Saturday, 27 June 2020

AMMONITES OF THE VOLGA REGION

The Heteromorph, Jaubertites (Audouliceras) renauxianum
A stunningly beautiful example of the heteromorph ammonite Jaubertites (Audouliceras) renauxianum (d'Orbigny, 1842) from the Volga region in Russia. The Volga region encompasses the drainage basin of the Volga River, the longest river in Europe, in central and southern European Russia. The area is well-known for the beautiful fossil assemblages found here.

These magnificent Jaubertites (Audouliceras) renauxianum heteromorph ammonites are often composites — built with exceptional artful skill from various partial specimens.

We sometimes see them cut in two symmetrical parts and glued into a matrix then doctored up a bit for sale. The practice is frowned upon both scientifically and commercially but continues as does the demand for these exceptional specimens. This beauty is in the collection of José Juárez Ruiz and is complete with some minor restorations. I love these chunky Jaubertites and particularly appreciate the beautiful oil in water colouring in the nacre.

The second photo here shows a lovely busy block of ammonites with Deshayesites volgensis (Sasonova, 1958), and Aconeceras (Sinzovia) trautscholdi (Sinzow. 1870) from Lower Cretaceous, Aptian, (120 - 112 MYA), deposits in the v. Shilovka, Ulyanovsk Region of Russia. This beauty is in the collections of Emil Black. While Emil has counselled me that there are some fundamental challenges with the interpretation of these faunal groups, I will share what is available from the current literature.

Aptian deposits near the Volga River between Ul'yanovsk and Saratov have been studied for more than a century. The area produces some of the most beautiful and sought after ammonite specimens in the world. I've never had the pleasure of collecting in this region but follow the literature and local collectors with enthusiastic interest. Looking at the specimens from here, I'm sure you can appreciate why.

Deshayesites volgensis & Aconeceras trautscholdi
The age of lower Aptian deposits was traditionally established based on changing ammonite assemblages of the family Deshayesitidae. The beauty you see to the right with the lovely ribbing and coloured from cream through to pink and blue is the hallmark species of this area.

But Deshayesitidae are not the only specimens found here. The vast array of heteromorphic ammonites  —  the Ancyloceratidae, inhabitants of relatively deep basins, has made it possible to propose a new scheme of ammonoid zonation in the lower Aptian epipelagic deposits of the Russian plate.

Many of the identified ancyloceratids were established here for the first time. The analysis of coexisting deshayesitids and heteromorphs enables a correlation of stratigraphic schemes for the monomorphic Deshayesitidae and heteromorphic Ancyloceratidae.

The described generic taxa and species are Volgoceratoides I. Michailova et Baraboshkin, gen. nov., V. schilovkensis I. Michailova et Baraboshkin, sp. nov., Koeneniceras I. Michailova et Baraboshkin, gen. nov., K. tenuiplicatum (von Koenen, 1902), K. rareplicatum I. Michailova et Baraboshkin, sp. nov.

In some sections of the Saratov Volga area, specifically in the central part of the Russian Platform, we find both offshore and nearshore lithofacies of the epicontinental Middle Russian Sea. Here we see simultaneous changes in ammonite and belemnite successions that speak to an environmental shift. The significant influence of anoxic events on faunal turnovers in marine communities is well-established. However, many studies are focused on the impact of anoxic conditions on benthic organisms, not on the hunter-gatherers living higher up in the sea column and food chain. For this reason, coeval changes in pelagic cephalopod assemblages remain relatively poorly studied and marginally understood.

Belemnites, represented by the late members of the family Oxyteuthididae, are common in the interval directly preceding the anoxic event, but totally disappear with the onset of the black shale deposition. We see a reduction in the shell size of the Deshayesites ammonites across the mudstone – black shale boundary (maximum shell diameter of adults reduces from ∼20 cm to 7–8 cm).

Some other ammonites become numerous (Sinzovia) within the black shale interval or show the first occurrence in it (Koeneniceras and Volgoceratoides). The diminishing of Deshayesites shell size during the early Aptian OAE may have been caused by palaeoenvironmental factors such as progressive warming and regional input of brackish water.

The significant influence of anoxic events on faunal turnovers in marine communities is well-established. However, many studies are focused on the impact of anoxic conditions on benthic organisms, not on the hunter-gatherers living higher up in the sea column. This means that coeval changes in pelagic cephalopod assemblages remain relatively poorly understood.

Photo: Jaubertites (Audoulicerasrenauxianum (d'Orbigny, 1842) collection of José Juárez Ruiz.
Photo: Deshayesites volgensis (Sasonova, 1958), and Aconeceras (Sinzovia) trautscholdi (Sinzow. 1870) collections of Emil Black. The diameter on the Deshayesites shown here is 70 mm.

Rogov, Mikhail & Shchepetova, Elena & Ippolitov, Alexei & Seltser, Vladimir & Mironenko, Aleksandr & Pokrovsky, Boris & Desai, Bhawanisingh. (2019). Response of cephalopod communities on abrupt environmental changes during the early Aptian OAE1a in the Middle Russian Sea. Cretaceous Research. 10.1016/j.cretres.2019.01.007.

E. Yu. Baraboshkin and I. A. Mikhailova. New Stratigraphic Scheme of the Lower Aptian in the Volga River Middle Courses. Stratigraphy arid Geological Correlation, Vol 10, No 6, 2002, pp 603-626 Translated from Stratigrafiy a Geologicheskaya Korrelyatsiya, Vol 10, No 6, 2002, pp 82-105