FOSSILS AND FAUNA OF MADAGASGAR

Aioloceras besairiei (Collingnon, 1949)

A stunning example of the internal suturing with calcite infill in this sliced Aioloceras besairiei (Collingnon, 1949) ammonite from the Upper Cretaceous (Lower Albian) Boeny region of Madagascar. 

This island country is 400 kilometres off the coast of East Africa in the Indian Ocean and a wonderful place to explore off the beaten track.

Madagascar has some of the most spectacular of all the fossil specimens I have ever seen. This beauty is no exception. The shell has a generally small umbilicus, arched to acute venter, and typically at some growth stage, falcoid ribs that spring in pairs from umbilical tubercles, usually disappearing on the outer whorls. I had originally had this specimen marked as a Cleoniceras besairiei, except Cleoniceras and Grycia are not present in Madagascar. 

This lovely, seen in cross-section, is now far from home and in the collection of a wonderful friend. It is an especially lovely example of the ammonite, Aioloceras besairiei, making it a beudanticeratinae. Cleoniceras and Grycia are the boreal genera. If you'd like to see (or argue) the rationale on the name, consider reading Riccardi and Medina's riveting work from back in 2002, or Collingnon from 1949.

The beauty you see here measures in at a whopping 22 cm, so quite a handful. This specimen is from the youngest or uppermost subdivision of the Lower Cretaceous. I'd originally thought this locality was older, but dating reveals it to be from the Lower Albian, so approximately 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma.

Aioloceras are found in the Cretaceous of Madagascar at geo coordinates 16.5° S, 45.9° E: paleo-coordinates 40.5° S, 29.3° E.; and in four localities in South Africa: at locality 36, near the Mzinene River at 28.0° S, 32.3° E: paleo-coordinates 48.6° S, 7.6° E. 

We find them near the Mziene River, at a second locality north of Hluhluwe where the Mzinene Formation overlies the Aptian-Albian Makatini Formation at 28.0° S, 32.3° E: paleo-coordinates 48.6° S, 7.6° E; and at Haughton Z18, on the Pongola River in the Albian III, Tegoceras mosense beds at 27.3° S, 32.2° E: paleo-coordinates 48.0° S, 7.8° E.

If you happen to be trekking to Madagascar, know that it's big. It’s 592,800 square kilometres (or  226,917 square miles), making it the fourth-largest island on the planet — bigger than Spain, Thailand, Sweden and Germany. The island has an interesting geologic history.

Although there has been a geological survey, which was active extending back well into French colonial times, in the non-French-speaking world our geological understanding of the island is still a bit of a mystery. 

Plate tectonic theory had its beginnings in 1915 when Alfred Wegener proposed his theory of "continental drift." 

Wegener proposed that the continents ploughed through the crust of ocean basins, which would explain why the outlines of many coastlines (like South America and Africa) look like they fit together like a puzzle. Half a century after Wegener there is still no agreement as to whether in continental reconstructions Madagascar should be placed adjacent to the Tanzanian coast to the north (e.g., McElhinny and Embleton,1976), against the Mozambique-Natal coast (Flores 1970), or basically left where it is (Kent 1974, Nairn 1978).

There have been few attempts apart from McKinley’s (1960) comparison of the Karoo succession of southwestern Tanzania with that of Madagascar to follow the famous geological precept of “going to sea.” One critical reason is that although there may be a bibliography of several thousand items dealing with Madagascan geology as Besairie (1971) claims, they are items not generally available to the general public. The vital information gained of the geology of the offshore area by post-World War II petroleum exploration has remained largely proprietary. 

Without this data to draw upon, our understanding remains incomplete. I don't actually mind a bit of a mystery here. It is interesting to speculate on how these geologic puzzle pieces fit together and wait for the big reveal. Still, we have good old Besairie from his 1971, Geologie de Madagascar, and a later précis (Besairie, 1973).

We do know that Madagascar was carved off from the African-South American landmass early on. The prehistoric breakup of the supercontinent Gondwana separated the Madagascar–Antarctica–India landmass from the Africa–South America landmass around 135 million years ago. Madagascar later split from India about 88 million years ago, during the Late Cretaceous, so the native plants and animals on the island evolved in relative isolation. 

It is a green and lush island country with more than its fair share of excellent fossil exposures. Along the length of the eastern coast runs a narrow and steep escarpment containing much of the island's remaining tropical lowland forest. If you could look beneath this lush canopy, you'd see rocks of the Precambrian age stretching from the east coast all the way to the centre of the island. The western edge is made up of sedimentary rock from the Carboniferous to the Quaternary.

Red-Tailed Lemurs, Waiwai & Hedgehog

Madagascar is a biodiversity hotspot. Just as Darwin's finches on the Galápagos were isolated, evolving into distinct species (hello, adaptive radiation), over 90% of the wildlife from Madagascar is found nowhere else. 

The island's diverse ecosystems, like so many on this planet, are threatened by Earth's most deadly species, homo sapien sapiens. 

We arrived back in 490 CE and have been chopping down trees and eating our way through the island's tastier populations ever since. Still, they have cuties like this Red-Tailed Lemur. Awe, right?

Today, beautiful outcrops of wonderfully preserved fossil marine fauna hold appeal for me. The material you see from Madagascar is distinctive — and prolific.

Culturally, you'll see a French influence permeating the language, architecture and legal process. There is a part of me that pictures these lovely Lemurs chatting away in French. "Ah, la vache! Regarde le beau fossile, Hérissonne!"

We see the French influence because good 'ol France invaded sleepy Madagascar back in 1883, during the first Franco-Hova War. Malagasy (the local Madagascarian residents) were enlisted as troops, fighting for France in World War I.  During the Second World War, the island was the site of the Battle of Madagascar between the Vichy government and the British. By then, the Malagasy had had quite enough of colonization and after many hiccuping attempts, reached full independence in 1960. Colonization had ended but the tourist barrage had just begun. You can't stop progress.

If you're interested in learning more about this species, check out the Treatise on Invertebrate Paleontology, Part L (Ammonoidea). R.C. Moore (ed). Geological Soc of America and Univ. Kansas Press (1957), p L394. Or head over to look at the 2002 paper from Riccardi and Medina. 2002. Riccardi, A., C. & Medina, F., A. The Beudanticeratinae and Cleoniceratinae (Ammonitina) from the Lower Albian of Patagonia in Revue de Paléobiologie - 21(1) - Muséum d’Histoire Naturelle de la ville de Genève, p 313-314 (=Aioloceras besairiei (COLLIGNON, 1949). You have Bertrand Matrion to thank for the naming correction. Good to have friends in geeky places!

Collignon, M., 1933, Fossiles cenomaniens d’Antmahavelona (Province d’ Analalave, Madagascar), Ann. Geol. Serv. Min. Madagascar, III, 1934 Les Cephalopods du Trias inferieur de Madagascar, Ann. Paleont. XXII 3 and 4, XXII 1.

Besairie, H., 1971, Geologie de Madagascar, 1. Les terrains sedimentaires, Ann. Geol. Madagascar, 35, p. 463.

J. Boast A. and E. M. Nairn collaborated on a chapter in An Outline of the Geology of Madagascar, that is very readable and cites most of the available geologic research papers. It is an excellent place to begin a paleo exploration of the island.

If you happen to parle français, check out: Madagascar ammonites: http://www.ammonites.fr/Geo/Madagascar.htm

THE SLOW RACE OF TIME: LIVING FOSSILS

Horseshoe crabs are marine and brackish water arthropods of the order Xiphosura — a slowly evolving, conservative taxa.

Much like (slow) Water Striders (Aquarius remigis), (relatively sluggish) Coelacanth (Latimeria chalumnae) and (the current winner on really slow evolution) Elephant Sharks (Callorhinchus milii), these fellows have a long history in the fossil record with very few anatomical changes. 

But slow change provides loads of great information. It makes our new friend, Yunnanolimulus luoingensis, an especially interesting and excellent reference point for how this group evolved. 

We can examine their genome today and make comparisons all the way back to the Middle Triassic (with this new find) and other specimens from further back in the Ordovician — 445 million years ago. 

These living fossils have survived all five mass extinction events. They are generalists who can live in shallow or deep water and will eat pretty much anything they can find on the seafloor.

The oldest horseshoe crab fossil, Lunataspis aurora, is found in outcrops in Manitoba, Canada. Charmingly, the name means crescent moon shield of the dawn. It was palaeontologist Dave Rudkin and team who chose that romantic name. Finding them as fossils is quite remarkable as their shells are made of protein which does not mineralized like typical fossils.

Even so, the evolution of their exoskeleton is well-documented by fossils, but appendage and soft-tissue preservation are extremely rare. 

A new study analyzes details of the appendage and soft-tissue preservation in Yunnanolimulus luoingensis, a Middle Triassic (ca. 244 million years old) horseshoe crab from Yunnan Province, SW China. The remarkable anatomical preservation includes the chelicerae, five pairs of walking appendages, opisthosomal appendages with book gills, muscles, and fine setae permits comparison with extant horseshoe crabs.

The close anatomical similarity between the Middle Triassic horseshoe crabs and their recent analogues documents anatomical conservatism for over 240 million years, suggesting persistence of lifestyle.

The occurrence of Carcinoscorpius-type claspers on the first and second walking legs in male individuals of Y. luoingensis tells us that simple chelate claspers in males are plesiomorphic for horseshoe crabs, and the bulbous claspers in Tachypleus and Limulus are derived.

As an aside, if you hadn't seen an elephant shark before and were shown a photo, you would likely say, "that's no freaking shark." You would be wrong, of course, but it would be a very clever observation.

Callorhinchus milii look nothing like our Great White friends and they are not true sharks at all. Rather, they are ghost sharks that belong to the subclass Holocephali (chimaera), a group lovingly known as ratfish. They diverged from the shark lineage about 400 million years ago.

If you have a moment, do a search for Callorhinchus milii. The odd-looking fellow with the ironic name, kallos, which means beautiful in Greek, sports black blotches on a pale silver elongate body. And their special feature? It is the fishy equivalent of business in the front, party in the back, with a dangling trunk-like projection at the tip of their snout and well-developed rectal glands near the tail.

As another small point of interest with regards to horseshoe crabs, John McAllister collected several of these while working on his MSc to see if they had microstructures similar to trilobites (they do) and whether their cuticles were likewise calcified. He found no real calcification in their cuticles, in fact, he had a rather frustrating time getting anything measurable to dissolve in acid in his hunt for trace elements. 

Likewise, when looking at oxygen isotopes (16/18) to get a handle on water salinity and temperature, his contacts at the University of Waterloo had tons of fun getting anything at all to analyze. It made for some interesting findings. Sadly, for a number of reasons, he abandoned the work, but you can read his very interesting thesis here: https://dr.library.brocku.ca/handle/10464/1959

Ref: Hu, Shixue & Zhang, Qiyue & Feldmann, Rodney & Benton, Michael & Schweitzer, Carrie & Huang, Jinyuan & Wen, Wen & Zhou, Changyong & Xie, Tao & Lü, Tao & Hong, Shuigen. (2017). Exceptional appendage and soft-tissue preservation in a Middle Triassic horseshoe crab from SW China. Scientific Reports. 7. 10.1038/s41598-017-13319-x.

OIL IN WATER BEAUTY: FOSSILS OF FOLKSTONE

Sheer beauty — a beautiful Euhoplites ammonite from Folkstone, UK. I've been really enjoying looking at all oil-in-water colouring and chunkiness of these ammonites.

Euhoplites is an extinct ammonoid cephalopod from the Lower Cretaceous, characterized by strongly ribbed, more or less evolute, compressed to inflated shells with flat or concave ribs, typically with a deep narrow groove running down the middle.

In some, ribs seem to zigzag between umbilical tubercles and parallel ventrolateral clavi. In others, the ribs are flexious and curve forward from the umbilical shoulder and lap onto either side of the venter.

Its shell is covered in the lovely lumps and bumps we associate with the genus. The function of these adornments are unknown. I wonder if they gave them greater strength to go deeper into the ocean to hunt for food. 

They look to have been a source of hydrodynamic drag, likely preventing Euhoplites from swimming at speed. Studying them may give some insight into the lifestyle of this ancient marine predator. Euhoplites had shells ranging in size up to a 5-6cm. 

We find them in Lower Cretaceous, middle to upper Albian age strata. Euhoplites has been found in Middle and Upper Albian beds in France where it is associated respectively with Hoplites and Anahoplites, and Pleurohoplites, Puzosia, and Desmoceras; in the Middle Albian of Brazil with Anahoplites and Turrilites; and in the Cenomanian of Texas.

This species is the most common ammonite from the Folkstone Fossil Beds in southeastern England where a variety of species are found, including this 37mm beauty from the collections of José Juárez Ruiz.

FIRST DINOSAUR FROM VANCOUVER ISLAND

This dapper fellow is a pine needle and horsetail connoisseur. He's a hadrosaurus — a duck-billed dinosaur. They were a very successful group of plant-eaters that thrived throughout western Canada during the late Cretaceous, some 70 to 84 million years ago.

Hadrosaurs lived as part of a herd, dining on pine needles, horsetails, twigs and flowering plants.

Hadrosaurs are ornithischians — an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. They are close relatives and possibly descendants of the earlier iguanodontid dinosaurs. 

They had slightly webbed, camel-like feet with pads on the bottom for cushioning and perhaps a bit of extra propulsion in water. They were primarily terrestrial but did enjoy feeding on plants near and in shallow water. There had a sturdy build with a stiff tail and robust bone structure. 

At their emergence in the fossil record, they were quite small, roughly three meters long. That's slightly smaller than an American bison. They evolved during the Cretaceous with some of their lineage reaching up to 20 meters or 65 feet.

Hadrosaurs are very rare in British Columbia but a common fossil in our provincial neighbour, Alberta, to the east. Here, along with the rest of the world, they were more abundant than sauropods and a relatively common fossil find. They were common in the Upper Cretaceous of Europe, Asia, and North America.

There are two main groups of Hadrosaurs, crested and non-crested. The bony crest on the top of the head of the hadrosaurs was hollow and attached to the nasal passages. It is thought that the hollow crest was used to make different sounds. These sounds may have signalled distress or been the hadrosaur equivalent of a wolf whistle used to attract mates. Given their size it would have made for quite the trumpeting sound.

This beautiful specimen graces the back galleries of the Courtenay and District Museum on Vancouver Island, British Columbia, Canada. I was very fortunate to have a tour this past summer with the deeply awesome Mike Trask joined by the lovely Lori Vesper. The museum houses an extensive collection of palaeontological and archaeological material found on Vancouver Island, many of which have been donated by the Vancouver Island Palaeontological Society.

Dan Bowen, Chair of the Vancouver Island Palaeontological Society, shared the photo you see here of the first partly articulated dinosaur from Vancouver Island ever found. The vertebrate photo and illustration are from a presentation by Dr. David Evans at the 2018 Paleontological Symposium in Courtenay.  The research efforts of the VIPS run deep in British Columbia and this new very significant find is no exception. A Hadrosauroid dinosaur is a rare occurrence and further evidence of the terrestrial influence in the Upper Cretaceous, Nanaimo Group, Vancouver Island — outcrops that we traditionally thought of as marine from years of collecting well-preserved marine fossil fauna.

CDM 002 / Hadrosauroid Caudal Vertebrae

The fossil bone material was found years ago by Mike Trask of the Vancouver Island Palaeontological Society. You may recall that he was the same fellow who found the Courtenay Elasmosaur on the Puntledge River.

Mike was leading a fossil expedition on the Trent River. While searching through the Upper Cretaceous shales, the group found an articulated mass of bones that looked quite promising.

Given the history of the finds in the area, the bones were thought to be from a marine reptile.

Since that time, we've found a wonderful terrestrial helochelydrid turtle, Naomichelys speciosa, but up to this point, the Trent had been known for its fossil marine fauna, not terrestrial. Efforts were made to excavate more of the specimen, and in all more than 25 associated vertebrae were collected with the help of some 40+ volunteers. Identifying fossil bone is a tricky business. Encased in rock, the caudal vertebrae were thought to be marine reptile in origin. Some of these were put on display in the Courtenay Museum and mislabeled for years as an unidentified plesiosaur.

In 2016, after years of collecting dust and praise in equal measure, the bones were reexamined. They didn't quite match what we'd expect from a marine reptile. Shino Sugimoto, Fossil Preparator, Vertebrate Palaeontology Technician at the Royal Ontario Museum was called in to work her magic — painstakingly prepping out each caudal vertebrae from the block.

Once fully prepped, seemingly unlikely, they turned out to be from a terrestrial hadrosauroid. This is the second confirmed dinosaur from the Upper Cretaceous Nanaimo Group. The first being a theropod from Sucia Island consisting of a partial left thigh bone — the first dinosaur fossil ever found in Washington state.

Dr. David Evans, Temerty Chair in Vertebrate Palaeontology, Department of Natural History, Palaeobiology from the Royal Ontario Museum, confirmed the ID and began working on the partial duck-billed dinosaur skeleton to publish on the find.

Drawing of Trent River Hadrosauroid Caudal Vertebrae

Now fully prepped, the details of this articulated Hadrosauriod caudal vertebrae come to light. We can see the prominent chevron facets indicative of caudal vertebrae with a nice hexagonal centrum shape on its anterior view.

There are well-defined long, raked neural spines that expand distally — up and away from the acoelous centrum. 

Between the successive vertebrae, there would likely have been a fibrocartilaginous intervertebral body with a gel-like core —  the nucleus pulposus — which is derived from the embryonic notochord. This is a handy feature in a vertebrate built as sturdily as a hadrosaur. Acoelous vertebrae have evolved to be especially well-suited to receive and distribute compressive forces within the vertebral column.

This fellow has kissing cousins over in the state of New Jersey where this species is the official state fossil. The first of his kind was found by John Estaugh Hopkins in New Jersey back in 1838. Since that time, we've found many hadrosaurs in Alberta, particularly the Edmontosuaurs, another member of the subfamily Hadrosaurine.

In 1978, Princeton University found fifteen juvenile hadrosaurs, Maiasaura ("good mother lizard") on a paleontological expedition to the Upper Cretaceous, Two Medicine Formation of Teton County in western Montana. 

Their initial finds of several small skeletons had them on the hunt for potential nests — and they found them complete with wee baby hatchlings!

Photo One: Fossil Huntress / Heidi Henderson, VIPS

Photo Two / Sketch Three: Danielle Dufault, Palaeo-Scientific Ilustrator, Research Assistant at the Royal Ontario Museum, Host of Animalogic. 

The vertebrate photo and illustration were included in a presentation by Dr. David Evans at the 2018 BCPA Paleontological Symposium in Courtenay, British Columbia, Canada.

Photo Four: Illustration by the talented Greer Stothers, Illustrator & Natural Science-Enthusiast.

QUENSTEDTOCERAS WITH PATHOLOGY

What you are seeing here is a protuberance extruding from the venter of Quenstedtoceras cf. leachi (Sowerby). It is a pathology in the shell from hosting immature bivalves that shared the seas with these Middle Jurassic, Upper Callovian, Lamberti zone fauna from the Volga River basin. The collecting site is the now inactive Dubki commercial clay quarry and brickyard near Saratov, Russia. 

The site has produced thousands of ammonite specimens. A good 1,100 of those ended up at the Black Hills Institute of Geological Research in Hill City, South Dakota. 

Roughly 1,000 of those are Quenstedtoceras (Lamberticeras) lamberti and the other 100 are a mix of other species found in the same zone. These included Eboraciceras, Peltoceras, Kosmoceras, Grossouvria, Proriceras, Cadoceras and Rursiceras. 

What is especially interesting is the volume of specimens — 167 Quenstedtoceras (Lamberticeras) lamberti and 89 other species in the Black Hills collection — with healed predation injuries. It seems Quenstedtoceras (Lamberticeras) lamberti are the most common specimens found here and so not surprisingly the most common species found injured. Of the 1,000, 655 of the Quenstedtoceras (Lamberticeras) lamberti displayed some sort of deformation or growth on the shell or had grown in a tilted manner. 

Again, some of the Q. lamberti had small depressions in the centre likely due to a healed bite and hosting infestations of the immature bivalve Placunopsis and some Ostrea. 

The bivalves thrived on their accommodating hosts and the ammonites carried on, growing their shells right up and over their bivalve guests. This relationship led to some weird and deformities of their shells. They grow in, around, up and over nearly every surface of the shell and seem to have lived out their lives there. It must have gotten a bit unworkable for the ammonites, their shells becoming warped and unevenly weighted. Over time, both the flourishing bivalves and the ammonite shells growing up and over them produced some of the most interesting pathology specimens I have ever seen.    

In the photo here from Emil Black, you can see some of the distorted shapes of Quenstedtoceras sp. Look closely and you see a trochospiral or flattened appearance on one side while they are rounded on the other. 

All of these beauties hail from the Dubki Quarry near Saratov, Russia. The ammonites were collected in marl or clay used in brick making. The clay particles suggest a calm, deep marine environment. One of the lovely features of the preservation here is the amount of pyrite filling and replacement. It looks like these ammonites were buried in an oxygen-deficient environment. 

The ammonites were likely living higher in the water column, well above the oxygen-poor bottom. An isotopic study would be interesting to prove this hypothesis. There's certainly enough of these ammonites that have been recovered to make that possible. It's estimated that over a thousand specimens have been recovered from the site but that number is likely much higher. But these are not complete specimens. We mostly find the phragmocones and partial body chambers. Given the numbers, this may be a site documenting a mass spawning death over several years or generations.

If you fancy a read on all things cephie, consider picking up a copy of Cephalopods Present and Past: New Insights and Fresh Perspectives edited by Neil Landman and Richard Davis. Figure 16.2 is from page 348 of that publication and shows the hosting predation quite well. 

Photos: Courtesy of the deeply awesome Emil Black. These are in his personal collection that I hope to see in person one day. 

It was his sharing of the top photo and the strange anomaly that had me explore more about the fossils from Dubki and the weird and wonderful hosting relationship between ammonites and bivalves. Thank you, my friend!

HUMPBACK WHALES: MEGAPTERA NOVAENGLIAE

Look deep into the knowing eye of this magnificent one. He is a Humpback whale, Megaptera novaeangliae, a species of baleen whale for whom I hold a special place in my heart. 

Baleens are toothless whales who feed on plankton and other wee oceanic tasties that they consume through their baleens, a specialised filter of flexible keratin plates that frame their mouth and fit within their robust jaws.

Baleen whales, the mysticetes, split from toothed whales, the Odontoceti, around 34 million years ago. The split allowed our toothless friends to enjoy a new feeding niche and make their way in a sea with limited food resources. There are fifteen species of baleen whales who inhabit all major oceans. Their number include our humbacks, grays, right whales and the massive blue whale. Their territory runs as a wide band running from the Antarctic ice edge to 81°N latitude. These filter feeders

In the Kwak̓wala language of the Kwakiutl or Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, and my cousins on my father's side, whales are known as g̱wa̱'ya̱m. Both the California grey and the Humpback whale live on the coast. Only a small number of individuals in First Nation society had the right to harpoon a whale. This is a practice from many years ago. It was generally only the Chief who was bestowed this great honour. Humpback whales like to feed close to shore and enter the local inlets. Around Vancouver Island and along the coast of British Columbia, this made them a welcome food source as the long days of winter passed into Spring.

Humpback whales are rorquals, members of the Balaenopteridae family that includes the blue, fin, Bryde's, sei and minke whales. The rorquals are believed to have diverged from the other families of the suborder Mysticeti during the middle Miocene. 

While cetaceans were historically thought to have descended from mesonychids—which would place them outside the order Artiodactyla—molecular evidence supports them as a clade of even-toed ungulates—our dear Artiodactyla. 

It is one of the larger rorqual species, with adults ranging in length from 12–16 m (39–52 ft) and weighing around 25–30 metric tons (28–33 short tons). The humpback has a distinctive body shape, with long pectoral fins and a knobbly head. It is known for breaching and other distinctive surface behaviours, making it popular with whale watchers and the lucky few who see them from the decks of our local ferries.

Both male and female humpback whales vocalize, but only males produce the long, loud, complex "song" for which the species is famous. Males produce a complex soulful song lasting 10 to 20 minutes, which they repeat for hours at a time. I imagine Gregorian Monks vocalizing their chant with each individual melody strengthening and complimenting that of their peers. All the males in a group produce the same song, which differed in each season. Its purpose is not clear, though it may help induce estrus in females and bonding amongst the males.

Humpback Whale, Megaptera novaeangliae

Found in oceans and seas around the world, humpback whales typically migrate up to 25,000 km (16,000 mi) each year. 

They feed in polar waters and migrate to tropical or subtropical waters to breed and give birth, fasting and living off their fat reserves. Their diet consists mostly of krill and small fish. 

Humpbacks have a diverse repertoire of feeding methods, including the bubble net technique.

Humpbacks are a friendly species that interact with other cetaceans such as bottlenose dolphins. They are also friendly and oddly protective of humans. You may recall hearing about an incident off the Cook Islands a few years back. Nan Hauser was snorkelling and ran into a tiger shark. Two adult humpback whales rushed to her aid, blocking the shark from reaching her and pushing her back towards the shore. We could learn a thing or two from their kindness. We have not been as good to them as they have been to us.

Like other large whales, the humpback was a tasty and profitable target for the whaling industry. My grandfather and uncle participated in that industry out of Coal Harbour on northern Vancouver Island back in the 1950s. So did many of my First Nation cousins. My cousin John Lyon has told me tales of those days and the slippery stench of that work.

Six whaling stations operated on the coast of British Columbia between 1905 and 1976. Two of these stations were located at Haida Gwaii, one at Rose Harbour and the other at Naden Harbour. Over 9,400 large whales were taken from the waters around Haida Gwaii. The catch included blue whales, fin whales, sei whales, humpback whales, sperm whales and right whales. In the early years of the century, primarily humpback whales were taken. In later years, fin whales and sperm whales dominated the catch. 

Whales were hunted off South Moresby in Haida Gwaii, on the north side of Holberg Inlet in the Quatsino Sound region. It was the norm at the time and a way to make a living, especially for those who had hoped to work in the local coal mine but lost their employment when it shut down. 

While my First Nations relatives hunted whales in small numbers and many years ago, my Norwegian relatives participated in the hunt on a scale that nearly led to their extinction before the process was banned. The Coal Harbour Whaling Station closed in 1967. Once it had closed, my grandfather Einar Eikanger, my mother's father, took to fishing and my uncle Harry lost his life the year before when he slipped and fell over the side of the boat. He was crushed between the hull and a Humpback in rough seas. 

Humpback populations have partially recovered since that time to build their population up to 80,000 animals worldwide—but entanglement in fishing gear, collisions with ships, and noise pollution continue to negatively impact the species. So be kind if you see them. Turn your engine off and see if you can hear their soulful cries echoing in the water.

I did up a video on Humpback Whales over on YouTube so you could see them in all their majesty. Here is the link: https://youtu.be/_Vbta7kQNoM

ATURIA: MIOCENE NAUTILOID

Aturia angustata, Lower Miocene, WA

This lovely Lower Miocene nautiloid is Aturia angustata collected on the foreshore near Clallam Bay, Olympic Peninsula, northwestern Washington. 

Aturia is an extinct genus of Paleocene to Miocene nautiloid within Aturiidae, a monotypic family, established by Campman in 1857 for Aturia (Bronn, 1838), and is included in the superfamily Nautilaceae (Kümmel,  1964).

There are seven living nautiloid species in two genera: Nautilus pompilius, N. macromphalus, N. stenomphalus, N. belauensis, and the three new species being described from Samoa, Fiji, and Vanuatu (Ward et al.). We have specimens of fossil nautiloids dating to the Turonian of California, and possibly the Cenomanian of Australia. There has also been a discovery of what might be the only known fossil of Allonautilus (Ward and Saunders, 1997), from the Nanaimo Group of British Columbia, Canada.

Aturia in the Collection of Rick Ross, VIPS

The exquisite shell preservation of many Nanaimo nautilids has opened up a lens into paleotemperatures and accurate Nitrogen isotope analyses. 

Nautilus and all other known Cretaceous through Paleogene nautiloids were shallow water carnivores. We may see their shells as beautiful bits of art and science today, but they were seen in our ancient oceans as small yet mighty predators. Preferring to dine on shrimp, crab, fish and on occasion, a friendly cousin nautiloid to two.

Aturia lived in cooler water in the Cenozoic, preferring it over the warmer waters chosen by their cousins. Aturia, are commonly found as fossils from Eocene and Miocene outcrops. That record ends with their extinction in the late Miocene. This was a fierce little beast with jaws packed with piranha-like teeth. They grew at least twice that of the largest known Nautilus living today. 

Aturia is characterized by a smooth, highly involute, discoidal shell with a complex suture and subdorsal siphuncle. The shell of Aturia is rounded ventrally and flattened laterally; the dorsum is deeply impressed. The suture is one of the most complex within the subclass Nautiloidea. Of all the nautiloids, he may have been able to go deeper than his brethren.

Nautiloids are known for their simple suturing in comparison to their ammonite cousins. This simplicity of design limited their abilities in terms of withstanding the water pressure experienced when several atmospheres below the sea. Nautiloids were not able to compete with their ammonite cousins in this regard. 

Instead of elaborate and complex sutures capable of withstanding the pressures of the deep, nautiloids have simpler sutures that would have them enfold on themselves and crush at depth.  

Aturia angustata; Rick Ross Collection

It has a broad flattened ventral saddle, narrow pointed lateral lobes, broad rounded lateral saddles, broad lobes on the dorso-umbilical slopes, and a broad dorsal saddle divided by a deep, narrow median lobe. 

The siphuncle is moderate in size and located subdorsally in the adapical dorsal flexure of the septum. Based on the feeding and hunting behaviours of living nautiluses, Aturia most likely preyed upon small fish and crustaceans. 

I've found a few of these specimens along the beaches of Clallam Bay and nearby in a local clay quarry. I've also seen calcified and chalcedony — microcrystalline quartz — agatized beauties of this species collected from river sites within the Olympic Peninsula range. In the bottom photos, you can see Aturia from Washington state and one (on the stand on the left) from Oregon, USA. These beauties are in the collections of the deeply awesome Rick Ross, Vancouver Island Palaeontological Society.

References: Ward, P; Haggart, J; Ross, R; Trask, P; Beard, G; Nautilus and Allonautilus in the Nanaimo Group, and in the modern oceans; 12th British Columbia Paleontological Symposium, 2018, Courtenay, abstracts; 2018 p. 10-11

YING YANG: ZYCHASPISES

Zenaspis pololica

A Devonian bony fish mortality plate showing a lower shield of Zenaspis podolica (Lankester, 1869) from Lower Devonian deposits of Podolia, Ukraine.

While war rages on in the Ukraine, our hearts go out to those who live and work here contributing much to our understanding of Podolia, a historic region in Eastern Europe, located in the west-central and south-western parts of Ukraine, in northeastern Moldova. 

As I write this, thousands of Ukrainians are arriving at the Palanca border crossing in Moldova on Ukraine's western border to escape the fighting and pincher advance of the Russian military in the north, south and east. 

Looking at a map of the Ukraine, I imagine a torch touching the edges that mental map alighting it with flame — the edges slowly burning and curling in as the world watches and good people on both sides feel that burn.

We look to the Ukraine with our modern lens as we are want to do. Ukraine emerged as the concept of a nation and the Ukrainians as a nationality with the Ukrainian National Revival in the mid-18th century, in the wake of the peasant revolt of 1768-1769 and the eventual partition of the Polish–Lithuanian Commonwealth. They have known a tense love/hate relationship with their neighboring nations for a long while, sharing culture and language but desiring freedom and independence. 

It became an independent state in 1991 with the collapse of the Soviet Union. This fertile part of the world has an archaeology record of human habitation that goes back to the first millennium BC — the oldest known main inhabitants of Ukraine were Cimmerians. They were replaced in Fifth Century BC by Scythians who ruled till 2nd Century BC. The Sarmatian tribes replaced them. Wars, battles and skirmishes ensued until the tribesmen of the dominant horde, the Alanis, could be said to rule in the First Century AD. While our recorded history, our ancestors likely hunted and past through here far earlier. 

Beneath our human habitation and current military action is bedrock that tells the story of the Earth's violent past. The fauna here are from the Silurian and record a time in our Earth's history where the planet suffered a major mass extinction event that wiped out 23 percent of all marine life. It is the only region in Ukraine where 420 million-year-old remains of ichthyofauna can be found near the surface, making them accessible to collection and study. 

Zenaspis is an extinct genus of jawless fish which thrived during the early Devonian. Being jawless, Zenaspis was probably a bottom feeder, dining on debris from the seafloor similar to how flounder, groupers, bass and other bottom-feeding fish make a living.

For the past 150 years, vertebrate fossils have been found in more than 90 localities situated in outcrops along banks of the Dniester River and its northern tributaries, and in sandstone quarries. 

At present, the faunal list of Early Devonian agnathans and fishes from Podolia number seventy-two species, including 8 Thelodonti, 39 Heterostraci, 19 Osteostraci, 4 Placodermi, 1 Acanthodii, and 1 Holocephali (Voichyshyn 2001a).

In Podolia, the Lower Devonian Redbeds strata (the Old Red Formation or Dniester Series) are 1800 metres thick and range from Lochkovian to Eifelian in age (Narbutas 1984; Drygant 2000, 2003).

In their lower part, the Ustechko and Khmeleva members of the Dniester Series are built from lovely multicoloured, mainly red, fine-grained cross-bedded massive quartz sandstones and siltstones with seams of argillites (Drygant 2000).

We see fossils of Zenaspis in the early Devonian of Western Europe. Both Zenaspis pagei and Zenaspis poweri can be found up to 25 centimetres long in Devonian outcrops of Scotland.

Reference: Voichyshyn, V. 2006. New osteostracans from the Lower Devonian terrigenous deposits of Podolia, Ukraine. Acta Palaeontologica Polonica 51 (1): 131–142. Photo care of the awesome Fossilero Fisherman, a fossil hunter devoted to collecting the Lower Devonian Creek exposures of the Ukraine

WEST COAST OYSTERS: T'LOXT'LOX

One of the now rare species of oysters in the Pacific Northwest is the Olympia oyster, Ostrea lurida, (Carpenter, 1864).  

While rare today, these are British Columbia’s only native oyster. 

Had you been dining on their brethren in the 1800s or earlier, it would have been this species you were consuming. Middens from Port Hardy to California are built from Ostrea lurida.

These wonderful invertebrates bare their souls with every bite. Have they lived in cold water, deep beneath the sea, protected from the sun's rays and heat? Are they the rough and tumble beach denizens whose thick shells tell us of a life spent withstanding the relentless pounding of the sea? Is the oyster in your mouth thin and slimy having just done the nasty—spurred by the warming waters of Spring? 

Is this oyster a local or was it shipped to your current local and, if asked, would greet you with "Kon'nichiwa?" Not if the beauty on your plate is indeed Ostrea lurida. 

Oyster in Kwak'wala is t̕łox̱t̕łox̱

We have been cultivating, indeed maximizing the influx of invasive species to the cold waters of the Salish Sea for many years. 

But in the wild waters off the coast of British Columbia is the last natural abundant habitat of the tasty Ostrea lurida in the pristine waters of  Nootka Sound. 

The area is home to the Nuu-chah-nulth First Nations who have consumed this species boiled or steamed for thousands of years. Here these ancient oysters not only survive but thrive — building reefs and providing habitat for crab, anemones and small marine animals. 

Oysters are in the family Ostreidae — the true oysters. Their lineage evolved in the Early Triassic — 251 - 247 million years ago. 

In the Kwak̓wala language of the Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest and my family, an oyster is known as t̕łox̱t̕łox̱. 

I am curious to learn if any of the Nuu-chah-nulth have a different word for an oyster. If you happen to know, I would be grateful to learn.

COAHUILACERATOPS MAGNACUERNA

Coahuilaceratops or "Coahuila Horn Face," is a relatively new genus of Ornithischia Ceratopsidae, a  herbivorous ceratopsian dinosaur who lived during the Upper Cretaceous (late Campanian) near the town of Porvenir de Jalpa (about 64 km / 40 miles west of Saltillo) in what is now southern Coahuila (formerly Coahuila de Zaragoza), northern Mexico.

The Sierra Madre Oriental mountain range runs northwest to southwest forming a spine through the centre of the State. East of the range, the arid landscape slopes gently through the desert terrain down to the Rio Grande. It is home to wonderful common, rare and endangered cacti, beautiful (and one of my favourite) raptors, Aquila chrysaetos and the evolutionarily unlikely pronghorn, Antilocapra americana (if a monkey/owl/ antelope had a baby...)

The world was a much wetter warmer place when these big beauties roamed. Picture them ambling through lush vegetation and rearing young next to freshwater rivers, brackish swamps and salty ancient seas. Many of the dinosaur remains from the area bear the marks or remains of fossilized snails and clams. Perhaps predation vs a symbiotic relationship as proximity isn't always intimacy. Coahuilaceratops magnacuerna is known from holotype CPC 276, a partial skeleton of an adult along with bits and pieces of skull, a section of horn, pretty complete lower jaw, a smidge of the upper jaw and part of the frill.

Another specimen, CPS 277, has been touted as a possible juvenile Coahuilaceratops. All the specimens from Coahuilaceratops come from a single Upper Cretaceous (Campanian) locality of the Cerro del Pueblo Formation, northern Mexico.

This particular species of Coahuilaceratops was formally named C. magnacuerna by Mark A. Loewen, Scott D. Sampson, Eric K. Lund, Andrew A. Farke, Martha C. Aguillón-Martínez, C.A. de Leon, R.A. Rodríguez-de la Rosa, Michael A. Getty and David A. Eberth in 2010. Though the name was in circulation informally by those working in the study of ceratopsian dinosaurs as early as 2008.

Though challenged by examining and interpreting mere bits and pieces, the team posed estimates on the overall size of this new rather largish, 6.7 m / 22 ft, chasmosaurine. Coahuilaceratops' horns are also impressively large, estimated at 1.2 m / 4 feet. Rather long for a ceratopsian (consider that a Triceratops distinctive horn generally comes in under 115 cm / 45 inches and interesting in terms of evolutionary design. The holotypes are available for viewing at the Museo del Desierto in Saltillo, Coahuila. Photo credit: José F. Ventura

THE GIANT FOSSIL AMMONITE OF FERNIE

Titanites occidentalis, Fernie Ammonite

The Fernie ammonite, Titanites occidentalis, from outcrops on Coal Mountain near Fernie, British Columbia, Canada. 

This beauty is the remains of a carnivorous cephalopod within the family Dorsoplanitidae that lived and died in a shallow sea some 150 million years ago.

If you would like to get off the beaten track and hike up to see this ancient beauty, you will want to head to the town of Fernie in British Columbia close to the Alberta border. 

This is the traditional territory of the the Yaq̓it ʔa·knuqⱡi ‘it First Nation who have lived here since time immemorial. There was some active logging along the hillside in 2021, so if you are looking at older directions on how to get to the site be mindful that many of the trailheads have been altered and a fair bit of bushwhacking will be necessary to get to the fossil site proper. That being said, the loggers from CanWel may have clear-cut large sections of the hillside but they did give the ammonite a wide berth and have left it intact.

Wildsight, a non-profit environmental group out of the Kimberly Cranbrook area has been trying to gain grant funding to open up the site as an educational hike with educational signage for folks visiting the Fernie area. It is likely the province of British Columbia would top up those funds if they are able to place the ammonite under the Heritage Conservation Act. CanWel would remain the owners of the land but the province could assume the liability for those visiting this iconic piece of British Columbia's palaeontological history. 

Driving to the trail base is along an easy access road just east of town along Fernie Coal Road. There are some nice exposures of Cretaceous plant material on the north side (left-hand side) of the road as you head from Fernie towards Coal Creek. I recently drove up to Fernie to look at Cretaceous plant material and locate the access point to the now infamous Late Jurassic (Tithonian) Titanites (S.S. Buckman, 1921) site. While the drive out of town is on an easy, well-maintained road, the slog up to the ammonite site is often a wet, steep push.

Fernie, British Columbia, Canada

The first Titanites occidentalis was about one-third the size and was incorrectly identified as Lytoceras, a fast-moving nektonic carnivore. The specimen you see here is significantly larger at 1.4 metres (about four and a half feet) and rare in North America. 

Titanites occidentalis, the Western Giant, is the second known specimen of this extinct fossil species. 

The first was discovered in 1947 in nearby Coal Creek by a British Columbia Geophysical Society mapping team. When they first discovered this marine fossil high up on the hillside, they could not believe their eyes — both because it is clearly marine at the top of a mountain and the sheer size of this ancient beauty.

In the summer of 1947, a field crew was mapping coal outcrops for the BC Geological Survey east of Fernie. One of the students reported finding “a fossil truck tire.” Fair enough. The similarity of size and optics are pretty close to your average Goodridge. 

A few years later, GSC Paleontologist Hans Frebold described and named the fossil Titanites occidentalis after the large Jurassic ammonites from Dorset, England. The name comes from Greek mythology. Tithonus, as you may recall, was the Prince of Troy. He fell in love with Eos, the Greek Goddess of the Dawn. Eos begged Zeus to make her mortal lover immortal. Zeus granted her wish but did not grant Tithonus eternal youth. He did indeed live forever — ageing hideously. Ah, Zeus, you old trickster. It is a clever play on time placement. Dawn is the beginning of the day and the Tithonian being the latest age of the Late Jurassic. Clever Hans!

HIKING TO THE FERNIE AMMONITE

From the town of Fernie, British Columbia, head east along Coal Creek Road towards Coal Creek. The site is 3.81 km from the base of Coal Creek Road to the trailhead as the crow flies. I have mapped it here for you in yellow and added the wee purple GPS marker for the ammonite site proper. There is a nice, dark grey to black roadcut exposure of Cretaceous plants on the north side of the dirt road that is your cue to pull over and park.  

You access what is left of the trailhead on the south side of the road. You will need to cross the creek to begin your ascent. There is no easy way across the creek and you'll want to tackle this one with a friend when the water level is low. 

The beginning of the trail is not clear but a bit of searching will reveal the trailhead with its telltale signs of previous hikers. This is a moderate 6.3-kilometre hike up & back bushwhacking through scrub and fallen trees. Heading up, you will make about a 246-metre elevation gain. You will likely not have a cellular signal up here but if you download the Google Map to your mobile, you will have GPS to guide you. The area has been recently logged so much of the original trail has been destroyed. There may now be easier vehicle access up the logging roads but I have not driven them since the logging and new road construction.

If you are coming in from out of town, the closest airport is Cranbrook. Then it is about an hour and change to Fernie and another 15-minutes or so to park near the site.

You will want to leave your hammers with your vehicle (no need to carry the weight and this lovely should never be struck with anything more than a raindrop) as this site is best enjoyed with a camera. 

This is a site you will want to wear hiking boots to access. Know that these will get wet as you cross the creek. 

If you would like to see the ammonite but are not keen on the hike, a cast has been made by fossil preparator Rod Bartlett is on display at the Courtenay Museum in Courtenay, Vancouver Island, Canada. 

Respect for the Land / Leave No Trace

As your feet move up the hillside, you can imagine this land 10,000 years ago, rising above great glaciers. Where footfalls trace the steps of those that came before you. This land has been home to the Yaq̓it ʔa·knuqⱡi ‘it First Nation and Ktunaxa or Kukin ʔamakis First Nations whose oral history have them living here since time immemorial. Like them, take only what you need and no more than the land offers — packing out anything that you packed in. 

Fernie Ammonite Palaeo Coordinates: 49°29'04"N 115°00'49"W

JELLYFISH: DANCERS OF THE DEEP

This lovely ocean dancer—with her long delicate tentacles or lappets and thicker ruched oral arms—is a jellyfish. 

Her brethren are playing in the waters of the deep all over the world, from surface waters to our deepest seas — and they are old. They are some of the oldest animals in the fossil record.

Jellyfish, or sea jellies, are the informal common names given to the medusa-phase or adult phase of certain gelatinous members of the subphylum Medusozoa, a major part of the phylum Cnidaria — closely related to anemones and corals.

While the name is embedded, Jellyfish are not fish at all. They evolved millions of years before true fish. The oldest conulariid scyphozoans appeared between 635 and 577 million years ago in the Neoproterozoic of the Lantian Formation, a 150-meter-thick sequence of rocks deposited in southern China. 

Others are found in the youngest Ediacaran rocks of the Tamengo Formation of Brazil, c. 505 mya, through to the Triassic. Cubozoans and hydrozoans appeared in the Cambrian of the Marjum Formation in Utah, USA, c. 540 million years ago.

I have seen all sorts of their brethren growing up on the west coast of Canada. I have seen them in tide pools, washed up on the beach and swam amongst thousands of Moon Jellyfish while scuba diving in the Salish Sea. Their movement in the water is marvelous.  

In the Kwak̓wala language of the Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, jellyfish are known as ǥaǥisama—enjoyed as a tasty snack or used as bait to entice larger marine animals.

The watercolour ǥaǥisama you see here in dreamy pink and white is but one colour variation. They come in blue, purple, orange, yellow and clear — and are often luminescent. They produce light by the oxidation of a substrate molecule, luciferin, in a reaction catalyzed by a protein, luciferase.

NEVADA FOSSILS: CARNIAN-NORIAN BOUNDARY

Time Slows at Berlin-Ichthyosaur State Park

High on the hillside up a long entry road sits the entrance to Berlin-Ichthyosaur State Park in central Nevada.

A worn American flag and sun bleached outbuildings greet you on your way to the outcrops. Away from the hustle and bustle that define the rest of Nevada this place feels remarkably serene. Your eyes squint against the sun as you search for ammonoids and other marine fossil fauna while your nose tends to the assault from the bracing smell of sage brush.

This site holds many stories. The interpretive centre displays wonderful marine reptiles, ichthyosaurs in situ, as you might expect from the name of the park — but it also showcases years of history lovingly tended. This stretch of dry golden low hills dappled with the yellow of creosote and desert grasses is an important locality for our understanding of the Carnian-Norian boundary (CNB) in North America.

The area is known worldwide as one of the most important ichthyosaur Fossil-Lagerstätte because of the sheer volume of remarkably well-preserved, fully articulated (all the sweet bones laid out all in a row...) specimens of Shonisaurus popularis.

Rich ammonoid faunas outcrop in the barren hills of the Upper Triassic (Early Norian, Kerri zone), Luning Formation, West Union Canyon, Nevada. They were studied by N. J. Silberling (1959) and provide support for the definition of the Schucherti and Macrolobatus zones of the latest Carnian — which are here overlain by well-preserved faunas of the earliest Norian Kerri Zone. 

The genus Gonionotites, very common in the Tethys and British Columbia, is for the moment, unknown in Nevada. The Upper Carnian faunas are dominated by Tropitidae, while Juvavitidae are conspicuously lacking. 

Middle Triassic Ammonoids

Despite its importance, no further investigations had been done at this site for a good 50 years. That changed in 2010 when Jim Haggart, Mike Orchard and Paul Smith — all local Vancouverites — collaborated on a project that took them down to Nevada to look at the conodonts and ammonoids. They did a bed-by-bed sampling of ammonoids and conodonts in West Union Canyon during October of that year.

October is an ideal time to do fieldwork in this area. There are a few good weeks between screaming hot and frigid cold. It is also tarantula breeding season so keep your eyes peeled. Those sweet little burrows you see are not from rodents but rather largish arachnids. 

The eastern side of the canyon provides the best record of the Macrolobatus Zone, which is represented by several beds yielding ammonoids of the Tropites group, together with Anatropites div. sp. 

Conodont faunas from both these and higher beds are dominated by ornate metapolygnthids that would formerly have been collectively referred to Metapolygnathus primitius, a species long known to straddle the CNB. Within this lower part of the section, they resemble forms that have been separated as Metapolygnathus mersinensis. Slightly higher, forms close to Epigondolella' orchardi and a single Orchardella n. sp. occur. This association can be correlated with the latest Carnian in British Columbia.

Higher in the section, the ammonoid fauna shows a sudden change and is dominated by Tropithisbites. Few tens of metres above, but slightly below the first occurrence of Norian ammonoids Guembelites jandianus and Stikinoceras, two new species of conodonts (Gen et sp. nov. A and B) appear that also occur close to the favoured Carnian/Norian boundary at Black Bear Ridge, British Columbia. Stratigraphically higher collections continue to be dominated by forms close to M. mersinensis and E. orchardi after BC's own Mike Orchard.

The best exposure of the Kerri Zone is on the western side of the West Union Canyon. Ammonoids, dominated by Guembelites and Stikinoceras div. sp., have been collected from several fossil-bearing levels. Conodont faunas replicate those of the east section. The collected ammonoids fit perfectly well with the faunas described by Silberling in 1959, but they differ somewhat from coeval faunas of the Tethys and Canada. 

The ammonoid fauna paints a compelling picture of Tethyan influence with a series of smoking guns. We see an abundance of Tropitidae in the Carnian, a lack of Pterosirenites in the Norian, copious Guembelites, the Tethyan species G. philostrati, the stratigraphic position of G. clavatus and the rare occurrence of Gonionotites. Their hallelujah moment was likely finding an undescribed species of the thin-shelled bivalve Halobia similar to Halobia beyrichi — the clincher that perhaps seals this deal on Tethyan influence. 

I'll take a boo to see what Christopher McRoberts published on the find. A jolly good idea to have him on this expedition as it would have been easy to overlook if the focus remained solely on the conodonts and ammonoids. McRoberts has published on the much-studied Pardonet Formation up in the Willison Lake Area of Northeastern, British Columbia. He knows a thing or two about Upper Triassic Bivalvia and the correlation to coeval faunas elsewhere in the North American Cordillera, and to the Boreal, Panthalassan and Tethyan faunal realms. 

If you fancy a read, they published a paper: "Towards the definition of the Carnian/Norian Boundary: New data on Ammonoids and Conodonts from central Nevada," which you can find in the proceedings of the 21st Canadian Paleontology Conference; by Haggart, J W (ed.); Smith, P L (ed.); Canadian Paleontology Conference Proceedings no. 9, 2011 p. 9-10.

Fig. 1. Location map of Berlin-Ichthyosaur State Park

Marco Balini, James Jenks, Riccardo Martin, Christopher McRoberts, along with Mike Orchard and Norman Siberling, did a bed by bed sampling in 2013 and published on The Carnian/Norian boundary succession at Berlin-Ichthyosaur State Park (Upper Triassic, central Nevada, USA) and published in January 2014 in Paläontologische Zeitschrift 89:399–433. That work is available for download from ResearchGate. The original is in German, but there is a translation available.

After years of reading about the correlation between British Columbia and Nevada, I had the very great pleasure of walking through these same sections in October 2019 with members of the Vancouver Paleontological Society and Vancouver Island Palaeontological Society. It was with that same crew that I'd originally explored fossil sites in the Canadian Rockies in the early 2000s. Those early trips led to paper after paper and the exciting revelations that inspired our Nevada adventure.

If you plan your own adventure, you'll want to keep an eye out for some of the other modern fauna — mountain lions, snakes, lizards, scorpions, wolves, coyotes, foxes, ground squirrels, rabbits, falcons, hawks, eagles, bobcats, sheep, deer and pronghorns.

Figure One: Location map of Berlin-Ichthyosaur State Park. A detailed road log with access information for this locality is provided in Lucas et al. (2007).

SAKARA MADAGASGAR: OXFORDIAN OUTCROPS

This big beastie is a superb specimen of the ammonite Lobolytoceras costellatum showing the intricate fractal pattern of its septa. This lovely measures to a whopping 230 mm and hails from Oxfordian outcrops near Sakara, Madagascar. Lovingly prepped by the supremely talented José Juárez Ruiz.

Ammonites were predatory, squidlike creatures that lived inside coil-shaped shells. Like other cephalopods, ammonites had sharp, beak-like jaws inside a ring of squid-like tentacles that extended from their shells. They used these tentacles to snare prey — plankton, vegetation, fish and crustaceans — similar to the way a squid or octopus hunt today.

Catching a fish with your hands is no easy feat, as I'm sure you know. Ammonites did the equivalent, catching prey in their tentacles. They were skilled and successful hunters. They caught their prey while swimming and floating in the water column. Within their shells, they had a number of chambers, called septa, filled with gas or fluid that were interconnected by a wee air tube. By pushing air in or out, they were able to control their buoyancy in the water column.

They lived in the last chamber of their shells, continuously building new shell material as they grew. As each new chamber was added, the squid-like body of the ammonite would move down to occupy the final outside chamber.

They were a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids — octopuses, squid, and cuttlefish) then they are to shelled nautiloids such as the living Nautilus species.

Ammonites have intricate and complex patterns on their shells called sutures. The suture patterns differ across species and tell us what time period the ammonite is from. If they are geometric with numerous undivided lobes and saddles and eight lobes around the conch, we refer to their pattern as goniatitic, a characteristic of Paleozoic ammonites.

Ammonites first appeared about 240 million years ago, though they descended from straight-shelled cephalopods called bacrites that date back to the Devonian, about 415 million years ago, and the last species vanished in the Cretaceous–Paleogene extinction event.

They were prolific breeders that evolved rapidly. If you could cast a fishing line into our ancient seas, it is likely that you would hook an ammonite, not a fish. They were prolific back in the day, living (and sometimes dying) in schools in oceans around the globe. We find ammonite fossils (and plenty of them) in sedimentary rock from all over the world.

In some cases, we find rock beds where we can see evidence of a new species that evolved, lived and died out in such a short time span that we can walk through time, following the course of evolution using ammonites as a window into the past.

For this reason, they make excellent index fossils. An index fossil is a species that allows us to link a particular rock formation, layered in time with a particular species or genus found there. Generally, deeper is older, so we use the sedimentary layers rock to match up to specific geologic time periods, rather the way we use tree-rings to date trees. A handy way to compare fossils and date strata across the globe.

GRAY WHALES: ESCHRICHTIUS ROBUSTUS

Young Gray Whale, Eschrichtius robustus

The lovely fellow you see here is a young Gray Whale, Eschrichtius robustus, with a wee dusting of barnacles and his mouth ajar just enough to show his baleen.

Two Pacific Ocean populations are known to exist: one of about 200 individuals whose migratory route is presumed to be between the Sea of Okhotsk off Russia's south coast and southern Korea, and a larger one with a population of about 27,000 individuals in the eastern Pacific.

This second group are the ones we see off the shores of British Columbia as they travel the waters from northernmost Alaska down to Baja California. Gray whale mothers make this journey accompanied by their calves, hugging the shore in shallow kelp beds and providing rare but welcome glimpses of this beauty.

The gray whale is traditionally placed as the only living species in its genus and family, Eschrichtius and Eschrichtiidae, but an extinct species was discovered and placed in the genus in 2017 — the Akishima whale, E. akishimaensis. Some recent DNA analyses suggest that certain rorquals of the family Balaenopteridae, such as the humpback whale, Megaptera novaeangliae, and fin whale, Balaenoptera physalus, are more closely related to the gray whale than they are to some other rorquals, such as minke. Still, others place gray whales as outside the rorqual clade, a kissing cousin if you will.

John Edward Gray placed it in its own genus in 1865, naming it in honour of physician and zoologist Daniel Frederik Eschricht. The common name of the whale comes from its colouration. The subfossil remains of now-extinct gray whales from the Atlantic coasts of England and Sweden were used by Gray to make the first scientific description of a species then surviving only in Pacific waters. The living Pacific species was described by American palaeontologist, Edward Drinker Cope as Rhachianectes glaucus in 1869.

Fin Whale, Balaenoptera physalus

Skeletal comparisons showed the Pacific species to be identical to the Atlantic remains in the 1930s, and Gray's naming has been generally accepted since. Although identity between the Atlantic and Pacific populations cannot be proven by anatomical data, its skeleton is distinctive and easy to distinguish from that of all other living whales.

In 1993, a twenty-seven million-year-old specimen was discovered in deposits in Washington state that represents a new species of early baleen whale. It is especially interesting as it is from a stage in the group’s evolutionary history when baleen whales transitioned from having teeth to filtering food with baleen bristles.

Visiting researcher Carlos Mauricio Peredo studied the fossil whale remains, publishing his research to solidify Sitsqwayk cornishorum (pronounced sits-quake) in the annals of history. The earliest baleen whales clearly had teeth, and clearly still used them. Modern baleen whales have no teeth and have instead evolved baleen plates for filter feeding. Look to the rather good close-up of this young Gray Whale here to see his baleen where once there was a toothy grin.

The baleen is the comb-like strainer that sits on the upper jaw of baleen whales and is used to filter food. We have to ponder when this evolutionary change —moving from teeth to baleen — occurred and what factors might have caused it. Traditionally, we have sought answers about the evolution of baleen whales by turning to two extinct groups: the aetiocetids and the eomysticetids.

The aetiocetids are small baleen whales that still have teeth, but they are very small, and it remains uncertain whether or not they used their teeth. In contrast, the eomysticetids are about the size of an adult Minke Whale and seem to have been much more akin to modern baleen whales; though it’s not certain if they had baleen. Baleen typically does not preserve in the fossil record being soft tissue; generally, only hard tissue, bones and teeth are fossilized.