ZENAPSIS MORTALITY PLATE

A Devonian fish mortality plate showing all lower shields of Zenaspis podolica (Lankester, 1869) and Stensiopelta pustulata (or Victoraspis longicornualis) from Lower Devonian deposits of Podolia, Ukraine.

Zenaspis is an extinct genus of jawless fish which existed during the early Devonian period. Due to it being jawless, Zenaspis was probably a bottom feeder.

The lovely 420 million-year-old plate you see here is from Podolia or Podilia, a historic region in Eastern Europe, located in the west-central and south-western parts of Ukraine, in northeastern Moldova. Podolia is the only region in Ukraine where Lower Devonian remains of ichthyofauna can be found near the surface.

For the past 150 years, vertebrate fossils have been found in more than 90 localities situated in outcrops along banks of the Dniester River and its northern tributaries, and in sandstone quarries. At present faunal list of Early Devonian agnathans and fishes from Podolia number 72 species, including 8 Thelodonti, 39 Heterostraci, 19 Osteostraci, 4 Placodermi, 1 Acanthodii, and 1 Holocephali (Voichyshyn 2001a, modified).

In Podolia, Lower Devonian redbeds strata (the Old Red Formation or Dniester Series) can be found up to 1800 m thick and range from Lochkovian to Eifelian in age (Narbutas 1984; Drygant 2000, 2003). In the lower part (Ustechko and Khmeleva members of the Dniester Series) they consist of multicoloured, mainly red, fine-grained cross-bedded massive quartz sandstones and siltstones with seams of argillites (Drygant 2000).

We see fossils beds of Zenaspis in the early Devonian of Western Europe. Both Zenaspis pagei and Zenaspis poweri can be found up to 25 centimetres long in Devonian outcrops of Scotland.

Reference: Voichyshyn, V. 2006. New osteostracans from the Lower Devonian terrigenous deposits of Podolia, Ukraine. Acta Palaeontologica Polonica 51 (1): 131–142. Photo care of Fossilero Fisherman.

JELLYFISH: GAGISAMA

These festive lovelies are jellyfish. Jellyfish are found all over the world, from surface waters to our deepest seas — and they are old. They are some of the oldest animals in the fossil record.

Sea jellies and jellyfish are the common names for the medusa-phase or adult phase of certain gelatinous members of the subphylum Medusozoa, a major part of the phylum Cnidaria — more closely related to anemones and corals.

Jellyfish are not fish at all. Jellyfish evolved millions of years before true fish. 

The oldest conulariid scyphozoans — picture an ice-cream cone with fourfold symmetry — appeared between 635 and 577 million years ago in the Neoproterozoic of the Lantian Formation a 150-meter-thick sequence of rocks deposited in southern China. 

Others are found in the youngest Ediacaran rocks of the Tamengo Formation of Brazil, c. 505 mya, through to the Triassic. Cubozoans and hydrozoans appeared in the Cambrian of the Marjum Formation in Utah, USA, c. 540 mya. Like other soft-bodied organisms, ctenophores (comb jellies), sea jellies and jellyfish only produce fossils only under exceptional taphonomic conditions — think rare.

I have seen all sorts of their brethren growing up on the west coast of Canada. I have seen them in tide pools, washed up on the beach and swam amongst thousands of Moon Jellyfish while scuba diving in the Salish Sea. Their movement in the water is marvellous.  

In the Kwak̓wala language of the Kwakiutl or Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, jellyfish are known as ǥaǥisama.

The watercolour ǥaǥisama you see here is a bit of fancy. While I chose blue, purple and pink for these lovelies, they also come in bright yellow, orange and relatively clear — and are often luminescent.

Jellyfish such as comb jellies produce bright flashes to startle a predator, others such as siphonophores can produce a chain of light or release thousands of glowing particles into the water as a mimic of small plankton to confuse the predator.

For most jellyfish bioluminescence is used for defence against predators — and about half of all jellyfish are bioluminescent. Some produce a glowing sticky slime that clings to predators making them vulnerable to other predators. Some jellyfish can release their tentacles as glowing decoys. So you see that there are many strategies for using bioluminescence by jellyfish.

All bioluminescence comes from energy released from a chemical reaction. This is very different from other sources of light, such as from the sun or a light bulb, where the energy comes from heat. In a luminescent reaction, two types of chemicals, called luciferin and luciferase, combine together. The luciferase acts as an enzyme, allowing the luciferin to release energy as it is oxidized. The colour of the light depends on the chemical structures of the chemicals. 

There are more than a dozen known chemical luminescent systems, indicating that bioluminescence evolved independently in different groups of organisms. One type of luciferin is called coelenterazine, found in jellyfish, shrimp, and fish. Dinoflagellates and krill share another class of unique luciferins, while ostracods (firefleas) and some fish have a completely different luciferin. The occurrence of identical luciferins for different types of organisms suggests a dietary source for some groups. Organisms such as bacteria and fireflies have unique luminescent chemistries. In many other groups, the chemistry is still unknown

Some of the most amazing deep-sea jellyfish are the comb jellies, which can get as large as a basketball, and are in some cases so fragile that they are almost impossible to collect intact.

Also spectacular are the siphonophores, some of which can reach several meters in length. Siphonophores deploy many tentacles like a gill net casting for small fish.

METASEQUOIA: A LIVING FOSSIL

Autumn is a wonderful time to explore Vancouver. It is a riot of yellow, orange and green. The fallen debris you crunch through send up wafts of earthy smells that whisper of decomposition, the journey from leaf to soil.

It is a wonderful time to be out and about. I do love the mountain trails but must confess to loving our cultivated gardens for their colour and variety. 

We have some lovely native plants and trees and more than a few exotics at Vancouver's arboreal trifecta — Van Dusen, Queen E Park and UBC Botanical Gardens. One of those exotics, at least exotic to me, is the lovely conifer you see here is Metasequoia glyptostroboides — the dawn redwood. 

Of this long lineage, this is the sole surviving species in the genus Metasequoia and one of three species of conifers known as redwoods. Metasequoia are the smaller cousins of the mighty Giant Sequoia, the most massive trees on Earth. 

As a group, the redwoods are impressive trees and very long-lived. The President, an ancient Giant Sequoia, Sequoiadendron giganteum, and granddaddy to them all has lived for more than 3,200 years. While this tree is named The President, a worthy name, it doesn't really cover the magnitude of this giant by half.   

This tree was a wee seedling making its way in the soils of the Sierra Nevada mountains of California before we invented writing. It had reached full height before any of the Seven Wonders of the Ancient World, those remarkable constructions of classical antiquity, were even an inkling of our budding human achievements. And it has outlasted them all save the Great Pyramid of Giza, the oldest and last of those seven still standing, though the tree has faired better. Giza still stands but the majority of the limestone façade is long gone.

Aside from their good looks (which can really only get you so far), they are resistant to fire and insects through a combined effort of bark over a foot thick, a high tannin content and minimal resin, a genius of evolutionary design. 

While individual Metasequoia live a long time, as a genus they have lived far longer. 

Like Phoenix from the Ashes, the Cretaceous (K-Pg) extinction event that wiped out the dinosaurs, ammonites and more than seventy-five percent of all species on the planet was their curtain call. The void left by that devastation saw the birth of this genus — and they have not changed all that much in the 65 million years since. Modern Metasequoia glyptostroboides looks pretty much identical to their late Cretaceous brethren.

Dawn Redwood Cones with scales paired in opposite rows

They are remarkably similar to and sometimes mistaken for Sequoia at first glance but are easily distinguishable if you look at their size (an obvious visual in a mature tree) or to their needles and cones in younger specimens. 

Metasequoia has paired needles that attach opposite to each other on the compound stem. Sequoia needles are offset and attached alternately. Think of the pattern as jumping versus walking with your two feet moving forward parallel to one another. 

Metasequoia needles are paired as if you were jumping forward, one print beside the other, while Sequoia needles have the one-in-front-of-the-other pattern of walking.

The seed-bearing cones of Metasequoia have a stalk at their base and the scales are arranged in paired opposite rows which you can see quite well in the visual above. Coast redwood cone scales are arranged in a spiral and lack a stalk at their base.

Although the least tall of the redwoods, it grows to an impressive sixty meters (200 feet) in height. It is sometimes called Shui-sa, or water fir by those who live in the secluded mountainous region of China where it was rediscovered.

Fossil Metasequoia, McAbee Fossil Beds

Metasequoia fossils are known from many areas in the Northern Hemisphere and were one of my first fossil finds as a teenager. 

And folk love naming them. More than twenty fossil species have been named over time —  some even identified as the genus Sequoia in error — but for all their collective efforts to beef up this genus there are just three species: Metasequoia foxii, Metasequoia milleri, and Metasequoia occidentalis.

During the Paleocene and Eocene, extensive forests of Metasequoia thrived as far north as Strathcona Fiord on Ellesmere Island and sites on Axel Heiberg Island in Canada's far north around 80° N latitude.

We find lovely examples of Metasequoia occidentalis in the Eocene outcrops at McAbee near Cache Creek, British Columbia, Canada. I shared a photo here of one of those specimens. Once this piece dries out a bit, I will take a dental pick to it to reveal some of the teaser fossils peeking out.

The McAbee Fossil Beds are known for their incredible abundance, diversity and quality of fossils including lovely plant, insect and fish species that lived in an old lake bed setting. While the Metasequoia and other fossils found here are 52-53 million years old, the genus is much older. It is quite remarkable that both their fossil and extant lineage were discovered in just a few years of one another. 

Metasequoia was first described as a new genus from a fossil specimen found in 1939 and published by Japanese paleobotanist Shigeru Miki in 1941. Remarkably, the living version of this new genus was discovered later that same year. 

Professor Zhan Wang, an official from the Bureau of Forest Research was recovering from malaria at an old school chum's home in central China. His friend told him of a stand of trees discovered in the winter of 1941 by Chinese botanist Toh Gan (干铎). The trees were not far away from where they were staying and Gan's winter visit meant he did not collect any specimen as the trees had lost their leaves. 

The locals called the trees Shui-sa, or water fir. As trees go, they were reportedly quite impressive with some growing as much as sixty feet tall. Wang was excited by the possibility of finding a new species and asked his friend to describe the trees and their needles in detail. Emboldened by the tale, Wang set off through the remote mountains to search for his mysterious trees and found them deep in the heart of  Modaoxi (磨刀溪; now renamed Moudao (谋道), in Lichuan County, in the central China province of Hubei. He found the trees and was able to collect living specimens but initially thought they were from Glyptostrobus pensilis (水松). 

A few years later, Wang showed the trees to botanist Wan-Chun Cheng and learned that these were not the leaves of s Glyptostrobus pensilis (水松 ) but belonged to a new species. 

While the find was exciting, it was overshadowed by China's ongoing conflict with the Japanese that was continuing to escalate. With war at hand, Wang's research funding and science focus needed to be set aside for another two years as he fled the bombing of Beijing. 

When you live in a world without war on home soil it is easy to forget the realities for those who grew up in it. 

Zhan Wang and his family lived to witness the 1931 invasion of Manchuria, then the 1937 clash between Chinese and Japanese troops at the Marco Polo Bridge, just outside Beijing. 

That clash sparked an all-out war that would grow in ferocity to become World War II. 

Within a year, the Chinese military situation was dire. Most of eastern China lay in Japanese hands: Shanghai, Nanjing, Beijing, Wuhan. As the Japanese advanced, they left a devastated population in their path where atrocity after atrocity was the norm. Many outside observers assumed that China could not hold out, and the most likely scenario was a Japanese victory over China.

Yet the Chinese hung on, and after the horrors of Pearl Harbor, the war became genuinely global. The western Allies and China were now united in their war against Japan, a conflict that would finally end on September 2, 1945, after Allied naval forces blockaded Japan and subjected the island nation to intensive bombing, including the utter devastation that was the Enola Gay's atomic payload over Hiroshima. 

With World War II behind them, the Chinese researchers were able to re-focus their energies on the sciences. Sadly, Wang was not able to join them. Instead, two of his colleagues, Wan Chun Cheng and Hu Hsen Hsu, the director of Fan Memorial Institute of Biology would continue the work. Wan-Chun Cheng sent specimens to Hu Hsen Hsu and upon examination realised they were the living version of the trees Miki had published upon in 1941. 

Hu and Cheng published a paper describing a new living species of Metasequoia in May 1948 in the Bulletin of Fan Memorial Institute of Biology.

That same year, Arnold Arboretum of Harvard University sent an expedition to collect seeds and, soon after, seedling trees were distributed to various universities and arboreta worldwide. 

Today, Metasequoia grow around the globe. When I see them, I think of Wang and all he went through. He survived the conflict and went on to teach other bright, young minds about the bountiful flora in China. I think of Wan Chun Cheng collaborating with Hu Hsen Hsu in a time of war and of Hu keeping up to date on scientific research, even published works from colleagues from countries with whom his country was at war. Deep in my belly, I ache for the huge cost to science, research and all the species impacted on the planet from our human conflicts. Each year in April, I plant more Metasequoia to celebrate Earth Day and all that means for every living thing on this big blue orb.  

References: 

• https://web.stanford.edu/group/humbioresearch/cgi-bin/wordpress/?p=297
• https://humboldtredwoods.org/redwoods

TETRAPODS & FIRST NATION FOSSILS

Elpistostege watsoni

In the late 1930s, our understanding of the transition of fish to tetrapods — and the eventual jump to modern vertebrates — took an unexpected leap forward. The evolutionary a'ha came from a single partial fossil skull found on the shores of a riverbank in Eastern Canada. 

Meet the Stegocephalian, Elpistostege watsoni, an extinct genus of finned tetrapodomorphs that lived during the Late Givetian to Early Frasnian of the Late Devonian — 382 million years ago. 

Elpistostege watsoni — perhaps the sister taxon of all other tetrapods — was first described in 1938 by British palaeontologist and elected Fellow of the Royal Society of London, Thomas Stanley Westoll. Westoll's research interests were wide-ranging. He was a vertebrate palaeontologist and geologist best known for his innovative work on Palaeozoic fishes and their relationships with tetrapods. 

As a specialist in early fish, Westoll was asked to interpret that single partial skull roof discovered at the Escuminac Formation in Quebec, Canada. His findings and subsequent publication named Elpistostege watsoni and helped us to better understand the evolution of fishes to tetrapods — four-limbed vertebrates — one of the most important transformations in vertebrate evolution. 

Hypotheses of tetrapod origins rely heavily on the anatomy of but a few tetrapod-like fish fossils from the Middle and Late Devonian, 393–359 million years ago. These taxa — known as elpistostegalians — include Panderichthys, Elpistostege and Tiktaalik — none of which had yet to reveal the complete skeletal anatomy of the pectoral fin. 

Elpistostege watsoni

None until 2010 that is, when a complete 1.57-metre-long articulated specimen was found and described by Richard Cloutier et al. in 2020. 

The specimen helped us to understand the origin of the vertebrate hand. Stripped from its encasing stone, it revealed a set of paired fins of Elpistostege containing bones homologous to the phalanges (finger bones) of modern tetrapods and is the most basal tetrapodomorph known to possess them. 

Once the phalanges were uncovered, prep work began on the fins. The fins were covered in wee scales and lepidotrichia (fin rays). The work was tiresome, taking more than 2,700 hours of preparation but the results were thrilling. 

Origin of the Vertebrate Hand

We could now clearly see that the skeleton of the pectoral fin has four proximodistal rows of radials — two of which include branched carpals — as well as two distal rows organized as digits and putative digits. 

Despite this skeletal pattern — which represents the most tetrapod-like arrangement of bones found in a pectoral fin to date blurring the line between fish and land vertebrates — the fin retained lepidotrichia (those wee fin rays) distal to the radials. 

This arrangement confirmed an age-old question — showing us for the first time that the origin of phalanges preceded the loss of fin rays, not the other way around.

E. watsoni is very closely related to Tiktaalik roseae found in 2004 in the Canadian Arctic — a tetrapodomorpha species also known as a Choanata. These were advanced forms transitional between fish and the early labyrinthodonts playfully referred to as fishapods — half-fish, half-tetrapod in appearance and limb morphology. 

Up to that point, the relationship of limbed vertebrates (tetrapods) to lobe-finned fish (sarcopterygians) was well known, but the origin of major tetrapod features remained obscure for lack of fossils that document the sequence of evolutionary changes — until Tiktaalik. While Tiktaalik is technically a fish, this fellow is as far from fish-like you can be and still be a card-carrying member of the group. 

Tiktaalik roseae

Complete with scales and gills, this proto-fish lacked the conical head we see in modern fish but had a rather flattened triangular head more like that of a crocodile. 

Tiktaalik had scales on its back and fins with fin webbing but like early land-living animals, it had a distinctive flat head and neck. He was a brawny brute. The shape of his skull and shoulder look part fish and part amphibian.

The watershed moment came as Tiktaalik was prepped. Inside Tiktaalik's fins, we find bones that correspond to the upper arm, forearm and even parts of the wrist — all inside a fin with webbing — remarkable! 

Its fins have thin ray bones for paddling like most fish, but with brawny interior bones that gave Tiktaalik the ability to prop itself up, using his limbs for support. I picture him propped up on one paddle saying, "how you doing?" 

Six years after Tiktaalik was discovered by Neil Shubin and team in the ice-covered tundra of the Canadian Arctic on southern Ellesmere Island, a team working the outcrops at Miguasha on the Gaspé Peninsula discovered the only fully specimen of E. watsoni found to date — greatly increasing our knowledge of this finned tantalizingly transitional tetrapodomorph. 

E. watsoni fossils are rare — this was the fourth specimen collected in over 130 years of hunting. Charmingly, the specimen was right on our doorstop — extracted but a few feet away from the main stairs descending onto the beach of Miguasha National Park. 

L'nu Mi’gmaq First Nations of the Gespe’gewa’gi Region

Miguasha is nestled in the Gaspésie or Gespe’gewa’gi region of Canada — home to the Mi’gmaq First Nations who self-refer as L’nu or Lnu. The word Mi’gmaq or Mi’kmaq means the family or my allies/friends in Mi'kmaw, their native tongue (and soon to be Nova Scotia's provincial first language). They are the people of the sea and the original inhabitants of Atlantic Canada having lived here for more than 10,000 years. 

The L'nu were the first First Nation people to establish contact and trade with European explorers in the 16th and 17th centuries — and perhaps the Norse as early as the turn of the Millenium. Sailing vessels filled with French, British, Scottish, Irish and others arrived one by one to lay claim to the region — settling and fighting over the land. As each group rolled out their machinations of discovery, tensions turned to an all-out war with the British and French going head to head. I'll spare you the sordid details but for everyone caught in the crossfire, it went poorly.

North America Map 1775 (Click to Enlarge)

Cut to 1760, the British tipped the balance with their win at the Battle of the Restigouche, the last naval battle between France and England for possession of the North American continent — Turtle Island. 

The bittersweet British victory sparked the American War of Independence. 

For the next twenty years, the L'nu would witness and become embroiled in yet another war for these lands, their lands — first as bystanders, then as American allies, then intimidated into submission by the British Royal Navy with a show of force by way of a thirty-four gun man-of-war, encouraging L'nu compliance — finally culminating in an end to the hostilities with the 1783 Treaty of Paris. 

The peace accord held no provisions for the L'nu, Métis and First Nations impacted. None of these newcomers was Mi'kmaq — neither friends nor allies.

It was to this area some sixty years later that the newly formed Geological Survey of Canada (GSC) began exploring and mapping the newly formed United Province of Canada. Geologists in the New Brunswick Geology Branch traipsed through the rugged countryside that would become a Canadian province in 1867. 

It was on one of these expeditions that the Miguasha fossil outcrops were discovered. They, too, would transform in time to become Miguasha National Park or Parc de Miguasha, but at first, they were simply the promising sedimentary exposures on the hillside across the water —  a treasure trove of  Late Devonian fauna waiting to be discovered.

In the summer of 1842, Abraham Gesner, New Brunswick’s first Provincial Geologist, crossed the northern part of the region exploring for coal. Well, mostly looking for coal. Gesner also had a keen eye for fossils and his trip to the Gaspé Peninsula came fast on the heels of a jaunt along the rocky beaches of Chignecto Bay at the head of the Bay of Fundy and home to the standing fossil trees of the Joggins Fossil Cliffs. 

Passionate about geology and chemistry, he is perhaps most famous for his invention of the process to distil the combustible hydrocarbon kerosene from coal oil — a subject on which his long walks exploring a budding Canada gave him a great deal of time to consider. We have Gesner to thank for the modern petroleum industry. He filed many patents for clever ways to distil the soft tar-like coal or bitumen still in use today.

He was skilled in a broad range of scientific disciplines — being a geologist, palaeontologist, physician, chemist, anatomist and naturalist — a brass tacks geek to his core. Gesner explored the coal exposures and fossil outcrops across the famed area that witnessed the region become part of England and not France — and no longer L'nu.

Following the Restigouche River in New Brunswick through the Dalhousie region, Gesner navigated through the estuary to reach the southern coast of the Gaspé Peninsula into what would become the southeastern coast of Quebec to get a better look at the cliffs across the water. He was the first geologist to lay eyes on the Escuminac Formation and its fossils.

In his 1843 report to the Geologic Survey, he wrote, “...I found the shore lined with a coarse conglomerate. Farther eastward the rocks are light blue sandstones and shales, containing the remains of vegetables. (...) In these sandstone and shales, I found the remains of fish and a small species of tortoise with fossil foot-marks.”

We now know that this little tortoise was the famous Bothriolepis, an antiarch placoderm fish. It was also the first formal mention of the Miguasha fauna in our scientific literature. Despite the circulation of his report, Gesner’s discovery was all but ignored — the cliffs and their fossil bounty abandoned for decades to come. Geologists like Ells, Foord and Weston, and the research of Whiteaves and Dawson, would eventually follow in Gesner's footsteps.

North America Map 1866 (Click to Enlarge)

Over the past 180 years, this Devonian site has yielded a wonderfully diverse aquatic assemblage from the Age of Fishes — five of the six fossil fish groups associated with the Devonian including exceptionally well-preserved fossil specimens of the lobe-finned fishes. 

This is exciting as it is the lobe-finned fishes — the sarcopterygians — that gave rise to the first four-legged, air-breathing terrestrial vertebrates – the tetrapods. 

Fossil specimens from Miguasha include twenty species of lower vertebrates — anaspids, osteostra-cans, placoderms, acanthodians, actinopterygians and sarcopterygians — plus a limited invertebrate assemblage, along with terrestrial plants, scorpions and millipedes.

Originally interpreted as a freshwater lacustrine environment, recent paleontological, taphonomic, sedimentological and geochemical evidence corroborates a brackish estuarine setting — and definitely not the deep waters of the sea. This is important because the species that gave rise to our land-living animals began life in shallow streams and lakes. It tells us a bit about how our dear Elpistostege watsoni liked to live — preferring to lollygag in cool river waters where seawater mixed with fresh. Not fully freshwater, but a wee bit of salinity to add flavour.  

• Photos: Elpistostege watsoni (Westoll, 1938 ), Upper Devonian (Frasnian), Escuminac formation, Parc de Miguasha, Baie des Chaleurs, Gaspé, Québec, Canada. John Fam, VanPS
• Origin of the Vertebrate Hand Illustration, https://www.nature.com/articles/s41586-020-2100-8
• Tiktaalik Illustration: By Obsidian Soul - Own work, CC BY 4.0, https://commons.wikimedia.org/w/index.php?curid=47401797

References & further reading:

• From Water to Land: https://www.miguasha.ca/mig-en/the_first_discoveries.php
• UNESCO Miguasha National Park: https://whc.unesco.org/en/list/686/
• Office of L'nu Affairs: https://novascotia.ca/abor/aboriginal-people/
• Cloutier, R., Clement, A.M., Lee, M.S.Y. et al. Elpistostege and the origin of the vertebrate hand. Nature 579, 549–554 (2020). https://doi.org/10.1038/s41586-020-2100-8
• Daeschler, E. B., Shubin, N. H. & Jenkins, F. A. Jr. A Devonian tetrapod-like fish and the evolution of the tetrapod body plan. Nature 440, 757–763 (2006).
• Shubin, Neil. Your Inner Fish: A Journey into the 3.5 Billion History of the Human Body.
• Evidence for European presence in the Americas in AD 1021: https://www.nature.com/articles/s41586-021-03972-8

EIFELIAN PARALEJURUS

This bronzed beauty is the Middle Devonian, Eifelian (~395 mya) trilobite, Paralejurus rehamnanus (Alberti, 1970) from outcrops near Issoumour, Alnif, Morocco in North Africa. 

It was the colour of this amazing trilobite that captured the eye of David Appleton in whose collection it now resides. He is an avid collector and coming into his own as a macro photographer. I have shared three of his delightful photos for you here.

It initially thought that the gold we see here was added during prep, particularly considering the colouration of the matrix, but macro views of the surface show mineralization and the veins running right through the specimen into the matrix. There is certainly some repairs but that is common in the restoration of these specimens. Many of the trilobites I have seen from Morocco have bronze on black colouring but not usually this pronounced. Even so, there is a tremendous amount of fine anatomy to explore and enjoy in this wonderfully preserved specimen.  

Paralejurus is a genus of trilobite in the phylum Arthropoda from the Late Silurian to the Middle Devonian of Africa and Europe. These lovelies grew to be up to nine centimetres, though the fellow you see here is a wee bit over half that size at 5.3 cm. 

Paralejurus specimens are very pleasing to the eye with their long, oval outline and arched exoskeletons. 

Their cephalon or head is a domed half circle with a smooth surface.  The large facet eyes have very pleasing crescent-shaped lids. You can see this rather well in the first of the photos here. The detail is quite remarkable.

As you move down from his head towards the body, there is an almost inconspicuous occipital bone behind the glabella in the transition to his burnt bronze thorax.

The body or thorax has ten narrow segments with a clearly arched and broad axial lobe or rhachis. The pygidium is broad, smooth and strongly fused in contrast to the genus Scutellum in the family Styginidae, which has a pygidium with very attractive distinct furrows that I liken to the look of icing ridges on something sweet — though that may just be me and my sweet tooth talking. In Paralejurus, they look distinctly fused — or able to fuse — to add posterior protection against predators with both the look and function of Roman armour.

In Paralejurus, the axillary lobe is rounded off and arched upwards. It is here that twelve to fourteen fine furrows extend radially to complete the poetry of his body design. 

Trilobites were amongst the earliest fossils with hard skeletons and they come in many beautiful forms. While they are extinct today, they were the dominant life form at the beginning of the Cambrian. 

As a whole, they were amongst some of the most successful of all early animals — thriving and diversifying in our ancient oceans for almost 300 million years. The last of their brethren disappeared at the end of the Permian — 252 million years ago. Now, we enjoy their beauty and the scientific mysteries they reveal about our Earth's ancient history.

Photos and collection of the deeply awesome David Appleton. Specimen: 5.3 cm. 

MIDDLE TRIASSIC PAPER CLAMS & AMMONOIDS

Paper clams or "flat clams" were widespread in the Triassic. They often dominate the rocks in which they are found, as in these specimens from the Daonella dubia zone of the West Humboldt Range and the Desatoya Range of Nevada where they are associated with Ceratities trinodosus, Nevadites Whitney, Daonella lindstomi, D. moussoni and other species.

This designation was coined by J. P. Smith in the early 1900s for specific localities in the Humboldt Mountain Range. Because of their widespread distribution and very high species turnover rates, they make for excellent biochronological macrofossils, helping us to correlate biological events through time.

We see the "cousins" of these Nevada specimens up in  Pine Pass near Chetwynd, British Columbia.

Pine Pass is part of the Pardonet Formation. Just a short hike from the road we were able to easily find the abundant outcroppings of the paper clam Monotis subcircularis, perfectly preserved and cemented in this strata from the Late Triassic.

THE ART OF THE PORTAGE: BOWRON LAKES

A cool morning breeze keeps the mosquitoes down as we pack our kayaks and gear for today’s paddling journey. 

It is day four of our holiday, with two days driving up from Vancouver to Cache Creek, past the Eocene insect and plant site at McAbee, the well-bedded Permian limestone near Marble Canyon and onto Bowron Provincial Park, a geologic gem near the gold rush town of Barkerville. 

The initial draw for me, given that collecting in a provincial park is forbidden and all collecting close at hand outside the park appears to amount to a handful of crushed crinoid bits and a few conodonts, was the gorgeous natural scenery and a broad range of species extant. 

It was also the proposition of padding the Bowron Canoe Circuit, a 149,207-hectare geologic wonderland, where a fortuitous combination of plate tectonics and glacial erosion have carved an unusual 116-kilometre near-continuous rectangular circuit of lakes, streams and rivers bound on all sides by snowcapped mountains. 

From all descriptions, something like heaven. 

The east and south sides of the route are bound by the imposing white peaks of the Cariboo Mountains, the northern boundary of the Interior wet belt, rising up across the Rocky Mountain Trench, and the Isaac Formation, the oldest of seven formations that make up the Cariboo Group (Struik, 1988). 

Some 270 million-plus years ago, had one wanted to buy waterfront property in what is now British Columbia, you’d be looking somewhere between Prince George and the Alberta border. The rest of the province had yet to arrive but would be made up of over twenty major terranes from around the Pacific. 

The rock that would eventually become the Cariboo Mountains and form the lakes and valleys of Bowron was far out in the Pacific Ocean, down near the equator. 

With tectonic shifting, these rocks drifted north-eastward, riding their continental plate, until they collided with and joined the Cordillera in what is now British Columbia. Continued pressure and volcanic activity helped create the tremendous slopes of the Cariboo Range we see today with repeated bouts of glaciation during the Pleistocene carving their final shape. 

We brace our way into a headwind along the east side of the fjord-like Isaac Lake. Paddling in time to the wind, I soak up the view of this vast, deep green, ocean-like expanse that runs L-shape for nearly 38 kilometres, forming nearly half of the total circuit. 

The rock we paddle past is primarily calcareous phyllite, limestone and quartzite, typical of the type locality for this group and considered upper Proterozoic (Young, 1969), the time in our geologic history between the first algae and the first multicellular animals.

It is striking how much this lake fits exactly how you might picture pristine wilderness paddling in your mind’s eye. No powerboats, no city hum, just pure silence, broken only by the sound of my paddle pulling through the water and the occasional burst of glee from one of the park’s many songbirds. 

We have chosen kayaks over the more-popular canoes for this journey, as I got to experience my first taste of the handling capabilities of a canoe last year in Valhalla Provincial Park. The raised sides acted like sails and kept us off course in all but the lightest conditions. 

This year, Philip Torrens, Leanne Sylvest, and I were making our trek in low profile, Kevlar style. 

One single & one double kayak would be our faithful companions and mode of transport. They would also be briefly conscripted into service as a bear shield later in the trip. Versatile those kayaks. 

The area is home to a variety of plant life. Large sections of the forest floor are carpeted in the green and white of dogwood, a prolific ground cover we are lucky enough to see in full bloom. Moss, mushrooms and small wild grown on every available surface. 

Yellow Lilies line pathways and float in the cold, clear lake water. Somewhere I read a suggestion to bring a bathing suit to the park, but at the moment, I cannot imagine lowering anything more than my paddle into these icy waters. To reach the west side of the paddling route, we must first face several kilometres portaging muddy trails to meet up with the Isaac River and then paddle rapids to grade two. 

At the launch site, we meet up with two fellow kayakers, Adele and Mary of Victoria, and take advantage of their preceding us to watch the path they choose through the rapids. It has been raining in the area for forty plus days, so the water they run is high and fast. Hot on their heels, our short, thrilling ride along the Isaac River, is a flurry of paddle spray and playing around amid all the stumps, silt and conglomerate. 

The accommodation gods smile kindly on us as we are pushed out from Isaac River and settle into McLeary Lake. An old trapper cabin built by local Freddie Becker back in the 1930s, sits vacant and inviting, providing a welcome place to hang our hats and dry out. 

From here we can see several moose, large, lumbering, peaceful animals, the largest members of the deer family, feeding on the grass-like sedge on the far shore. 

The next morning, we paddle leisurely down the slower, silt-laden Cariboo River, avoiding the occasional deadhead, and make our way into the milky, glacier-fed Lanezi Lake. Like most mountainous areas, Bowron makes its own weather system and it appears you get everything in a 24-hour period. 

In fact, whatever weather you are enjoying seems to change 40 minutes later; good for rain, bad for sun. Wisps of cloud that seemed light and airy only hours early have become dark. Careful to hug the shore, we are ready for a quick escape from lightning as thundershowers break. 

Paddling in the rain, I notice bits of mica in the water, playing in the light and the rock change here to greywacke, argillite, phyllite and schist. Past Lanezi, we continue onto Sandy Lake, where old-growth cedars line the south-facing slopes to our left and grey limestone, shale and dolostone line the shore. Mottled in with the rock, we sneak up on very convincing stumps posing as large mammals. 

Picking up the Cariboo River again, we follow it as it flows into Babcock Lake, an area edged with Lower Cambrian limestone, shale and argillite. At the time these rocks were laid down, the Earth was seeing our earliest relatives, the first chordates entering the geologic scene. 

Mamma Moose, Baby Moose... Grizzly!

As we reach the end of Babcock Lake and prepare for our next portage I get out my camera to take advantage of the angle of the sun and the eroded rounded hilltops of the Quesnel Highlands that stand as a backdrop.

Leanne remarks that she can see a moose a little ways off and that it appeared to be heading our way. 

Yes, heading our way quickly with her calf in tow. I lift my lens to immortalize the moment and we three realized they were headed our way in double time because they were being chased by a grizzly bear — at top speed. 

A full-grown moose can run up to fifty-five kilometres per hour, more or less neck and neck with the speed of a Grizzly. They are also strong swimmers. Had she been alone, Mamma moose would likely have tried to outswim the bear. 

From where we stand we can see the water turned to white foam at their feet as they fly towards us. 

We freeze bear spray in hand. 

In seconds the three were upon us. Mamma moose, using home-field advantage, runs straight for us and just reaching our boats, turned 90 degrees, bolting for the woods, baby moose fast on her heels. 

The Grizzly, caught up in the froth of running and thrill of the kill, doesn’t notice the deke, hits the brakes at the boats and stands up, confused. Her eyes give her away. 

This was not what she had planned and the whole moose-suddenly-transformed-into-human thing is giving her pause. Her head tilts back as she gets a good smell of us. 

Suddenly, a crack in the woods catches her attention. Her head snaps around and she drops back on all fours, beginning her chase anew. Somewhere there is a terrified mother moose and calf hoping the distance gained is enough to keep them from being lunch. Both moose got away with the unwitting distraction we provided. 

The Lakes are at an elevation of over 900 m (3000 ft) and both grizzly and black bear sightings are common. Both bear families descend from a common ancestor, Ursavus, a bear-dog the size of a raccoon who lived more than 20 million years ago. Seems an implausible lineage having just met one of the larger descendants.

While we had grumbled only hours earlier about how tired we were feeling, we now feel quite motivated and do the next two portages and lakes in good time. Aside from the gripping fear that another bear encounter is imminent, we enjoy the park-like setting, careful to scan the stands of birch trees for dark shapes now posing as stumps. Fortunately, the only wildlife we see are a few wily chipmunks, various reticent warblers and some equally shy spruce grouse. 

The wind favours us now as we paddle Skoi and Spectacle Lake, even giving us a chance to use the sails we’ve rigged to add an extra knot of oomph to our efforts. Reaching the golden land of safety-in-numbers, we leap from our kayaks, happy to see the smiling faces of Mary and Adele. Making it here is doubly thrilling because it means I’m sleeping indoors tonight and I can tell the bear story with adrenaline still pumping through my veins. Tonight is all about camaraderie and the warmth of a campfire. 

Gobbling down Philip’s famous pizza, Leanne impresses everyone further by telling of his adventures in the arctic and surviving a polar bear attack. This is our first starlit night without rain, a luxury everyone comments on, but quietly, not wanting to jinx it. We share a good laugh at the expense of the local common loons — both Homeo sapien sapien and Gavia immer. 

Common Loon / Gavia immer

The marshy areas of the circuit provide a wonderful habitat for the regions many birds including a host of sleek, almost regal black and white common loons. 

Their cool demeanour by day is reduced to surprisingly loud, maniacal hoots and yelps with undignified flapping and flailing by night. 

It seems hardly possible that these awful noises could be coming from the same birds and that this has been going for nearly 65 million years, since the end of the age of dinosaurs, as loons are one of the oldest bird families in the fossil record. 

A guitar is pulled out to liven the quiet night while small offerings, sacred and scare this late in our journey, are passed around. Tonight is a celebration that we have all, both separately and together, made our way around this immense mountain-edged circuit.

Know before you go

BC Parks Bowron Lake Circuit Link: https://bcparks.ca/reserve/bowron-lake/

• Reservations may be booked up to 2 days prior to the departure date.
• All canoe circuit users must attend a mandatory orientation session (9:00 am or 12:00 noon) and visitors must report to the registration centre by the specified time.
• Visitors who have not attended their mandatory orientation session are considered no-shows. No-shows’ reservations will be given to first-come, first-served visitors. No refunds are granted for no-shows.
• Bowron Lake canoe circuit paddlers are responsible for bringing or renting their own equipment (personal flotation devices, paddles, vessels etc). Reservation transactions only include your reservation charges and user (camping) fees and do not include equipment rental. Visitors must familiarize themselves with the mandatory equipment required to paddle the circuit.
• Reservations are not transferable. Any reservation owners or holders found to be transferring or selling their reservations to another party, risk the chance that their reservation may be cancelled without a refund. If a customer can no longer use their reservation, they are encouraged to cancel their reservation so that any unused User Fees that are not forfeited as a penalty, can be automatically refunded to the credit card that was used to make the original reservation. Only the person whose name is on the reservation has access to change or cancel a reservation.
• At least one of the reservation holders ("Occupant" or "Optional Authorized Person") must be present upon arrival and during the stay. Identification may be required to keep the reservation valid.
• Reservations for the Bowron Lake canoe circuit are based on a non-refundable per vessel charge of $18.00 (plus tax) in addition to user fees ($60.00 per person for the full canoe circuit and $30.00 per person for the West Side). There is a limit of three (3) people per vessel.
• Accepted payment types include Visa, Mastercard, American Express, Visa Debit, and Mastercard Debit.
• Changes to a reservation can be made for a charge of $6.00 (plus tax), provided that space is available. No changes are permitted 28 days or less before the departure date.
• Cancellations are subject to a $6.00 (plus tax) per cancellation charge.
• Refunds – If a trip is cancelled more than 28 days before the departure date, customers receive a full refund less cancellation and non-refundable reservation charges. If cancellations are made with 28 or less days notice, no refunds apply. Transaction (reservation, change, cancellation) charges are non-refundable.
• Visitors who have not attended the mandatory orientation session they registered for are considered no-shows. No-shows' reservations will be given to first-come, first-served visitors. No refunds are granted for no-shows.

ALCIDS AUKS

Puffins are any of three small species of alcids or auks in the bird genus Fratercula with a brightly coloured beak during the breeding season.

Their sexy orange beaks shift from a dull grey to bright orange when it is time to attract a mate. While not strictly monogamous, most Puffins choose the same mate year upon year producing adorable chicks or pufflings (awe) from their mating efforts.

Female Puffins produce one single white egg which the parents take turns to incubate over a course of about six weeks. Their dutiful parents share the honour of feeding the wee pufflings five to eight times a day until the chick is ready to fly. Towards the end of July, the fledgeling Puffins begin to venture from the safety of their parents and dry land. Once they take to the seas, mom and dad are released from duty and the newest members of the colony are left to hunt and survive on their own.

These are pelagic seabirds that feed primarily by diving in the water. They breed in large colonies on coastal cliffs or offshore islands, nesting in crevices among rocks or in burrows in the soil. Two species, the tufted puffin and horned puffin are found in the North Pacific Ocean, while the Atlantic puffin is found in the North Atlantic Ocean. This lovely fellow, with his distinctive colouring, is an Atlantic Puffin or "Sea Parrot" from Skomer Island near Pembrokeshire in the southwest of Wales. Wales is bordered by Camarthenshire to the east and Ceredigion to the northeast with the sea bordering everything else. It is a fine place to do some birding if it's seabirds you're after.

These Atlantic Puffins are one of the most famous of all the seabirds and form the largest colony in Southern Britain. They live about 25 years making a living in our cold seas dining on herring, hake and sand eels. Some have been known to live to almost 40 years of age. They are good little swimmers as you might expect, but surprisingly they are great flyers, too! They are hindered by short wings, which makes flight challenging but still possible with effort. Once they get some speed on board, they can fly up to 88 km an hour.

The oldest alcid fossil is Hydrotherikornis from Oregon dating to the Late Eocene while fossils of Aethia and Uria go back to the Late Miocene. Molecular clocks have been used to suggest an origin in the Pacific in the Paleocene. Fossils from North Carolina were originally thought to have been of two Fratercula species but were later reassigned to one Fratercula, the tufted puffin, and a Cerorhinca species. Another extinct species, Dow's puffin, Fratercula dowi,  was found on the Channel Islands of California until the Late Pleistocene or early Holocene.

The Fraterculini are thought to have originated in the Pacific primarily because of their greater diversity in the region. There is only one extant species in the Atlantic, compared to two in the Pacific. The Fraterculini fossil record in the Pacific extends at least as far back as the middle Miocene, with three fossil species of Cerorhinca, and material tentatively referred to that genus, in the middle Miocene to late Pliocene of southern California and northern Mexico.

Although there no records from the Miocene in the Atlantic, a re-examination of the North Carolina material indicated that the diversity of puffins in the early Pliocene was as great in the Atlantic as it is in the Pacific today. This diversity was achieved through influxes of puffins from the Pacific; the later loss of species was due to major oceanographic changes in the late Pliocene due to closure of the Panamanian Seaway and the onset of severe glacial cycles in the North Atlantic.

TETRALOPHODON

Tetralophodon ("four-ridged tooth") is an extinct proboscidean genus belonging to the superfamily Elephantoidea. 

Their fossils have been found from the late Miocene to the Middle Pliocene of Europe, Asia, and Africa, which means many of our ancestors would have had the opportunity to see and hunt them. Lucky for them, these large elephant-like animals preferred fruits and vegetables. 

Like the gomphotheres, to which it was not closely related, Tetralophodon had an impressive four tusks and a trunk. They were big and on size with our modern elephants. You and I would come up to their hip bone on average. They were about 2.58–3.45 m (8.5–11.3 ft) tall at the shoulder and up to 10 tonnes in weight, larger than the size of the present Asian elephant, with a long trunk and incisors ranging up to 2 m (6.6 ft) long. 

FOSSILS, LIMESTONE & SALT: HALLSTATT

Hallstatt Salt Mines, Austria / Permian Salt Diapir

The Hallstatt Limestone is the world's richest Triassic ammonite unit, yielding specimens of more than 500 ammonite species.

Along with diversified cephalopod fauna  — orthoceratids, nautiloids, ammonoids — we also see gastropods, bivalves, especially the late Triassic pteriid bivalve Halobia (the halobiids), brachiopods, crinoids and a few corals. We also see a lovely selection of microfauna represented. 

For microfauna, we see conodonts, foraminifera, sponge spicules, radiolaria, floating crinoids and holothurian sclerites —  polyp-like, soft-bodied invertebrate echinozoans often referred to as sea cucumbers because of their similarities in size, elongate shape, and tough skin over a soft interior. 

Franz von Hauer’s exhaustive 1846 tome describing Hallstatt ammonites inspired renowned Austrian geologist Eduard Suess’s detailed study of the area’s Mesozoic history. That work was instrumental in Suess being the first person to recognize the former existence of the Tethys Sea, which he named in 1893 after the sister of Oceanus, the Greek god of the ocean. As part of the Northern Limestone Alps, the Dachstein rock mass, or Hoher Dachstein, is one of the large karstic mountains of Austria and the second-highest mountain in the Northern Limestone Alps. It borders Upper Austria and Styria in central Austria and is the highest point in each of those states.

Parts of the massif also lie in the state of Salzburg, leading to the mountain being referred to as the Drei-Länder-Berg or three-state mountain. Seen from the north, the Dachstein massif is dominated by the glaciers with the rocky summits rising beyond them. By contrast, to the south, the mountain drops almost vertically to the valley floor. The karst limestones and dolomites were deposited in our Mesozoic seas. The geology of the Dachstein massif is dominated by the Dachstein-Kalk Formation — the Dachstein limestone — which dates back to the Triassic.

Hallstatt and the Hallstatt Sea, Austria

There were several phases of mountain building in this part of the world pushing the limestone deposits 3,000 metres above current sea level. The rock strata were originally deposited horizontally, then shifted, broken up and reshaped by the erosive forces of ice ages and erosion.

The Hallstatt mine exploits a Permian salt diapir that makes up some of this area’s oldest rock. 

The salt accumulated by evaporation in the newly opened, and hence shallow, Hallstatt-Meliata Ocean. This was one of several small ocean basins that formed in what is now Europe during the late Paleozoic and early Mesozoic when the world’s landmasses were welded together to form the supercontinent Pangea. 

Pangea was shaped like a crescent moon that cradled the famous Tethys Sea. Subduction of Tethyian oceanic crust caused several slivers of continental crust to separate from Pangea, forming new “back-arc basins” (small oceans formed by rifting that is associated with nearby subduction) between the supercontinent and the newly rifted ribbon continents.

The Hallstatt-Meliata Ocean was one such back-arc basin. As it continued to expand and deepen during the Triassic, evaporation ceased and reefs flourished; thick limestone deposits accumulated atop the salt. When the Hallstatt-Meliata Ocean closed in the Late Jurassic, the compression squeezed the low-density salt into a diapir that rose buoyantly, injecting itself into the Triassic limestones above.

The Hallstatt salt diapir and its overlying limestone cap came to rest in their present position in the northern Austrian Alps when they were shoved northward as nappes (thrust sheets) during two separate collision events, one in the Cretaceous and one in the Eocene, that created the modern Alps. It is from the Hallstatt salt diapir that Hallstatt, like so many cities and towns, gets its name.

Deposits of rock salt or halite, the mineral name of sodium chloride with the chemical formula of NaCl, are found and mined around the globe. These deposits mark the dried remains of ancient oceans and seas. Names of rivers, towns and cities in Europe — Salzburg, Halle, Hallstatt, Hallein, La Salle, Moselle — all pay homage to their connection to halite and salt production. The Greek word for salt is hals and the Latin is sal. The Turkish name for salt is Tuz, which we see in the naming of Tuzla, a salt-producing region of northeastern Bosnia-Herzegovina and in the names of towns that dot the coast of Turkey where it meets the Black Sea. Hallstatt with its salt diapir is no exception.

The salt-named town of Hallstatt sits on the shores of the idyllic Hallstätter Sea at the base of the Dachstein massif. Visiting it today, you experience a quaint traditional fishing village built in the typical upper Austrian style. Tourism drives the economy as much as salt as this area of the world is picture-perfect from every angle.

Space is at a minimum in the town. For centuries, every ten years the local cemetery exhumes the bones of those buried there and moves them to an ossuary to make room for new burials. The Hallstatt Ossuary is called Karner, Charnel House, or simply Beinhaus (Bone House). Karners are places of secondary burials. They were once common in the Eastern Alps, but that custom has largely disappeared.

Hallstatt Beinhaus Ossuary, Hallstatt, Austria

A collection of over 700 elaborately decorated skulls rest inside the ossuary. They are lined up on rows of wooden shelves that grace the walls of the chapel. Another 500 undecorated skulls, bare and without any kind of adornment, are stacked in the corners.

Each is inscribed and attached to a record with the deceased's name, profession and date of death. The Bone House is located in a chapel in the basement of the Church of Saint Michael. The church dates from the 12th century CE. 

Decorating the skulls was traditionally the job of the local gravedigger and an honour granted to very few. At the family's request, garlands of flowers were painted on the skulls of deceased as decorative crowns if they were female. The skulls of men and boys were painted wreaths of oak or ivy.

Every building in Hallstatt looks out over the Hallstätter Sea. This beautiful mountain lake considered one of the finest of Austria's Salzkammergut region. It lies at the northern foot of the Dachstein mountain range, sitting eight-and-a-half kilometres long and two kilometres wide. The shoreline is dotted by the villages of  Obertraun, Steeg, and Hallstatt.

The region is habitat to a variety of diverse flora and fauna, including many rare species such as native orchids, in the wetlands and moors in the south and north.

Linked by road to the cities of Salzburg and Graz, Hallstatt and its lake were declared one of the World Heritage sites in Austria in 1997 and included in the Hallstatt-Dachstein Salzkammergut Alpine UNESCO World Heritage Site. The little market village of Hallstatt takes its name from the local salt mine.

Hallstatt, Salzkammergut region, Austria

The town is a popular tourist destination with its quaint shops and terraced cafes. In the centre of town, the 19th-century Evangelical Church of Hallstatt with its tall, slender spire is a lakeside landmark. You can see it here in the photo on the left.

Above the town are the Hallstatt Salt mines located within the 1,030-meter-tall Salzburg Salt Mountain. They are accessible by cable car or a three-minute journey aboard the funicular railway. There is also a wonderful Subterranean Salt Lake.

In 1734, there was a corpse found here preserved in salt. The fellow became known as the Man in Salt. Though no archaeological analysis was performed at the time — the mummy was respectfully reburied in the Hallstatt cemetery — based on descriptions in the mine records, archaeologists suspect the miner lived during the Iron Age. This Old Father, Senos ph₂tḗr, 'ɸatīr 'father' may have been a local farmer, metal-worker, or both and chatted with his friends and family in Celtic or Proto-Celtic.

Salt mining in the area dates back to the Neolithic period, from the 8th to 5th Centuries BC. This is around the time that Roman legions were withdrawing from Britain and the Goths sacked Rome. In Austria, agricultural settlements were dotting the landscape and the alpine regions were being explored and settled for their easy access to valuable salt, chert and other raw materials.

The salt-rich mountains of Salzkammergut and the upland valley above Hallstatt were attractive for this reason. The area was once home to the Hallstatt culture, an archaeological group linked to Proto-Celtic and early Celtic people of the Early Iron Age in Europe, c.800–450 BC.

Bronze Age vessel with cow and calf

In the 19th century, a burial site was discovered with 2,000 individuals, many of them buried with Bronze Age artefacts of amber and ivory.

It was this find that helped lend the name Hallstatt to this epoch of human history. The Late Iron Age, between around 800 and 400 BC, became known as the Hallstatt Period.

For its rich history, natural beauty and breathtaking mountainous geology, Hallstatt is a truly irresistible corner of the world.

Salzbergstraße 1, 4830 Hallstatt.  https://www.salzwelten.at/en/home/

Photo: Bronze vessel with cow and calf, Hallstatt by Alice Schumacher - Naturhistorisches Museum Wien - A. Kern – K. Kowarik – A. W. Rausch – H. Reschreiter, Salz-Reich. 7000 Jahre Hallstatt, VPA 2 (Wien, 2008) Seite 133 Abbildung 6. Hallstatt Village & Ossuary Photos: P. McClure Photography ca. 2015.

Bernoulli D, Jenkyns HC (1974) Alpine, Mediterranean, and Central Atlantic Mesozoic facies in relation to the early evolution of the Tethys. Soc Econ Paleont Mineral Spec Publ 19:129–160

Bernoulli D, Jenkyns H (2009) Ancient oceans and continental margins of the Alpine-Mediterranean Tethys: deciphering clues from Mesozoic pelagic sediments and ophiolites. Sedimentology 56:149–190

WASHINGTON STATE PALAEONTOLOGY

North Cascades National Park, Washington State, USA

Over vast expanses of time, powerful tectonic forces have massaged the western edge of the continent, smashing together a seemingly endless number of islands to produce what we now know as North America and the Pacific Northwest.

Washington is home to a wide variety of fossils—from new species of fossil crabs to marine mollusks and the fossil palm fronds that symbolize the Chuckanut formation.

We also find fossil whales, bird trackways, fossil sockeye salmon, mammal footprints, mammoth bones & the trace fossil remains of ancient rhino. In the time expanse in which we live our very short human lives, the Earth's crust appears permanent.

A fixed outer shell – terra firma. Aside from the rare event of an earthquake or the eruption of Mount St. Helen’s in 1980, our world seems unchanging, the landscape constant. In fact, it has been on the move for billions of years and continues to shift each day. As the earth’s core began cooling, some 4.5 billion years ago, plates, small bits of continental crust, have become larger and smaller as they are swept up in or swept under their neighbouring plates. Large chunks of the ocean floor have been uplifted, shifted and now find themselves thousands of miles in the air, part of mountain chains far from the ocean today or carved by glacial ice into valleys and basins.

Two hundred million years ago, Washington was two large islands, bits of the continent on the move westward, eventually bumping up against the North American continent and calling it home. Even with their new fixed address, the shifting continues; the more extreme movement has subsided laterally and continues vertically. The upthrusting of plates continue to move our mountain ranges skyward, the path of least resistance.

Fossil Palm Front, Washington State

This dynamic movement has created the landscape we see today and helped form the fossil record that tells much of Washington’s relatively recent history – the past 50 million years. Chuckanut Drive is much younger than other parts of Washington.

The fossils found there lived and died some 40-55 million years ago, very close to where they are now, but in a much warmer, swampy setting. The exposures of the Chuckanut Formation were once part of a vast river delta; imagine, if you will, the bayou country of the Lower Mississippi.

The siltstones, sandstones, mudstones and conglomerates of this formation were laid down about 40-54 million years ago during the Eocene epoch, a time of luxuriant plant growth in the subtropical flood plain that covered much of the Pacific Northwest.

This ancient wetland provided ideal conditions to preserve the many trees, shrubs, and plants that thrived here. Plants are important in the fossil record because they are more abundant and can give us a lot of information about climate, temperature, the water cycle, and humidity of the region. The Chuckanut flora is made up predominantly of plants whose modern relatives live in tropical areas such as Mexico and Central America.

Shore Bird Trackway, Washington State

While less abundant, evidence of the animals that called this ancient swamp home are also found here. Rare bird, reptile, and mammal tracks have been immortalized in the outcrops of the Chuckanut Formation.

Tracks of a type of archaic mammal of the Orders Pantodonta or Dinocerata (blunt foot herbivores), footprints from a small shorebird, and tracks from an early equid or webbed bird track give evidence to the vertebrates that inhabited the swamps, lakes and riverways of the Pacific Northwest 50 million years ago.

Fossil mammals from Washington do get most of the press. The movement of these celebrity vertebrates captured in the soft mud on the banks of a river, one of the depositional environments favourable for track preservation.

The bone record is actually far less abundant than the plant record, except near shell middens, given the preserving qualities of calcium and an alkaline environment. While calcium-rich bones and teeth fossilize well, they often do not get laid down in a situation that makes this possible. Hence the terrestrial paleontological record of Washington State at sites like Chuckanut is primarily made up of plant material.

CANADA'S SMILODON

This fierce predator with the luxurious coat is Smilodon fatalis — a compact but robust killer that weighed in around 160 to 280 kg and was 1.5 - 2.2 metres long.

Smilodon is a genus of the extinct machairodont subfamily of the felids. It is one of the most famous prehistoric mammals and the best known saber-toothed cat. Although commonly known as the saber-toothed tiger, it was not closely related to the tiger or other modern cats.

Up until a few years ago, all the great fossil specimens of this apex predator were found south of us in the United States. That was until some interesting bones from Medicine Hat, Alberta got a second look.

A few years ago, a fossil specimen caught the eye of researcher Ashley Reynolds as she was rummaging through the collections at the Royal Ontario Museum in Toronto. 

Back in the 1960s,  University of Toronto palaeontologist C.S. Churcher and his team had collected and donated more than 1,200 specimens from their many field seasons scouring the bluffs of the South Saskatchewan River near Medicine Hat, Alberta.

Churcher is a delightful storyteller and a palaeontologist with a keen eye. I had the very great pleasure of listening to many of his talks out at the University of British Columbia and a few Vancouver Paleontological Society meetings in the mid-2000s. 

"Rufus" was a thoroughly charming storyteller and shared many of his adventures from the field. 

He moved out to the West Coast for his retirement, first to Gabriola Island then to Victoria, but his keen love of the science kept him giving talks to enthralled listeners keen to hear about his survey of the Dakhleh Oasis in the Western Desert of Egypt, geomorphology, stratigraphy, recent biology, Pleistocene and Holocene lithic cultures, insights learned from Neolithic Islamic pottery to Roman settlements.

The specimens he had collected had been roughly sorted but never examined in detail. Reynolds, who was researching the growth patterns and life histories of extinct cats saw a familiar-looking bone from an ancient cat's right front paw. That tiny paw bone had reached through time and was positively identified as Canada's first Smilodon.

These Apex Predators used their exceptionally long upper canine teeth to hunt large mammals. 

Isotopes preserved in the bones of S. fatalis in the La Brea Tar Pits in California tell us that they liked to dine on bison (Bison antiquus) and camels (Camelops) along with deer and tapirs. Smilodon is thought to have killed its prey by holding it still with its forelimbs and biting it. And that was quite the bite!

Their razor-sharp incisors were arranged in an arch. Once they bit down, the teeth would hold their prey still and stabilize it while the canine bite was delivered — and what a bite that was. They could open their mouths a full 120 degrees.

Smilodon died out at the same time that most North and South American megafauna disappeared, about 10,000 years ago. Its reliance on large animals has been proposed as the cause of its extinction, along with climate change and competition with other species.