Keuppia is in the family Palaeoctopodidae, and one of the earliest representatives of the order Octopoda. These ancient marine beauties are in the class Cephalopoda making them relatives of our modern octopus, squid and cuttlefish.
This fellow with his remarkable soft-bodied preservation and inks sack and beak clearly visible is Keuppia levante. He hails from Late Cretaceous (Upper Cenomanian) limestone deposits near Hâdjoula in northwestern Lebanon.
The vampyropod coleoid, Glyphiteuthis abisaadiorum n. sp., is also found at this locality. This specimen is in the collection of David Appleton. Photo credit: David Appleton.