite block was found there years ago by the deeply awesome Emil Black. 

In more recent years, the site has been closed to fossil collecting and is in use solely for the processing and extraction of iron ore deposits. Kursk Oblast is one of Russia's major producers of iron ore. The area of the Kursk Magnetic Anomaly has one of the richest iron-ore deposits in the world. Rare Earth minerals and base metals also occur in commercial quantities in several locations. Refractory loam, mineral sands, and chalk are quarried and processed in the region. 

The Kursk Magnetic Anomaly Quarry is not far from the Sekmenevsk Formation or Sekmenevska Svita in Russian, a Cretaceous (Albian to Cenomanian) terrestrial geologic formation where Pterosaur fossils have been found in the sandstones. 

If you head there for a visit, be sure to check out the Sekmenevska Svita and Oblast's artesian-well water — most refreshing!

JURASSIC SEA URCHIN: AM'DA'MA

This lovely little biscuit is a Holectypus sea urchin from 120 million-year-old deposits from the Lagniro Formation of Madagascar.

The specimen you see here is in the collections of my beautiful friend Ileana. She and I were blessed to meet in China many years ago and formed an unbreakable bond that happens so few times in one's life. 

Holectypus are a genus of extinct echinoids related to modern sea urchins and sand dollars. They were abundant from the Jurassic to the Cretaceous (between 200 million and 65.5 million years ago).

This specimen is typical of Holectypus with his delicate five-star pattern adorning a slightly rounded test and flattened bottom. The specimen has been polished and was harvested both for its scientific and aesthetic value. 

I have many wonderful memories of collecting their modern cousins that live on the north end of Vancouver Island and along the beaches of Balaklava Island. In the Kwak̓wala language of the Kwakiutl or Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, sea urchins are known as a̱m'da̱'ma and it is this name that I hear in my head when I think of them.

In echinoids, the skeleton is almost always made up of tightly interlocking plates that form a rigid structure or test — in contrast with the more flexible skeletal arrangements of starfish, brittle stars, and sea cucumbers. Test shapes range from nearly globular, as in some sea urchins, to highly flattened, as in sand dollars. 

Sea Urchin Detail

Living echinoids are covered with spines, which are movable and anchored in sockets in the test. These spines may be long and prominent, as in typical sea urchins and most have lovely raised patterns on their surface. 

In sand dollars and heart urchins, however, the spines are very short and form an almost felt-like covering. The mouth of most echinoids is provided with five hard teeth arranged in a circle, forming an apparatus known as Aristotle’s lantern.

Echinoids are classified by the symmetry of the test, the number and arrangement of plate rows making up the test, and the number and arrangement of respiratory pore rows called petals. Echinoids are divided into two subgroups: regular echinoids, with nearly perfect pentameral (five-part) symmetry; and irregular echinoids with altered symmetry.

Because most echinoids have rigid tests, their ability to fossilize is greater than that of more delicate echinoderms such as starfish, and they are common fossils in many deposits. The oldest echinoids belong to an extinct regular taxon called the Echinocystitoidea. 

They first appeared in the fossil record in the Late Ordovician. Cidaroids or pencil urchins appear in the Mississippian (Early Carboniferous) and were the only echinoids to survive the mass extinction at the Permo-Triassic boundary. Echinoids did not become particularly diverse until well after the Permo-Triassic mass extinction event, evolving the diverse forms we find them in today. 

True sea urchins first appear in the Late Triassic, cassiduloids in the Jurassic, and spatangoids or heart urchins in the Cretaceous. Sand dollars, a common and diverse group today, do not make an appearance in the fossil record until the Paleocene. They remain one of my favourite echinoderms and stand tall amongst the most pleasing of the invertebrates.

CORAL: CLIMATE CHANGE INDICATORS

A lovely specimen of fossilized coral. Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically live in compact colonies of many identical individual polyps.

Annual growth bands in some corals, such as the deep-sea bamboo coral, Isididae, may be among the first signs of the effects of ocean acidification on marine life. The growth rings allow geologists to construct year-by-year chronologies, a form of incremental dating, which underlies high-resolution records of past climatic and environmental changes using geochemical techniques.

Certain species form communities called microatolls, which are colonies whose top is dead and mostly above the waterline, but whose perimeter is mostly submerged and alive. The average tide level limits their height. By analyzing the various growth morphologies, microatolls offer a low-resolution record of sea-level change and help us to reconstruct Holocene sea levels. Fossilized microatolls can also be dated using Radiocarbon dating, a method of discovering how old something is by measuring the decay of the radioactive isotope Carbon-14. 

Increasing sea temperatures in tropical regions, ~1 degree C, over the last century have caused major coral bleaching, death, and collapsing of coral populations although they are able to adapt and acclimate. It is uncertain if this evolutionary process will happen quickly enough to prevent a major reduction in their numbers.

Though coral has large sexually-reproducing populations, their evolution can be slowed by abundant asexual reproduction. Gene flow is variable among our coral friends. According to the biogeography of coral species gene flow cannot be counted on as a dependable source of adaptation as they are very stationary organisms. Also, coral longevity might factor into their adaptivity.

However, adaptation to climate change has been demonstrated in many cases. These are usually due to a shift in coral and zooxanthellae genotypes. These shifts in allele frequency have progressed toward more tolerant types of zooxanthellae. Scientists found that a certain scleractinian zooxanthella is becoming more common where sea temperature is high. Symbionts able to tolerate warmer water seem to photosynthesize more slowly, implying an evolutionary trade-off.

In the Gulf of Mexico, where sea temperatures are rising, cold-sensitive staghorn and elkhorn coral have shifted in location. Not only have the symbionts and specific species been shown to shift, but there seems to be a certain growth rate favourable to selection. Slower-growing but more heat-tolerant corals have become more common. The changes in temperature and acclimation are complex. Some reefs in current shadows represent a refugium location that will help them adjust to the disparity in the environment even if eventually the temperatures may rise more quickly there than in other locations. This separation of populations by climatic barriers causes a realized niche to shrink greatly in comparison to the old fundamental niche.

MIGHTY EAGLE: KWIKW

Bald Eagle / Kwikw / Haliaeetus leucocephalus

A mighty Bald Eagle sitting with wings spread looks to be controlling the weather with his will as much as being subject to it. This fellow has just taken a dip for his evening meal and is drying his feathers in the wind. 

As you can imagine, waterlogged feathers make flight difficult. Their wings are built for graceful soaring and gliding on updrafts of warm air called thermals. 

Their long feathers are slotted, easily separating so air flows smoothly and giving them the added benefit of soaring at slower speeds. 

As well as his wings, this fellow is also drying off his white head feathers. A bald eagle's white head can make it look bald from a distance but that is not where the name comes from. It is from the old English word balde, meaning white.

In the Kwak'wala language of the Kwakiutl First Nations of the Pacific Northwest — or Kwakwaka'wakw, speakers of Kwak'wala — an eagle is known as kwikw (kw-ee-kw) and an eagle's nest is called a kwigwat̕si. 

Should you encounter an eagle and wish to greet them in Kwak'wala, you would just say yo. Yup, just yo. They would like your yo hello better if you offered them some fresh fish. They dine on all sorts of small mammals, fish and birds but are especially fond of pink salmon or ha̱nu'n (han-oon).

These living dinosaurs are a true homage to their lineage. They soar our skies with effortless grace. Agile, violent and beautiful, these highly specialized predators can catch falling prey mid-flight and dive-bomb into rivers to snag delicious salmon. 

Their beauty and agility are millions of years in the making. From their skeletal structure to their blood cells, today’s birds share a surprising evolutionary foundation with reptiles. 

Between 144 million and 66 million years ago, during the Mesozoic era, we see the first birds evolve. Eventually, tens of millions of years ago, an ancient group of birds called kites developed. Like today’s bald eagle, early kites are thought to have scavenged and hunted fish.

About 36 million years ago, the first eagles descended from kites, their smaller cousins. First to appear were the early sea eagles, which — like kites — continued to prey on fish and whose feet were free of feathers, along with booted eagles, which had feathers below the knee. Fossils of Bald Eagles are very rare and date to the late Pleistocene. Eagles are known from the early Pleistocene of Florida, but they are extinct species not closely related to the bald eagle.

Like the kites, bald eagles have featherless feet, but they also developed a range of other impressive adaptations that help them hunt fish and fowl in a watery environment. Each foot has four powerful toes with sharp talons. Tiny projections on the bottom of their feet called “spicules” help bald eagles grasp their prey. A bald eagle also has serrations on the roof of its mouth that help it hold slippery fish, and incredibly, the black pigment in its wing feathers strengthens them against breakage when they dive head first into water.

Obviously, there is much more than their striking white heads that sets these iconic raptors apart from the crowd. Their incredible physiology, built for life near the water, is literally millions of years in the making. 

FRAGILE BEAUTY: FOSSILIZED SCLERACTINIAN CORAL

Scleractinian Fossil Coral, Florida

The delicate wintery beauty you see here is a Scleractinian coral we find first in the fossil record in the Mesozoic. 

Corals first appeared in the Cambrian about 535 million years ago. Fossils are extremely rare until the Ordovician period, 100 million years later, when rugose and tabulate corals became widespread. 

Palaeozoic corals seem to make friends wherever they live and often contain numerous endobiotic symbionts.

Tabulate corals occur in limestones and calcareous shales of the Ordovician and Silurian periods, and often form low cushions or branching masses of calcite alongside rugose corals. 

Their numbers began to decline during the middle of the Silurian period, and they became extinct at the end of the Permian period, 250 million years ago.

Rugose or horn corals became dominant by the middle of the Silurian and became extinct early in the Triassic period. The rugose corals existed in solitary and colonial forms and were also composed of calcite.

The famous Great Barrier Reef is thought to have been laid down about two million years ago. If you have had the pleasure of scuba diving near it to take in its modern wonders, perhaps you will be interested to learn how it was formed. Over long expanses of time, the corals here have broken up, fragmented and died. Sand and rubble accumulate between the corals, and the shells of clams and other molluscs decay to form a gradually evolving calcium carbonate structure to what you view today. 

Coral reefs are extremely diverse marine ecosystems hosting over 4,000 species of fish, massive numbers of cnidarians, molluscs, crustaceans, and many other animals.

BLADDER-BEARERS: HOODED SEALS

If you frequent the eastern coast of North America north of Maine to the western tip of Europe, along the coast of Norway near Svalbard you may have glimpsed one of their chubby, dark silver-grey and white residents. 

Hooded seals, Cystophora cristata, are large phocid seals in the family Phocidae, who live in some of the chilliest places on Earth, from 47° to 80° N in latitude. 

These skilled divers are mainly concentrated around Bear Island, Norway, Iceland, and northeast Greenland. 

In rare cases, we find them in the icy waters in Siberia. They usually dive depths of 600 m (1,968 ft) in search of fishy treats but can go as deep as 1000 m (3,280 ft) when needed. That is deep into the cold, dark depths of our oceans. Sunlight entering the sea may travel as deep as 1,000 m (3,280 ft) under the right conditions, but there is rarely any significant light beyond 200 meters (656 ft). This is the dark zone and the place we find our bioluminescent friends. 

Hooded seals have a sparse fossil record. One of the first fossils found was a Pliocene specimen from Anvers, Belgium discovered in 1876. In 1983 a paper was published claiming there were some fossils found in North America thought to be from Cystophora cristata. Of the three accounts, the most creditable discovery was from a sewer excavation in Maine, the northeasternmost U.S. state, known for its rocky coastline, maritime history and nature areas like the granite and spruce islands of Acadia National Park. A scapula and humeri were found among other bones and thought to date to the post-Pleistocene. 

Of two other accounts, one was later reassigned to another species and the other left unsolved. (Folkow, et al., 2008; Kovacs and Lavigne, 1986; Ray, 1983)

The seals are typically silver-grey or white in colour, with black spots that vary in size covering most of the body. 

Hooded seal pups are known as, Blue-backs as their coats are blue-grey on the back with whitish bellies, though this coat is shed after 14 months of age when the pups moult.

FIRST NATION, INUIT, METIS, MI'KMAQ L'NU

In the Kwak̓wala language of the Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, seal are known as migwat — and fur seals are known as x̱a'wa.

Hooded seals live primarily on drifting pack ice and in deep water in the Arctic Ocean and North Atlantic. Although some drift away to warmer regions during the year their best survival rate is in colder climates. They can be found on four distinct areas with pack ice: near Jan Mayen Island, northeast of Iceland; off Labrador and northeastern Newfoundland; the Gulf of St. Lawrence; and the Davis Strait, off midwestern Greenland. 

The province of Newfoundland and Labrador is home to the Inuit, the Innu, the Mi'kmaq L'nu and the Southern Inuit of NunatuKavut, formerly the Labrador Inuit-Metis. The Hooded Seals that visit their traditional territory were a welcome source of food and clothing. In Mi'kmaw, the language spoken in Mi'kma'ki, the territory of the Mi'kmaq L'nu, the word for seal is waspu.

HOODED SEAL HABITAT

Males are localized around areas of complex seabeds, such as Baffin Bay, Davis Strait, and the Flemish Cap. Females concentrate their habitat efforts primarily on shelf areas, such as the Labrador Shelf. 

Females reach the age of sexual maturity between two and nine years old and it is estimated that most females give birth to their first young at around five years of age. Males reach sexual maturity a little later around four to six years old but often do not mate until much later. Females give birth to one young at a time through March and April. The gestation period is 240 to 250 days. 

Blue-back, Hooded Seal Pup

During this time the fetus, unlike those of other seals, sheds its lanugo — a covering of fine soft hair that is replaced by thicker pelage — in the uterus. 

These young are precocious and at birth are able to move about and swim with ease. They are independent and left to fend for themselves immediately after they have been weaned.

Hooded seals are known to be a highly migratory species that often wander long distances, as far west as Alaska and as far south as the Canary Islands and Guadeloupe. 

Prior to the mid-1990s, hooded seal sightings in Maine and the east Atlantic were rare but began increasing in the mid-1990s. From January 1997 to December 1999, a total of 84 recorded sightings of hooded seals occurred in the Gulf of Maine, one in France and one in Portugal. 

From 1996 to 2006, five strandings and sightings were noted near the Spanish coasts in the Mediterranean Sea. There is no scientific explanation for the increase in sightings and range of the hooded seal.

Cystophora means "bladder-bearer" in Greek and pays homage to this species' inflatable bladder septum on the heads of adult males. The bladder hangs between the eyes and down over the upper lip in a deflated state. 

The hooded seal can inflate a large balloon-like sac from one of its nostrils. This is done by shutting one nostril valve and inflating a membrane, which then protrudes from the other nostril. 

I was thinking of Hooded seals when contemplating the nasal bladders of Prosaurolophus maximum, large-headed duckbill dinosaurs, or hadrosaurid, in the ornithischian family Hadrosauridae. Perhaps both species used these bladders in a similar manner — to warn predators and attract mates.

Hooded seals are known for their uniquely elastic nasal cavity located at the top of their head, also known as the hood. Only males possess this display-worthy nasal sac, which they begin to develop around the age of four. The hood begins to inflate as the seal makes its initial breath prior to going underwater. It then begins to repetitively deflate and inflate as the seal is swimming. 

The purpose of this is acoustic signaling. It occurs when the seal feels threatened and attempt to ward off hostile species when competing for resources such as food and shelter. It also serves to communicate their health and superior status to both other males and females they are attempting to attract. 

In sexually mature males, a pinkish balloon-like nasal membrane comes out of the left nostril to further aid it in attracting a mate. This membrane, when shaken, is able to produce various sounds and calls depending on whether the seal is underwater or on land. Most of these acoustic signals are used in an acoustic situation (about 79%), while about 12% of the signals are used for sexual purposes.

References: Ray, C. 1983. Hooded Seal, Cystophora cristata: Supposed Fossil Records in North America. American Society of Mammalogists, Vol. 64 No. 3: 509-512; Cystophora cristata, Hooded Seal", 2007; "Seal Conservation Society", 2001; Kovacs and Lavigne, 1986.

Mi'kmaq Online Dictionary: https://www.mikmaqonline.org/servlet/dictionaryFrameSet.html?method=showCategory&arg0=animal

OYSTER: T'LOXT'LOX

One of the now rare species of oysters in the Pacific Northwest is the Olympia oyster, Ostrea lurida, (Carpenter, 1864).  

While rare today, these are British Columbia’s only native oyster. 

Had you been dining on their brethren in the 1800s or earlier, it would have been this species you were consuming. Middens from Port Hardy to California are built from Ostrea lurida.

These wonderful invertebrates bare their souls with every bite. Have they lived in cold water, deep beneath the sea, protected from the sun's rays and heat? Are they the rough and tumble beach denizens whose thick shells tell us of a life spent withstanding the relentless pounding of the sea? Is the oyster in your mouth thin and slimy having just done the nasty—spurred by the warming waters of Spring? 

Is this oyster a local or was it shipped to your current local and, if asked, would greet you with "Kon'nichiwa?" Not if the beauty on your plate is indeed Ostrea lurida. 

Oyster in Kwak'wala is t̕łox̱t̕łox̱

We have been cultivating, indeed maximizing the influx of invasive species to the cold waters of the Salish Sea for many years. 

But in the wild waters off the coast of British Columbia is the last natural abundant habitat of the tasty Ostrea lurida in the pristine waters of  Nootka Sound. 

The area is home to the Nuu-chah-nulth First Nations who have consumed this species boiled or steamed for thousands of years. Here these ancient oysters not only survive but thrive — building reefs and providing habitat for crab, anemones and small marine animals. 

Oysters are in the family Ostreidae — the true oysters. Their lineage evolved in the Early Triassic — 251 - 247 million years ago. 

In the Kwak̓wala language of the Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest and my family, an oyster is known as t̕łox̱t̕łox̱. 

I am curious to learn if any of the Nuu-chah-nulth have a different word for an oyster. If you happen to know, I would be grateful to learn.

ANCIENT BEAUTIES: FOSSIL PEARLS

One of my favourite pairs of earrings are a simple set of pearls. I have worn them pretty much every day since 2016 when I received them as a gift. What is it about pearls that makes them so appealing? I am certainly not alone in this. 

A simple search will show you a vast array of pearls being used for their ornamental value in cultures from all over the world. I suppose the best answer to why they are appealing is just that they are. 

If you make your way to Paris, France and happen to visit the Louvre's Persian Gallery, do take a boo at one of the oldest pearl necklaces in existence — the Susa necklace. It hails from a 2,400-year-old tomb of long lost Syrian Queen. It is a showy piece with three rows of 72 pearls per strand strung upon a bronze wire. 

A queen who truly knew how to accessorize. 

I imagine her putting the final touches of her outfit together, donning the pearls and making an entrance to wow the elite of ancient Damascus. The workmanship is superb, intermixing pure gold to offset the lustre of the pearls. It is precious and ancient, crafted one to two hundred years before Christ. Perhaps a gift from an Egyptian Pharaoh or from one of the Sumerians, Eblaites, Akkadians, Assyrians, Hittites, Hurrians, Mitanni, Amorites or Babylonian dignitaries who sued for peace but brought war instead. 

Questions, good questions, but questions without answers. So, what can we say of pearls? We do know what they are and it is not glamorous. Pearls form in shelled molluscs when a wee bit of sand or some other irritant gets trapped inside the shell, injuring the flesh. As a defensive and self-healing tactic, the mollusc wraps it in layer upon layer of mother-of-pearl — that glorious shiny nacre that forms pearls. 

They come in all shapes and sizes from minute to a massive 32 kilograms or 70 pounds. While a wide variety of our mollusc friends respond to injury or irritation by coating the offending intruder with nacre, there are only a few who make the truly gem-y pearls. 

These are the marine pearl oysters, Pteriidae and a few freshwater mussels. Aside from Pteriidae and freshwater mussels, we sometimes find less gem-y pearls inside conchs, scallops, clams, abalone, giant clams and large marine gastropods.

Pearls are made up mostly of the carbonate mineral aragonite, a polymorphous mineral — the same chemical formula but different crystal structure — to calcite and vaterite, sometimes called mu-calcium carbonate. These polymorphous carbonates are a bit like Mexican food where it is the same ingredients mixed in different ways. Visually, they are easy to tell apart — vaterite has a hexagonal crystal system, calcite is trigonal and aragonite is orthorhombic.

As pearls fossilize, the aragonite usually gets replaced by calcite, though sometimes by vaterite or another mineral. When we are very lucky, that aragonite is preserved with its nacreous lustre — that shimmery mother-of-pearl we know and love.  

Molluscs have likely been making pearls since they first evolved 530 million years ago. The oldest known fossil pearls found to date, however, are 230-210 million years old. 

This was the time when our world's landmass was concentrated into the C-shaped supercontinent of Pangaea and the first dinosaurs were calling it home. In the giant ancient ocean of Panthalassa, ecosystems were recovering from the high carbon dioxide levels that fueled the Permian extinction. Death begets life. With 95% of marine life wiped out, new species evolved to fill each niche.  

While this is where we found the oldest pearl on record, I suspect we will one day find one much older and hopefully with its lovely great-great grandmother-of-pearl intact. 

LOVE THE WILD: KU'MIS

Look how epic this little guy is! 

He is a crab — and if you asked him, the fiercest warrior that ever lived. While that may not be strictly true, crabs do have the heart of a warrior and will raise their claws, sometimes only millimetres into the air, to assert dominance over their world. 

Crabs are decapod crustaceans of the Phylum Arthropoda. 

In the Kwak'wala language of the Kwakwaka'wakw of the Pacific Northwest, this brave fellow is ḵ̓u'mis — both a tasty snack and familiar to the supernatural deity Tuxw'id, a female warrior spirit. Given their natural armour and clear bravery, it is a fitting role.

They inhabit all the world's oceans, sandy beaches, many of our freshwater lakes and streams. Some few prefer to live in forests.

Crabs build their shells from highly mineralized chitin — and chitin gets around. It is the main structural component of the exoskeletons of many of our crustacean and insect friends. Shrimp, crab, and lobster all use it to build their exoskeletons.

Chitin is a polysaccharide — a large molecule made of many smaller monosaccharides or simple sugars, like glucose. 

It is handy stuff, forming crystalline nanofibrils or whiskers. Chitin is actually the second most abundant polysaccharide after cellulose. It is interesting as we usually think of these molecules in the context of their sugary context but they build many other very useful things in nature — not the least of these are the hard shells or exoskeletons of our crustacean friends.

Crabs in the Fossil Record

The earliest unambiguous crab fossils date from the Early Jurassic, with the oldest being Eocarcinus from the early Pliensbachian of Britain, which likely represents a stem-group lineage, as it lacks several key morphological features that define modern crabs. 

Most Jurassic crabs are only known from dorsal — or top half of the body — carapaces, making it difficult to determine their relationships. Crabs radiated in the Late Jurassic, corresponding with an increase in reef habitats, though they would decline at the end of the Jurassic as the result of the decline of reef ecosystems. Crabs increased in diversity through the Cretaceous and represented the dominant group of decapods by the end.

We find wonderful fossil crab specimens on Vancouver Island. The first I ever collected was at Shelter Point, then again on Hornby Island, down on the Olympic Peninsula and along Vancouver Island's west coast near Nootka Sound. They are, of course, found globally and are one of the most pleasing fossils to find and aggravating to prep of all the specimens you will ever have in your collection. Bless them.

EXPLORING THE EAST KOOTENAY REGION

The East Kootenay region on the south-eastern edge of British Columbia is a land of colossal mountains against a clear blue sky. 

That is not strictly true, of course, as this area does see its fair share of rain and temperature extremes — but visiting in Autumn every view is a postcard of mountainous terrain.

Rocks from deep within the Earth's crust underlie the entire East Kootenay region and are commonly exposed in the areas majestic mountain peaks, craggy rocky cliffs, glaciated river canyons, and rock cuts along the highways. Younger Ice Age sediments blanket much of the underlying rock.

I've been heading to the Cranbrook and Fernie area since the early 1990s. My interest is the local geology and fossil history that these rocks have to tell. I'm also drawn to the warm and welcoming locals who share a love for the land and palaeontological treasures that open a window to our ancient past.  

Cranbrook is the largest community in the region and is steeped in mining history and the opening of the west by the railway. It is also a stone's throw away from Fort Steele and the Lower Cambrian exposures of the Eager Formation. These fossil beds rival the slightly younger Burgess Shale fauna and while less varied, produce wonderful examples of olenellid trilobites and weird and wonderful arthropods half a billion years old. 

Labiostria westriopi, McKay Group

The Lower Cambrian Eager Formation outcrops at a few localities close to Fort Steele, many known since the early 1920s, and up near Mount Grainger near the highway. 

Further east, the Upper Cambrian McKay Group near Tanglefoot Mountain is a palaeontological delight with fifteen known outcrops that have produced some of the best-preserved and varied trilobites in the province — many of them new species. 

The McKay Formation also includes Ordovician outcrops sprinkled in for good measure.

Other cities in the area and the routes to and from them produce other fossil fauna from Kimberley to Fernie and the district municipality of Invermere and Sparwood. This is an arid country with native grasslands and forests of semi-open fir and pine. Throughout there are a host of fossiliferous exposures from Lower Cretaceous plants to brachiopods. 

The area around Whiteswan Lake has wonderful large and showy Ordovician graptolites including Cardiograptus morsus and Pseudoclimacograptus angustifolius elongates — some of our oldest relatives. A drive down to Flathead will bring you to ammonite outcrops and you can even find Eocene fresh-water snails in the region. 

The drive from Cranbrook to Fernie is about an hour and change through the Cambrian into the Devonian which flip-flops and folds over revealing Jurassic exposures. 

Fernie Ichthyosaur Excavation, 1916

The Crowsnest Highway into Fernie follows Mutz Creek. From the highway, you can see the Fernie Group and the site along the Elk River where an ichthyosaur was excavated in 1916. 

The Fernie Formation is Jurassic. It is present in the western part of the Western Canada Sedimentary Basin in western Alberta and northeastern British Columbia. 

It takes its name from the town of Fernie, British Columbia, and was first defined by W.W. Leach in 1914. The town of Fernie is rimmed by rugged mountains tipped with Devonian marine outcrops. In essence, all these mountains are upside down with the oldest layers flipped to the top and a good 180 million years older than those they sit upon. 

Before they were mountains, these sedimentary rocks were formed as sediment collected in a shallow sea or inland basin. About 360 million years ago, the rocks that you see in Fernie today were down near the equator. They road tectonic plates, pushing northeast smashing into the coastline of what would become British Columbia. A little push here, shove there — compression and thrust faulting — and the rock was rolled over on its head — repeatedly. But that is how mountains are often formed, though not usually pushed so hard that they flip over. But still, it is a slow, relentless business. 

Cretaceous Plant Material, Fernie, BC

Within Fernie, there are small exposures of Triassic and Jurassic marine outcrops. East of the town there are Cretaceous plant sites, and of course, the Jurassic 1.4-metre Titanites occidentalis ammonite up on Coal Mountain.

The regional district's dominant landform is the Rocky Mountain Trench, which is flanked by the Purcell Mountains and the Rocky Mountains on the east and west, and includes the Columbia Valley region. The southern half of which is in the regional district — its northern half is in the Columbia-Shuswap Regional District. 

The regional district of Elk Valley in the southern Rockies is the entryway to the Crowsnest Pass and an important coal-mining area. 

Other than the Columbia and Kootenay Rivers, whose valleys shape the bottomlands of the Rocky Mountain Trench, the regional districts form the northernmost parts of the basins of the Flathead, Moyie and Yahk Rivers. 

The Moyie and Yahk are tributaries of the Kootenay, entering it in the United States, and the Flathead is a tributary of the Clark Fork into Montana.

Photo One: Tyaughton Mountain, Mckay Group; Photo Two: Labiostria westriopi, Upper Cambrian McKay Group, Site ML (1998); John Fam Collection; Photo Three: Ichthyosaur Excavation, Fernie, British Columbia, 1916; Photo Four: Cretaceous Plant Fossils, east of Fernie towards Coal Mountain. The deeply awesome Guy Santucci as hand-model for scale. 

LOVE LESSONS FROM THE NORTH

Nunatsiarmiut Mother and Child, Baffin Island, Nunavut

Warm light bathes this lovely Nunatsiarmiut mother and child from Baffin Island, Nunavut. 

They speak Inuktitut, the mother tongue of the majority of the Nunatsiarmiut who call Baffin Island home. 

Baffin is the largest island in the Arctic Archipelago in the territory of Nunavut in Canada's far north. 

As part of the Qikiqtaaluk Region of Nunavut, Baffin Island is home to a constellation of remote Inuit communities each with a deep cultural connection to the land—Iqaluit, Pond Inlet, Pangnirtung, Clyde River, Arctic Bay, Kimmirut and Nanisivik. 

The ratio of Inuit to non-Inuit here is roughly three to one and perhaps the reason why the Inuktitut language has remained intact and serves as the mother tongue for more than 36,000 residents. Inuktitut has several subdialects—these, along with a myriad of other languages—are spoken across the north.  

If you look at the helpful visual below you begin to get a feel for the diversity of these many tongues. The languages vary by region. There is the Iñupiaq of the Inupiatun/Inupiat; Inuvialuktun of the Inuinnaqtun, Natsilingmiutut, Kivallirmiutut, Aivilingmiutut, Qikiqtaaluk Uannanganii and Siglitun. Kalaallisut is spoken by many Greenlandic peoples though, in northwest Greenland, Inuktun is the language of the Inughuit.

We use the word Inuktitut when referring to a specific dialect and inuktut when referring to all the dialects of Inuktitut and Inuinnaqtun.

Northern Language Map (Click to Enlarge)

Should you travel to the serene glacier-capped wilds and rolling tundra of our far north, you will want to dress for the weather and learn a few of the basics to put your best mukluk shod feet forward. 

The word for hello or welcome in Inuktitut is Atelihai—pronounced ahh-tee-lee-hi. And thank you is nakurmiik, pronounced na-kur-MIIK.  

Perhaps my favourite Inuktitut expression is Naglingniq qaikautigijunnaqtuq maannakautigi, pronounced NAG-ling-niq QAI-kau-ti-gi-jun-naqtuq MAAN-na-KAU-ti-gi. This tongue-twister is well worth the linguistic challenge as it translates to love can travel anywhere in an instant. Indeed it can. 

So much of our Indigenous culture is passed through stories, so language takes on special meaning in that context. It is true for all societies but especially true for the Inuit. Stories help connect the past to the present and future. They teach how to behave in society, engage with the world and how to survive in the environment. They also help to create a sense of belonging. 

You have likely seen or heard the word Eskimo used in older books to refer to the Inuit, Iñupiat, Kalaallit or Yupik. This misnomer is a colonial term derived from the Montagnais or Innu word ayas̆kimew—netter of snowshoes. It is a bit like meeting a whole new group of people who happen to wear shoes and referring to them all as cobblers—not as a nickname, but as a legal term to describe populations from diverse communities disregarding the way each self-refer. 

Inukshuk / Inuksuk Marker Cairn

For those who identify as Inupiaq or Yupik, the preferred term is Inuit meaning people—though some lingering use of the term Eskimo lives on. The Inuit as a group are made up of many smaller groups. 

The Inuit of Greenland self-refer as Kalaallit or Greenlanders when speaking Kalaallisut. The Inupiat of Alaska, or real people, use Yupik as the singular for real person and yuk to simply mean person.

When taken all together, Inuit is used to mean all the peoples in reference to the Inuit, Iñupiat, Kalaallit and Yupik. Inuit is the plural of inuk or person. 

You likely recognize this word from inuksuk or inukshuk, pronounced ih-nook-suuk — the human-shaped stone cairns built by the Inuit, Iñupiat, Kalaallit, Yupik, and other peoples of the Arctic regions of northern Canada, Greenland, and Alaska—helpful reference markers for hunters and navigation. The word inuksuk means that which acts in the capacity of a human, combining inuk or person and suk, to substitute. 

A World of Confusion

You may be disappointed to learn that our northern friends do not live in igloos. I remember answering the phone as a child and the fellow calling was hoping to speak to my parents about some wonderful new invention perfect for use in an igloo. 

He was disappointed to hear that I was standing in a wooden house with the standard four walls to a room and a handy roof topping it off. "Well, what about your neighbours? Surely, a few of them live in igloos..." 

It seems that some atlases in circulation at the time, and certainly the one he was looking at, simply blanketed everything north of the 49th parallel in a snowy white. His clearly showed an igloo sitting proudly in the centre of the province.

My cousin Shawn brought one such simplified book back from his elementary school in California. British Columbia had a nice image of a grizzly bear and a wee bit further up, a polar bear grinned smugly. British Columbia's beaver population would be sad to know that they did not inhabit the province though there were two chipper beavers with big bright smiles—one in Ontario and another gracing the province of Quebec. Further north, where folk do build igloos, their icy domes were curiously lacking. 

Igloos are used for winter hunting trips much the same way we use tents for camping. The Inuit do not have fifty words for snow—you can thank the ethnographer Franz Boas for that wee fabrication—but within the collective languages of the frozen north there are more than fifty words to describe it. And kisses are not nose-to-nose. To give a tender kiss or kunik to a loved one, you press your nose and upper lip to their forehead or cheek and rub gently. 

Fancy trying a wee bit of Inuktitut yourself? This link will bring you to a great place to start: https://inhabitmedia.com/inuitnipingit/

Inuit Language Map:  By Noahedits - Own work, CC BY-SA 4.0. If you want to the image full size, head to this link: https://commons.wikimedia.org/w/index.php?curid=85587388

GULLS ON THE FORESHORE: T'SIK'WI

A gull cries in protest at not getting his share of a meal

Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. 

The Laridae are known from not-yet-published fossil evidence from the Early Oligocene — 30–33 million years ago. 

Three gull-like species were described by Alphonse Milne-Edwards from the early Miocene of Saint-Gérand-le-Puy, France. 

Another fossil gull from the Middle to Late Miocene of Cherry County, Nebraska, USA, has been placed in the prehistoric genus Gaviota. 

These fossil gulls, along with undescribed Early Oligocene fossils are all tentatively assigned to the modern genus Larus. Among those of them that have been confirmed as gulls, Milne-Edwards' "Larus" elegans and "L." totanoides from the Late Oligocene/Early Miocene of southeast France have since been separated in Laricola.

Gulls are most closely related to the terns in the family Sternidae and only distantly related to auks, skimmers and distantly to waders. 

A historical name for gulls is mews, which is cognate with the German möwe, Danish måge, Swedish mås, Dutch meeuw, Norwegian måke/måse and French mouette. We still see mews blended into the lexicon of some regional dialects.

In the Kwak̓wala language of the Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest and my family, gulls are known as t̕sik̕wi. Most folk refer to gulls from any number of species as seagulls. This name is a local custom and does not exist in the scientific literature for their official naming. Even so, it is highly probable that it was the name you learned for them growing up.

If you have been to a coastal area nearly everywhere on the planet, you have likely encountered gulls. They are the elegantly plumed but rather noisy bunch on any beach. You will recognize them both by their size and colouring. 

Gulls are typically medium to large birds, usually grey or white, often with black markings on the head or wings. They typically have harsh shrill cries and long, yellow, curved bills. Their webbed feet are perfect for navigating the uneven landscape of the foreshore when they take most of their meals. 

Most gulls are ground-nesting carnivores that take live food or scavenge opportunistically, particularly the Larus species. Live food often includes crab, clams (which they pick up, fly high and drop to crack open), fish and small birds. Gulls have unhinging jaws which allow them to consume large prey which they do with gusto. 

Their preference is to generally live along the bountiful coastal regions where they can find food with relative ease. Some prefer to live more inland and all rarely venture far out to sea, except for the kittiwakes. 

The larger species take up to four years to attain full adult plumage, but two years is typical for small gulls. Large white-headed gulls are typically long-lived birds, with a maximum age of 49 years recorded for the herring gull.

Gulls nest in large, densely packed, noisy colonies. They lay two or three speckled eggs in nests composed of vegetation. The young are precocial, born with dark mottled down and mobile upon hatching. Gulls are resourceful, inquisitive, and intelligent, the larger species in particular, demonstrating complex methods of communication and a highly developed social structure. Many gull colonies display mobbing behaviour, attacking and harassing predators and other intruders. 

Certain species have exhibited tool-use behaviour, such as the herring gull, using pieces of bread as bait with which to catch goldfish. Many species of gulls have learned to coexist successfully with humans and have thrived in human habitats. Others rely on kleptoparasitism to get their food. Gulls have been observed preying on live whales, landing on the whale as it surfaces to peck out pieces of flesh. They are keen, clever and always hungry.

WEYLA OF THE SUNRISE FORMATION

Weyla (Nielsen, 1963) New York Canyon, Nevada

A lovely example of the large bivalve, Weyla (Nielsen, 1963), from the earliest known Jurassic Ferguson Hill Member (Hettangian and Sinemurian) of the Sunrise Formation in the New York Canyon area of west-central Nevada, USA.

The end-Triassic mass extinction was global, severe, and accompanied by worldwide disturbance to carbonate ramp and platform sedimentation. We see the effects played out in the Ferguson Hill Member of the Sunrise Formation. These outcrops are the result of the earliest known Jurassic carbonate ramp produced in the back-arc basin along NE Panthalassa following the extinction event to determine the biotic constituents and timing of local ecological recovery.

The Ferguson Hill Member (Hettangian and Sinemurian) of the Sunrise Formation in the New York Canyon area of west-central Nevada, USA has a lovely counterpart in the Rockies of British Columbia, Canada, explored over three field seasons in the early 2000s before being closed off as a provincial park.

In the Hettangian, post-extinction biosiliceous sedimentation extended to the inner ramp, where an ooid and grapestone shoal marked the outermost extent of a narrow belt of carbonate sedimentation. An early recovery phase in the late Hettangian is characterized by widespread, laterally homogeneous, demosponge-dominated level-bottom sedimentation across the mid- to inner-ramp, in addition to limited trophic tiering (sessile epifaunal suspension-feeding and mobile infaunal deposit-feeding), substantial ramp aggradation, and poor settling conditions for competitive benthic colonizers (e.g., corals, crinoids, infaunal bivalves).

Within 1.6–2 Myr after the extinction (in the early Sinemurian), a late recovery phase is recognized by the appearance of epifaunal grazers (gastropods, echinoids) and suspension feeders (crinoids, solitary scleractinian corals), phototrophic microbialites (oncoids, and possibly photosymbionts within corals), and infaunal deposit or suspension feeders (bivalves).

Although the late recovery faunas included more trophic levels than pre-extinction carbonate ramp habitats, development and progradation of the first Jurassic carbonate ramp still relied heavily on sponge, microbialite, and abiotic mineralization.

AMMONOIDS AND ICHTHYOSAURS: NEVADA PALAEONTOLOGY

Time Slows at Berlin-Ichthyosaur State Park

Around half past five, as the first flush of dawn appears, our group was up and ready to visit a site that has been on my bucket list for some time.

High on the hillside up a long entry road sits the entrance to Berlin-Ichthyosaur State Park in central Nevada. It is a short 2-hour drive from our lodgings through orange-hued grasslands and low sloping hills.

A worn American flag and sun-bleached outbuildings greet you on your way to the outcrops. Away from the hustle and bustle that defines the rest of Nevada this place feels remarkably serene. Your eyes squint against the sun as you search for ammonoids and other marine fossil fauna while your nose tends to the assault from the bracing smell of sagebrush.

This site holds many stories. The interpretive centre displays wonderful marine reptiles, ichthyosaurs in situ, as you might expect from the name of the park — but it also showcases years of history lovingly tended. This stretch of dry golden low hills dappled with the yellow of creosote and desert grasses is an important locality for our understanding of the Carnian-Norian boundary (CNB) in North America.

The area is known worldwide as one of the most important ichthyosaur Fossil-Lagerstätte because of the sheer volume of remarkably well-preserved, fully articulated (all the sweet bones laid out all in a row...) specimens of Shonisaurus popularis.

Rich ammonoid faunas outcrop in the barren hills of the Upper Triassic (Early Norian, Kerri zone), Luning Formation, West Union Canyon, Nevada. They were studied by N. J. Silberling (1959) and provide support for the definition of the Schucherti and Macrolobatus zones of the latest Carnian — which are here overlain by well-preserved faunas of the earliest Norian Kerri Zone. 

The genus Gonionotites, very common in the Tethys and British Columbia, is for the moment, unknown in Nevada. The Upper Carnian faunas are dominated by Tropitidae, while Juvavitidae are conspicuously lacking. 

Middle Triassic Ammonoids

Despite its importance, no further investigations had been done at this site for a good 50 years. That changed in 2010 when Jim Haggart, Mike Orchard and Paul Smith — all local Vancouverites — collaborated on a project that took them down to Nevada to look at the conodonts and ammonoids. They did a bed-by-bed sampling of ammonoids and conodonts in West Union Canyon during October of that year.

October is an ideal time to do fieldwork in this area. There are a few good weeks between screaming hot and frigid cold. It is also tarantula breeding season so keep your eyes peeled. Those sweet little burrows you see are not from rodents but rather largish arachnids. 

The eastern side of the canyon provides the best record of the Macrolobatus Zone, which is represented by several beds yielding ammonoids of the Tropites group, together with Anatropites div. sp. 

Conodont faunas from both these and higher beds are dominated by ornate metapolygnthids that would formerly have been collectively referred to Metapolygnathus primitius, a species long known to straddle the CNB. Within this lower part of the section, they resemble forms that have been separated as Metapolygnathus mersinensis. Slightly higher, forms close to Epigondolella' orchardi and a single Orchardella n. sp. occur. This association can be correlated with the latest Carnian in British Columbia.

Higher in the section, the ammonoid fauna shows a sudden change and is dominated by Tropithisbites. Few tens of metres above, but slightly below the first occurrence of Norian ammonoids Guembelites jandianus and Stikinoceras, two new species of conodonts (Gen et sp. nov. A and B) appear that also occur close to the favoured Carnian/Norian boundary at Black Bear Ridge, British Columbia. Stratigraphically higher collections continue to be dominated by forms close to M. mersinensis and E. orchardi after BC's own Mike Orchard.

The best exposure of the Kerri Zone is on the western side of the West Union Canyon. Ammonoids, dominated by Guembelites and Stikinoceras div. sp., have been collected from several fossil-bearing levels. Conodont faunas replicate those of the east section. The collected ammonoids fit perfectly well with the faunas described by Silberling in 1959, but they differ somewhat from coeval faunas of the Tethys and Canada. 

The ammonoid fauna paints a compelling picture of Tethyan influence with a series of smoking guns. We see an abundance of Tropitidae in the Carnian, a lack of Pterosirenites in the Norian, copious Guembelites, the Tethyan species G. philostrati, the stratigraphic position of G. clavatus and the rare occurrence of Gonionotites. Their hallelujah moment was likely finding an undescribed species of the thin-shelled bivalve Halobia similar to Halobia beyrichi — the clincher that perhaps seals this deal on Tethyan influence. 

I'll take a boo to see what Christopher McRoberts published on the find. A jolly good idea to have him on this expedition as it would have been easy to overlook if the focus remained solely on the conodonts and ammonoids. McRoberts has published on the much-studied Pardonet Formation up in the Willison Lake Area of Northeastern, British Columbia. He knows a thing or two about Upper Triassic Bivalvia and the correlation to coeval faunas elsewhere in the North American Cordillera, and to the Boreal, Panthalassan and Tethyan faunal realms. 

If you fancy a read, they published a paper: "Towards the definition of the Carnian/Norian Boundary: New data on Ammonoids and Conodonts from central Nevada," which you can find in the proceedings of the 21st Canadian Paleontology Conference; by Haggart, J W (ed.); Smith, P L (ed.); Canadian Paleontology Conference Proceedings no. 9, 2011 p. 9-10.

Fig. 1. Location map of Berlin-Ichthyosaur State Park

Marco Balini, James Jenks, Riccardo Martin, Christopher McRoberts, along with Mike Orchard and Norman Siberling, did a bed by bed sampling in 2013 and published on The Carnian/Norian boundary succession at Berlin-Ichthyosaur State Park (Upper Triassic, central Nevada, USA) and published in January 2014 in Paläontologische Zeitschrift 89:399–433. That work is available for download from ResearchGate. The original is in German, but there is a translation available.

After years of reading about the correlation between British Columbia and Nevada, I had the very great pleasure of walking through these same sections in October 2019 with members of the Vancouver Paleontological Society and Vancouver Island Palaeontological Society. It was with that same crew that I'd originally explored fossil sites in the Canadian Rockies in the early 2000s. Those early trips led to paper after paper and the exciting revelations that inspired our Nevada adventure.

If you plan your own adventure, you'll want to keep an eye out for some of the other modern fauna — mountain lions, snakes, lizards, scorpions, wolves, coyotes, foxes, ground squirrels, rabbits, falcons, hawks, eagles, bobcats, sheep, deer and pronghorns.

Figure One: Location map of Berlin-Ichthyosaur State Park. A detailed road log with access information for this locality is provided in Lucas et al. (2007).