Geologically, the Caucasus Mountains belong to a system that extends from southeastern Europe into Asia and is considered a border between them. The Greater Caucasus Mountains are mainly composed of Cretaceous and Jurassic rocks with the Paleozoic and Precambrian rocks in the higher regions.
Some volcanic formations are found throughout the range. On the other hand, the Lesser Caucasus Mountains are formed predominantly of the Paleogene rocks with a much smaller portion of the Jurassic and Cretaceous rocks.
The evolution of the Caucasus began from the Late Triassic to the Late Jurassic during the Cimmerian orogeny at the active margin of the Tethys Ocean while the uplift of the Greater Caucasus is dated to the Miocene during the Alpine orogeny.
The Caucasus Mountains formed largely as the result of a tectonic plate collision between the Arabian plate moving northwards with respect to the Eurasian plate. As the Tethys Sea was closed and the Arabian Plate collided with the Iranian Plate and was pushed against it and with the clockwise movement of the Eurasian Plate towards the Iranian Plate and their final collision, the Iranian Plate was pressed against the Eurasian Plate.
As this happened, the rocks that had been deposited in this basin from the Jurassic to the Miocene were folded to form the Greater Caucasus Mountains. This collision also caused the uplift and the Cenozoic volcanic activity in the Lesser Caucasus Mountains.
The preservation of this Russian specimen is outstanding. Acanthohoplites bigoureti are also found in Madagascar, Mozambique, in the Rhone-Alps of France and the Western High Atlas Mountains and near Marrakech in Morocco. This specimen measures 55mm and is in the collection of the deeply awesome Emil Black.