TRIASSIC OF NORTH AMERICA

In the early 1980s, Tim Tozer, Geological Survey of Canada, looked at the distribution of marine invertebrate fauna in the Triassic of North America.

Tozer's interest in our marine invert friends was their distribution and what those occurrences could tell us. How and when did certain species migrate, cluster, evolve — and for those that were prolific, how could their occurrence — and therefore significance — aide in an assessment of plate and terrane movements that would help us to determine paleolatitudinal significance.

In the western terranes of the Cordillera, marine faunas from southern Alaska and Yukon to Mexico are known from the parts that are obviously allochthonous with regard to the North American plates. Lower and upper Triadic faunas of these areas, as well as some that are today up to 63 ° North, have the characteristics of the lower paleo latitudes. As far as is known, Middle Triadic faunas in these zones do not provide any significant data. In the western Cordillera, the faunas of the lower paleo latitudes can be found up to 3000 km north of their counterparts on the American plate. This indicates a tectonic shift of this magnitude.

There are marine triads on the North American plate over 46 latitudes from California to Ellesmere Island. For some periods, two to three different fauna provinces can be distinguished from one another. The differences in fauna are obviously linked to the paleolatitude. They are called LPL, MPL, HPL (lower, middle, higher paleolatitude). Nevada provides the diagnostic features of the lower; northeastern British Columbia that of the middle and Sverdrup Basin that of the higher paleolatitude. A distinction between the provinces of the middle and the higher paleo-situations can not be made for the lower Triassic and lower Middle Triassic (anise). However, all three provinces can be seen in the deposits of Ladin, Kam and Nor.

Diatoms / Microalgae dominant components of phytoplankton

If one looks at the fauna and the type of sediment, the paleogeography of the Triassic can be interpreted as follows: a tectonically calm west coast of the North American plate that bordered on an open sea; in the area far from the coast, a series of volcanic archipelagos delivered sediment to the adjacent basins. Some were lined or temporarily covered with coral wadding and carbonate banks.

Deeper pools were in between. The islands were probably within 30 degrees of the triadic equator. They moved away from the coast up to about 5000 km from the forerunner of the East Pacific Ridge. The geographical situation west of the back was probably similar.

Jurassic and later generations of the crust from near the back have brought some of the islands to the North American plate; some likely to South America; others have drifted west, to Asia. There are indications that New Guinea, New Caledonia and New Zealand were at a northern latitude of 30 ° or more during the Triassic period. The terranes that now form the western Cordillera were probably welded together and reached the North American plate before the end of the Jurassic period.

Tozer, ET (Tim): Marine Triassic faunas of North America: Their significance for assessing plate and terrane movements. Geol Rundsch 71, 1077-1104 (1982). https://doi.org/10.1007/BF01821119

Danner, W. (Ted): Limestone resources of southwestern British Columbia. Montana Bur. Mines & Geol., Special publ. 74: 171-185, 1976.

Davis, G., Monger, JWH & Burchfiel, BC: Mesozoic construction of the Cordilleran “collage”, central British Columbia to central California. Pacific Coast Paleography symposium 2, Soc. Economic Paleontologists and Mineralogists, Los Angeles: 1-32, 1978.

Gibson, DW: Triassic rocks of the Rocky Mountain foothills and front ranges of northeastern British Columbia and west-central Alberta. Geol. Surv. Canada Bull. 247, 1975.

HYPHANTOCERAS ORIENTALE

A stunning example of the heteromorph ammonite, Hyphantoceras orientale macroconch. This beauty corresponds to 'Morphotype C' from Aiba (2017). 

The specimen is a handful at 136 mm and was lovingly prepared by the hand holding it, that of the talented José Juárez Ruiz.

This an adult specimen (not the juvenile stage) from Upper Santonian outcrops near Ashibetsu, Hokkaido, Japan.

Aiba published on a possible phylogenetic relationship of two species of Hyphantoceras (Ammonoidea, Nostoceratidae) earlier this year, proposing that a phylogenetic relationship may exist based on newly found specimens with precise stratigraphic occurrences in the Kotanbetsu and Obira areas, northwestern Hokkaido.

Two closely related species, Hyphantoceras transitorium and H. orientale, were recognized in the examined specimens from the Kotanbetsu and Obira areas. Specimens of H. transitorium show wide intraspecific variation in the whorl shape. The stratigraphic occurrences of the two species indicate that they occur successively in the Santonian–lowermost Campanian, without stratigraphic overlapping. 

The similarity of their shell surface ornamentations and the stratigraphic relationships possibly suggest that H.orientale was derived from H. transitorium. The presumed lineage is likely indigenous to the northwestern Pacific realm in the Santonian–earliest Campanian. Hyphantoceras venustum and H. heteromorphum might stand outside a H. transitorium–H. orientale lineage, judging from differences of their shell surface ornamentation.

Aiba, Daisuke. (2019). A Possible Phylogenetic Relationship of Two Species of Hyphantoceras (Ammonoidea, Nostoceratidae) in the Cretaceous Yezo Group, Northern Japan. Paleontological Research. 23. 65-80. 10.2517/2018PR010.

SOFT-BODIED PRESERVATION: THE MIGUASHA COLLECTION

An exquisite fossil specimen of an Eusthenopteron Fordi from the upper Devonian (Frasnian), Eescuminac Formation, Miguasha Park, Bay of Heat, Gaspé, Quebec, Canadian Museum of Natural History, Miguasha Collection.

If you look closely at this specimen, you can see the remarkable 3-D and soft-bodied preservation. This fish specimen reminds me of the ray-finned fossil fish you see in carbonate concretion from Lower Cretaceous deposits in the Santana Formation, Brazil.

Eusthenopteron would have shared our ancient seas with the first ammonites and primitive sharks, along with well-established fauna including the trilobites, brachiopods, coral reefs and a whole host of interesting arthropods.

Miguasha National Park / Parc National de Miguasha, is a protected area near Carleton-sur-Mer on the Gaspé Peninsula along the south side of the Saint Lawrence River to the east of the Matapedia Valley in Quebec, Canada. It was created in 1985 by the Government of Quebec and designated as a World Heritage Site in 1999 in honour of paleontological significance for Devonian fish, flower and spore fossils.

These fossils represent five of the six main fossil fish groups recorded from the Devonian (370 million years ago) including specimens of the lobe-finned fish and tetrapods. We see the placoderms, armoured prehistoric fish, in their heyday, dominating almost every known aquatic environment. The Devonian is known as the 'Age of Fishes,' but it could have equally been called the 'Age of Spores,' as this was a time of significant adaptive radiation of terrestrial biota and free-sporing vascular plants. Immense forests carpeted the continents and we see the first of the plant groups evolving leaves, true roots and seeds.

The site was discovered in 1842 by a local geologist and medical doctor, Abraham Gesner. He shared much of his collection with both the British Museum and Royal Scottish Museum for further study.  Other names for this site are the Miguasha Fossil Site, the Bay of Escuminac Fossil Site, the Upper Devonian Escuminac Formation, and the Hugh-Miller Cliffs. It is also sometimes referred to on fossil specimens as 'Scaumenac Bay' or 'Scaumenac Bay P.Q. Photo credit to the deeply awesome John Fam

OSTRACODERMS TO ANGLERFISH

The festive lassie you see here is an Anglerfish. They always look to be celebrating a birthday of some kind, albeit solo. This party is happening deep in our oceans and for those that join in, I hope they like it rough.

The wee candle you see on her forehead is a photophore, a tiny bit of luminous dorsal spine. Many of our sea dwellers have these candle-like bits illuminating the depths. You may have noticed them glowing around the eyes of many of our cephalopod friends. 

These light organs can be a simple grouping of photogenic cells or more complex with light reflectors, lenses, colour filters able to adjust the intensity or angular distribution of the light they produce. Some species have adapted their photophores to avoid being eaten, in others, it's an invitation to lunch.

In anglerfish' world, this swaying light is dead sexy. It's an adaptation used to attract prey and mates alike.

Deep in the murky depths of the Atlantic and Antarctic oceans, hopeful female anglerfish light up their sexy lures. When a male latches onto this tasty bit of flesh, he fuses himself totally. He might be one of several potential mates. She's not picky, just hungry. Lure. Feed. Mate. Repeat.

A friend asked if anglerfish mate for life. Well, yes.... yes, indeed they do.

Mating is a tough business down in the depths. Her body absorbs all the yummy nutrients of his body over time until all that's left are his testes. While unusual, it is only one of many weird and whacky ways our fishy friends communicate, entice, hunt and creatively survive and thrive. The deepest, darkest part of the ocean isn't empty — its hungry.

The evolution of fish began about 530 million years ago with the first fish lineages belonging to the Agnatha, a superclass of jawless fish. We still see them in our waters as cyclostomes but have lost the conodonts and ostracoderms to the annals of time. Like all vertebrates, fish have bilateral symmetry; when divided down the middle or central axis, each half is the same. Organisms with bilateral symmetry are generally more agile, making finding a mate, hunting or avoiding being hunted a whole lot easier.

When we envision fish, we generally picture large eyes, gills, a well-developed mouth. The earliest animals that we classify as fish appeared as soft-bodied chordates who lacked a true spine. While they were spineless, they did have notochords, a cartilaginous skeletal rod that gave them more dexterity than the cold-blooded invertebrates who shared those ancient seas and evolved without a backbone. 

Fish would continue to evolve throughout the Paleozoic, diversifying into a wide range of forms. Several forms of Paleozoic fish developed external armour that protected them from predators. The first fish with jaws appeared in the Silurian period, after which many species, including sharks, became formidable marine predators rather than just the prey of arthropods.

Fish in general respire using gills, are most often covered with bony scales and propel themselves using fins. There are two main types of fins, median fins and paired fins. The median fins include the caudal fin or tail fin, the dorsal fin, and the anal fin. Now there may be more than one dorsal, and one anal fin in some fishes.

The paired fins include the pectoral fins and the pelvic fins. And these paired fins are connected to, and supported by, pectoral and pelvic girdles, at the shoulder and hip; in the same way, our arms and legs are connected to and supported by, pectoral and pelvic girdles. This arrangement is something we inherited from the ancestors we share with fish. They are homologous structures.

When we speak of early vertebrates, we are often talking about fish. Fish is a term we use a lot in our everyday lives but taxonomically it is not all that useful. When we say, fish we generally mean an ectothermic, aquatic vertebrate with gills and fins.

Fortunately, many of our fishy friends have ended up in the fossil record. We may see some of the soft bits from time to time, as in the lovely fossil fish found in concretion in Brazil, but we often see fish skeletons. Vertebrates with hard skeletons had a much better chance of being preserved. 

Eohiodon Fish, McAbee Fossil Beds

In British Columbia, we have lovely two-dimensional Eocene fossil fish well-represented from the Allenby of Princeton and the McAbee Fossil Beds. 

We have the Tiktaalik roseae, a large freshwater fish, from 375 million-year-old Devonian deposits on Ellesmere Island in Canada's Arctic. Tiktaalik is a wonderfully bizarre creature with a flat, almost reptilian head but also fins, scales and gills. We have other wonders from this time. 

Canada also boasts spectacular antiarch placoderms, Bothriolepsis, found in the Upper Devonian shales of Miguasha in Quebec.

There are fragments of bone-like tissues from as early as the Late Cambrian with the oldest fossils that are truly recognizable as fishes come from the Middle Ordovician from North America, South America and Australia. At the time, South America and Australia were part of a supercontinent called Gondwana. North America was part of another supercontinent called Laurentia and the two were separated by deep oceans.

These two supercontinents and others that were also present were partially covered by shallow equatorial seas and the continents themselves were barren and rocky. Land plants didn't evolve until later in the Silurian Period. In these shallow equatorial seas, a large diverse and widespread group of armoured, jawless fishes evolved: the Pteraspidomorphi. The first of our three groups of ostracoderms. The Pteraspidomorphi are divided into three major groups: the Astraspida, Arandaspida and the Heterostraci.

The oldest and most primitive pteraspidomorphs were the Astraspida and the Arandaspida. You'll notice that all three of these taxon names contain 'aspid', which means shield. This is because these early fishes — and many of the Pteraspidomorphi — possessed large plates of dermal bone at the anterior end of their bodies. This dermal armour was very common in early vertebrates, but it was lost in their descendants. 

Arandaspida is represented by two well-known genera: Sacabampaspis, from South America and Arandaspis from Australia. Arandaspis have large, simple, dorsal and ventral head shields. Their bodies were fusiform, which means they were shaped sort of like a spindle, fat in the middle and tapering at both ends. Picture a sausage that is a bit wider near the centre with a crisp outer shell.

If you're a keen bean to see an anglerfish that recently washed up on the shores of Newport Beach this past May, hit this link: https://www.theguardian.com/us-news/2021/may/11/deep-sea-anglerfish-california-beach-finding-nemo. Kudos for my colleague, Giovanni, bringing this gloriously horrific lovely to my attention. 

ANCIENT SEAS: HOKKAIDO

A very beautiful Lower Campanian block from Haroto, Hokkaido, Japan. This specimen contains an ancient undersea world at a glance.

The beautiful block you see here was prepared, photographed and is in the collections of José Juárez Ruiz. In it, you can see a lovely Pseudoxybeloceras (Parasolenoceras) soyaense (143 mm), Polyptychoceras jimboi (134 mm), Polyptychoceras sp. (114 mm), Gaudryceras mite (48 and 45 mm), Gaudryceras tenuiliratum (Hirano, 1978) at (48 and 20 mm), and a wee fragment of wood (69 mm).

Matsumoto published on the ammonites from the Campanian (Upper Cretaceous) of northern Hokkaido back in 1984, in the Palaeontological Society of Japan Special Series Papers, Number #27.

This was my first look at the glorious fauna from northern Japan. The species and preservation are truly outstanding. Since then, many of the Japanese palaeontologists have made their way over to Vancouver Island, to look at ammonites, inoceramids and coleoid jaws from the Nanaimo Group and compare them to the Japanese species.

Rick Ross and Pat Trask, both of Courtenay on Vancouver Island, collaborated with Dr. Kazushige Tanabe and Yoshinori Hikida of Japan, to produce a wonderful paper in the Journal of Paleontology, 82 (2), 2008, pp 398-408, on Late Cretaceous Octobrachiate Coleoid Lower Jaws from the North Pacific Regions. They compared eight well-preserved cephalopod jaws from Upper Cretaceous (Santonian and Campanian) deposits of Vancouver Island, Canada, and Hokkaido, Japan. Seven of these were from Santonian to lower Campanian strata of the Nanaimo Group in the northeastern region of Vancouver Island. The eighth specimen was from Santonian strata of the Yezo Group in the Nakagawa area, northern Hokkaido, Japan. 

While they were collaborating on identifying coleoid jaws from the Comox Valley, Rick was visited twice by Dr. Kazushige Tanabe who was joined by his colleague Akinori Takahashi. Takahashi is an expert on temporal species-diversity changes in Japanese Cretaceous inoceramid bivalves.

They had the very great pleasure of visiting many fossil sites and seeing personal and museum collections. If you'd like to read Matsumoto's paper, here is the link: http://www.palaeo-soc-japan.jp/download/SP/SP27.pdf  I have a pdf copy of the Coleoid paper from Rick. It has very nice photos and illustrations, including a drawing of the holotypes of Paleocirroteuthis haggerti n. gen. and Paleocirroteuithis pacifica.

Here's a link to one of Takahashi's papers: https://bioone.org/journals/paleontological-research/volume-9/issue-3/prpsj.9.217/Diversity-changes-in-Cretaceous-inoceramid-bivalves-of-Japan/10.2517/prpsj.9.217.short

ANCIENT OCTOPUS: KEUPPIA

A sweet as you please example of Keuppia levante (Fuchs, Bracchi & Weis, 2009), an extinct genus of octopus that swam our ancient seas back in the Cretaceous. 

The dark black and brown area you see here is his ink sac which has been preserved for a remarkable 95 million years.

This cutie is in the family Palaeoctopodidae, and one of the earliest representatives of the order Octopoda — and perhaps my favourite fossil. It was this perfect specimen that inspired the logo for the Fossil Huntress brand.  

These ancient marine beauties are in the class Cephalopoda making them relatives of our modern octopus, squid and cuttlefish.

There are two species of Keuppia, Keuppia hyperbolaris and Keuppia levante, both of which we find as fossils. We find their remains, along with those of the genus Styletoctopus, in Cretaceous-age Hâqel and Hjoula localities in Lebanon. 

For many years, Palaeoctopus newboldi (Woodward, 1896) from the Santonian limestones at Sâhel Aalma, Lebanon, was the only known pre‐Cenozoic coleoid cephalopod believed to have an unambiguous stem‐lineage representative of Octobrachia fioroni. 

With the unearthing of some extraordinary specimens with exquisite soft‐part preservation in the Lebanon limestones, our understanding of ancient octopus morphology has blossomed. The specimens are from the sub‐lithographical limestones of Hâqel and Hâdjoula, in northwestern Lebanon. These localities are about 15 km apart, 45 km away from Beirut and 15 km away from the coastal city of Jbail. Fuchs et al. put a nice little map in their 2009 paper that I've included and referenced here.

Palaeoctopus newboldi had a spherical mantle sac, a head‐mantle fusion, eight equal arms armed with suckers, an ink sac, a medially isolated shell vestige, and a pair of (sub‐) terminal fins. The bipartite shell vestige suggests that Palaeoctopus belongs to the octopod stem‐lineage, as the sister taxon of the Octopoda, the Cirroctopoda, is characterized by an unpaired clasp‐like shell vestige (Engeser 1988; Haas 2002; Bizikov 2004).

It is from the comparisons of Canadian fauna combined with those from Lebanon and Japan that things really started to get interesting with Octobrachia. Working with fossil specimens from the Campanian of Canada, Fuchs et al. (2007a ) published on the first record of an unpaired, saddle‐shaped shell vestige that might have belonged to a cirroctopod. 

Again from the Santonian–Campanian of Canada and Japan, Tanabe et al. (2008) reported on at least four different jaw morphotypes. Two of them — Paleocirroteuthis haggarti (Tanabe et al., 2008) and Paleocirroteuthis Pacifica  (Tanabe et al ., 2008) — have been interpreted as being of cirroctopod type, one of octopod type, and one of uncertain octobrachiate type. 

Interestingly Fuchs et al. have gone on to describe the second species of Palaeoctopus, the Turonian Palaeoctopus pelagicus from limestones at Vallecillo, Mexico. While more of this fauna will likely be recovered in time, their work is based solely on a medially isolated shell vestige.

Five new specimens have been found in the well-known Upper Cenomanian limestones at Hâqel and Hâdjoula in Lebanon that can be reliably placed within the Octopoda. Fuchs et al. described these exceptionally well‐preserved specimens and discuss their morphology in the context of phylogeny and evolution in their 2008 paper (2009 publishing) in the Palaeontology Association Journal, Volume 51, Issue 1.

The presence of a gladius vestige in this genus shows a transition from squid to octopus in which the inner shell has divided into two parts in early forms to eventually be reduced to lateralized stylets, as can be seen in Styletoctopus.

The adorable fellow you see here with his remarkable soft-bodied preservation and inks sack and beak clearly visible is Keuppia levante. He hails from Late Cretaceous (Upper Cenomanian) limestone deposits near Hâdjoula, northwestern Lebanon. The vampyropod coleoid, Glyphiteuthis abisaadiorum n. sp. is also found at this locality. This specimen is about 5 cm long.

Fuchs, D.; Bracchi, G.; Weis, R. (2009). "New octopods (Cephalopoda: Coleoidea) from the Late Cretaceous (Upper Cenomanian) of Hâkel and Hâdjoula, Lebanon". Palaeontology. 52: 65–81. doi:10.1111/j.1475-4983.2008.00828.x.

Photo one: Fossil Huntress. Figure Two: Topographic map of north‐western Lebanon with the outcrop area in the upper right-hand corner. Fuchs et al, 2009.  

BRACHIOPODA: LOPHOTROCHOZOANS

Looking down at the pebbly sand, you see just the wee top of this lovely fossil brachiopod poking out. Glee, delight and wonder follow as you roll it over in your hands and notice how it differs from clams you may be more familiar with.

Clams or bivalves are molluscs, the second-largest phylum of invertebrates with about 85,000 extant species. While brachiopods share some similarities with their molluscan friends they are in a phylum all their own.

Brachiopods are small marine shellfish that are not so common today but back in the Palaeozoic they were plentiful the world over. The two valves that make up a brachiopods shell are of different sizes and if you look closely you'll see that the hinge runs top and bottom  -- versus left and right like a clam.

Brachiopods had evolved from an ancestor similar to Halkieria, a slug-like Cambrian animal with "chain mail" on its back and a shell at the front and rear end; it was thought that the ancestral brachiopod converted its shells into a pair of valves by folding the rear part of its body under its front. 

New fossils found in the last few years have shown us that the "chain mail" of tommotiids formed the tube of a sessile animal; one tommotiid resembled phoronids, which are close relatives or a subgroup of brachiopods, while the other tommotiid bore two symmetrical plates that might be an early form of brachiopod valves. Lineages of brachiopods that have both fossil and extant taxa appeared in the early Cambrian, Ordovician, and Carboniferous periods, respectively. 

Other lineages have arisen and then become extinct, sometimes during severe mass extinctions. At their peak in the Paleozoic era, the brachiopods were among the most abundant filter-feeders and reef-builders and occupied other ecological niches, including swimming in the jet-propulsion style of scallops. Brachiopod fossils have been useful indicators of climate changes during the Paleozoic. However, after the Permian–Triassic extinction event, brachiopods recovered only a third of their former diversity. 

Brachiopods were especially vulnerable to the Permian–Triassic extinction, as they built calcareous hard parts — made of calcium carbonate — and had low metabolic rates and weak respiratory systems. It was often thought that brachiopods went into decline after the Permian–Triassic extinction, and were out-competed by bivalves, but both brachiopod and bivalve species increased from the Paleozoic to modern times, with bivalves increasing faster; after the Permian–Triassic extinction, brachiopods became for the first time less diverse than bivalves.

Their lineage dates back to the Cambrian with over 12,000 fossil species and 350 living relatives sorted between 6,000 genera. There are two major groups of brachiopods, articulate with toothed hinges and simple open and closing muscles to manipulate their shells and inarticulate brachiopods with untoothed hinges and a more complex set of muscles used to control the brachial supports used to open and close their shells.

AMAZING AMMONITE: HOLCOPHYLLOCERAS

What is wonderful about natural science is exploring new species. Take a look at this tremendously robust suturing on this lovely ammonite, Holcophylloceras mediterraneum, (Neumayr, 1871) from Late Jurassic (Oxfordian) deposits near Sokoja, Madagasgar. This particular specimen and post goes out to Susan Gerard who has provided lovely cabinetry that will become home for so many of these wonderfully preserved specimens.  

Madagascar is a treasure trove of outstanding fossil species and this Holcophylloceras ammonite is no exception.

The shells had many chambers divided by walls called septa. The chambers were connected by a tube called a siphuncle which allowed for the control of buoyancy with the hollow inner chambers of the shell acting as air tanks to help them float.

We can see the edges of this specimen's shell where it would have continued out to the last chamber, the body chamber, where the ammonite lived. Picture a squid or octopus, now add a shell and a ton of water.

LOWER LIAS LYTOCERAS

A superbly prepped and extremely rare Lytoceras (Suess, 1865) ammonite found as a green ammonite nodule by Matt Cape in the Lower Lias of Dorset. 

Lytoceras are rare in the Lower Lias of Dorset — apart from the Belemnite Stone horizon — so much so that Paul Davis, whose skilled prep work you see here, initially thought it might be a Becheiceras hidden within the large, lumpy nodule. 

One of the reasons these lovelies are rarely found from here is that they are a Mediterranean Tethyian genus. The fossil fauna we find in the United Kingdom are dominated by Boreal Tethyian genera. 

We do find Lytoceras sp. in the Luridum subzone of the Pliensbachian showing that there was an influx of species from the Mediterranean realm during this time. This is the first occurrence of a Lytoceras that he has ever seen in a green nodule and Paul's seen quite a few. 

This absolutely cracking specimen was found and is in the collections of the awesome Matt Cape. Matt recognized that whatever was hidden in the nodule would take skilled and careful preparation using air scribes. Indeed it did. It took more than five hours of time and skill to unveil the lovely museum-worthy specimen you see here. 

We find Lytoceras in more than 1,000 outcrops around the globe ranging from the Jurassic through to the Cretaceous, some 189.6 to 109.00 million years ago. Once this specimen is fully prepped with the nodule material cut or scraped away, you can see the detailed crinkly growth lines or riblets on the shell and none of the expected coarse ribbing. 

Lytoceras sp. Photo: Craig Chivers

If you imagine running your finger along these, you would be tracing the work of decades of growth of these cephalopods. 

While we cannot know their actual lifespans, but we can make a healthy guess. 

The nautilus, their closest living cousins live upwards of 20 years — gods be good — and less than three years if conditions are poor.

The flanges, projecting flat ribs or collars, develop at the edge of the mouth border on the animal's mantle as they grow each new chamber. 

Each delicate flange grows over the course of the ammonites life, marking various points in time and life stages as the ammonite grew. There is a large variation within Lytoceras with regards to flanges. They provide both ornamentation and strength to the shell to protect it from water pressure as they moved into deeper seas.

The concretion prior to prep

This distinctive genus with its evolute shells are found in the Cretaceous marine deposits of: 

Antarctica (5 collections), Austria (19), Colombia (1), the Czech Republic (3), Egypt (2), France (194), Greenland (16), Hungary (25), Italy (11), Madagascar (2), Mexico (1), Morocco (4), Mozambique (1), Poland (2), Portugal (1), Romania (1), the Russian Federation (2), Slovakia (3), South Africa (1), Spain (24), Tanzania (1), Trinidad and Tobago (1), Tunisia (25); and the United States of America (17: Alaska, California, North Carolina, Oregon).

We also find them in Jurassic marine outcrops in:

Austria (15), Canada (9: British Columbia), Chile (6), France (181), Germany (11), Greenland (1), Hungary (189), India (1), Indonesia (1), Iran (1), Italy (50), Japan (14), Kenya (2), Luxembourg (4), Madagascar (2), Mexico (1), Morocco (43), New Zealand (15), Portugal (1), Romania (5), the Russian Federation (1), Slovakia (1), Spain (6), Switzerland (2), Tunisia (11), Turkey (12), Turkmenistan (1), Ukraine (5), the United Kingdom (12), United States (11: Alaska, California) — in at least 977 known collections. 

References:

Sepkoski, Jack (2002). "A compendium of fossil marine animal genera (Cephalopoda entry)". Bulletins of American Paleontology. 363: 1–560. Archived from the original on 2008-05-07. Retrieved 2017-10-18.

Paleobiology Database - Lytoceras. 2017-10-19.

Systematic descriptions, Mesozoic Ammonoidea, by W.J Arkell, Bernhard Kummel, and C.W. Wright. 1957. Treatise on Invertebrate Paleontology, Part L. Geological Society of America and University of Kansas press.

PORT HARDY: TIME AND TIDE

One of the most beautiful areas of Vancouver Island is the town of Port Hardy on the north end of the island. 

Just outside Port Hardy further south on the west coast is the area known as Fort Rupert or Tsaxis. 

It was here that the Hudson's Bay Company built Fort Rupert both for trade with the local First Nation population and the allure of potential coal deposits. 

I headed up to the north island this past week to stomp around my old haunts, visit with family and get in a bit of late season kayaking. The town was much as I remembered it. There have been changes, of course. I lived up on Wally's hill above the reserve at Tsaxis beside the old cemetery. 

My wee childhood home is still there and I am very pleased to see that the earthly home of my ancestors is well maintained. The cemetery is groomed and cared for but the land surrounding it is overgrown and it took me a few minutes to orient myself to see where things used to be. Where the old Hudson's Bay Company Fort and its iconic chimney were in relation to the graveyard. 

A lifetimes worth of memories came flooding back. Those from my earliest years and then later when I returned to kayak, fish and scuba dive in these rich waters.

My plans of blissful days kayaking and taking photos of the scenery were altered by hurricane-force winds. Still beautiful, but chilly and choppy.

The beachhead here was clocking 120 km winds so I did a brief visit to the homestead, the graveyard and Jokerville then headed home to light the fire and hunker in as the storm blew through. 

Port Harty and Fort Rupert have an interesting history and how you read it or hear it truly depends on the lens that is applied. This has been the ancestral home to many First Nation groups. Mostly they were passing through and coming here to dig up delicious butter clams, roots, berries and other natural yummy goodness. Years before Port Hardy was settled at the turn of the century it was the home to the Kwakiutl or Kwagu’ł and part of my heritage. 

Alec and Sarah Lyon operated a store and post office on the east side of Hardy Bay. A 1912 land deal promoted by the Hardy Bay Land Co., put the area on the map and increased its population. By 1914, 12 families had settled, built a school, sawmill, church and hotel. 

The community of Port Hardy is situated within traditional Kwagu’ł First Nation territory. It is also home to the Gwa’sala-‘Nakwaxda’xw First Nation. In 1964 all the First Nations communities were amalgamated and forced to relocate from their traditional territories by the federal government, for administrative reasons. 

The First Nation families were told that it would cost less for education, easier for medical help, and the government would help with housing, but it turned out to be a hidden agenda designed to assimilate the various groups into Canadian society — or face extermination. Several years of threats and promises later, the Gwa’sala and ‘Nakwaxda’xw reluctantly gave in to the relocation, but the government didn’t keep their promise for adequate housing. 

There were five homes for over 200 people on the Tsulquate Reservation. The Gwa’sala traditional territory is Smith Inlet and surrounding islands. ‘Nakwaxda’xw traditional territory is Seymour Inlet, the Deserter’s Group, Blunden Harbour, and surrounding islands.

There was limited access to the community until the logging road connecting Port Hardy to Campbell River was paved in December of 1979.

Port Hardy’s population grew to a little over 5,000 residents during the Island Copper Mine years (1971-1995). The former mine site is located 16 kilometres south of Port Hardy on the shores of Rupert Inlet. The open-pit porphyry copper mine employed over 900 employees from Port Hardy and the surrounding communities. Today, the former mine has been transformed into a wildlife habitat and pit lake biological treatment system (BHP Copper Inc., 2010). The Quatsino First Nation manage the property and their Economic Development Board is exploring options for its use. 

The Quatsino First Nations have conducted several feasibility studies around the implementation of a puck or brickett mill onsite, utilizing the existing infrastructure, which includes six industrial buildings.

Today, Port Hardy serves as the crossroads for air, ferry and marine transportation networks, and serves as the gateway to the fast-growing Central Coast, the Cape Scott and North Coast Trails, and BC Ferry’s northern terminus for the Discovery Coast run and Prince Rupert. It supports several traditional and emerging sectors and remains rich in natural resources and community spirit.

Every corner of the Port Hardy region is enriched with culture and history. Starting with the two welcome poles in Carrot Park, both carved and replicated by Calvin Hunt, a Kwagu’ł artist who is based in Tsax̱is. 

From here and along the seawall are interpretive signs with Kwak’wala words for various wildlife, such as salmon, bear, wolf, and orca. At the end of this walk is Tsulquate Park. 

From here you can see across Queen Charlotte Strait; the ocean highway and lands of the Kwakwa̱ka̱ʼwakw. Port Hardy was named after Vice-Admiral Sir Thomas Masterman Hardy (5 April 1769 – 20 September 1839) who served as the captain of H.M.S. Victory in the Royal Navy. 

He served at the Battle of Trafalgar and held Lord Nelson at the end of that battle where Nelson died in his arms. Though he never visited this island community, it bears his name today. 

A ten-minute drive from downtown Port Hardy, in the neighbouring community of Fort Rupert, is the village of Tsax̱is. This is the current home of the Kwagu’ł First Nation. Here lies elaborated totem poles and the big house; a venue where First Nations ceremonies take place, such as the potlatch. 

The potlatch is a First Nations constitution that determines our politics, our government, our education, our medicine, our territory, and our jurisdiction. Potlatch is a complex event with several ceremonies, which are still practiced in buildings like the Tsax̱is big house.

On the front porch of the village of Tsax̱is is Tayaguł (Storey’s Beach). Along this waterfront were several villages, which are depicted on map (pictured below) by Mervyn Child, a Kwagu’ł artist. 

Across the way and middle of K’ak’a (Beaver Harbour) are Atłanudzi (Cattle Island), Ḵ’ut’sa̱dze (Peel Island), Ḵ’a̱msa̱x̱tłe (Shell Island), and Uxwiwe’ (Deer Island). Once the words are broken down and translated; the names of these islands are unique to their environment, as they’re part of a story that belongs to the Kwagu’ł.

Where: Port Hardy, British Columbia. 50°43'27"N, 127°29'52"W

DACTYLIOCERAS AMMONITE PREPPED BY HARRY TABINER

Dactylioceras ammonite, Photo: Harry Tabiner

A lovely Dactylioceras ammonite from the Lower Jurassic Upper Lias Holderness of the Yorkshire Coast. This beauty measures over 8cm with especially attractive colouring.

Holderness is an area of the East Riding of Yorkshire, on the east coast of England. An area of rich agricultural land, Holderness was marshland until it was drained in the Middle Ages. Topographically, Holderness has more in common with the Netherlands than with other parts of Yorkshire. To the north and west are the Yorkshire Wolds.

Geologically, Holderness is underlain by Cretaceous chalk but in most places, it is so deeply buried beneath glacial deposits that it has no influence on the landscape.

The landscape is dominated by deposits of till, boulder clays and glacial lake clays. These were deposited during the Devensian glaciation. The glacial deposits form a more or less continuous lowland plain which has some peat filled depressions (known locally as meres) which mark the presence of former lake beds. There are other glacial landscape features such as drumlin mounds, ridges and kettle holes scattered throughout the area.

Dactylioceras ammonite, Photo: Harry Tabiner

The well-drained glacial deposits provide fertile soils that can support intensive arable cultivation. Fields are generally large and bounded by drainage ditches. There is very little woodland in the area and this leads to a landscape that is essentially rural but very flat and exposed. The coast is subject to rapid marine erosion.

The Geology of Yorkshire in northern England shows a very close relationship between the major topographical areas and the geological period in which their rocks were formed. The rocks of the Pennine chain of hills in the west are of Carboniferous origin whilst those of the central vale are Permo-Triassic.

The North York Moors in the north-east of the county are Jurassic in age while the Yorkshire Wolds to the southeast are Cretaceous chalk uplands. The plain of Holderness and the Humberhead levels both owe their present form to the Quaternary ice ages.

The strata become gradually younger from west to east. Much of Yorkshire presents heavily glaciated scenery as few places escaped the direct or indirect impact of the great ice sheets as they first advanced and then retreated during the last ice age. This beauty is in the collection of the deeply awesome Harry Tabiner.

HARRISON LAKE: FOSSILS AND THE BEATING HEART

Located three hours east of Vancouver, most folks head to Harrison Lake to enjoy its crisp waters, soak in the hot springs, camp or four-wheel-drive immersed in rugged scenery, or look for the elusive Sasquatch reported to live in the area. 

But there are some who come to Harrison Lake and miss the town entirely. Instead, they favour the upper west side of the lake and the fossiliferous bounty found here.

Indeed, this is the perfect location for local citizen scientists to strut their stuff. Harrison is a perfect family day trip, where you can discover wonderful marine fossil specimens as complete or partially crushed fossilized shells embedded in rock. 

It is truly amazing that we can find them at all. These beauties range in age from Jurassic to Cretaceous, with most being Lower Callovian, meaning the ammonites here swam our ancient oceans more than 160 million years ago. 

The area around Harrison Lake has been home to the Sts’ailes, a sovereign Coast Salish First Nation for thousands of years. Sts’ailes’ means, “the beating heart,” and it sums up this glorious wilderness perfectly. They describe their ancient home as Xa’xa Temexw or Sacred Earth. 

With the settling of Canada, Geologists began exploring the area in the 1880s, calling upon the Sts’ailes to help them look for coal and a route for the Canadian Pacific Railway. Coal was the aim, but happily, they also found fossils. Sacred Earth, indeed.  

Belemnite Fossils

In my favourite outcrops, you can find large, smooth inflated Jurassic ammonites along with their small grey and brown cousins. 

Further up the road, you will see Cretaceous cigar-shaped squid-like cephalopods called Belemnites, and the bivalve (clam) Buchia — gifts deposited by glaciers. Here are the most common.

Ammonites

Almost all of the ammonite specimens found near Harrison Lake are the toonie sized Cadoceras (Paracadoceras) tonniense with well-preserved outer whorls but flattened inner whorls. We find semi-squished elliptical specimens here, too. If you see a large, smooth, inflated grapefruit-sized ammonite, you are holding a rare prize — a Cadoceras comma ammonite, the macroconch or female of the species.  

Ammonites were predatory, squid-like creatures that lived inside coil-shaped shells. Like other cephalopods, ammonites had sharp, beak-like jaws inside a ring of squid-like tentacles that extended from their shells. They used these tentacles to snare prey — plankton, vegetation, fish and crustaceans — similar to the way a squid or octopus hunts today.

Within their shells, ammonites had a number of chambers called septa filled with gas or fluid, and they were interconnected through a wee air tube. By pushing air in or out, they were able to control their buoyancy. 

These small but mighty marine predators lived in the last chamber of their shell and continuously built new shell material as they grew. As they added each new chamber, they would move their squid-like body down to occupy the final outside chamber.

Interestingly, ammonites from Harrison Lake are quite similar to the ones found within the lower part of the Chinitna Formation near Cook Inlet, Alaska, and Jurassic Point, Kyuquot, on the west coast of Vancouver Island — some of the most beautiful places on Earth. 

Buchia (bivalve) Clams

The bivalve or clam Buchia are commonly found at Harrison Lake. You will see them cemented together en masse. . They populated Upper Jurassic–Lower Cretaceous waters like a team sport. When they thrived, they really thrived, building up large coquinas of material. Large boulders of Buchia cemented together en masse hitched a ride with the glaciers and were deposited around Harrison Lake. Some kept going and we find similar erratics or glacier-deposited boulders as far south as Washington state. 

Buchia is used as Index Fossils. Index fossils help us to figure out the age of the rock we are looking at because they are abundant, populate an area en masse, and then die out quickly. In other words, they make it easy to identify a geologic time span.

So what does this mean to you? Now, when you are out and about with friends and discover rocks with Buchia, or made entirely of Buchia, you can say, “Oh, this looks to be Upper Jurassic or Lower Cretaceous. Come take a look! We're likely the first to lay eyes on this little clam since dinosaurs roamed the Earth.” 

Fossil Collecting at Harrison Lake Fossil Field Trip — Getting there

This Harrison Lake site is a great day trip from Vancouver or the Fraser Valley. You will need a vehicle with good tires for travel on gravel roads. Search out the route ahead of time and share your trip plan with someone you trust. If you can pre-load the Google Earth map of the area, you will thank yourself. 

Heading east on from Vancouver, it will take you 1.5-2 hours to reach Harrison Mills. 

Access Forestry Road #17 at the northeast end of the parking lot from the Sasquatch Inn at 46001 Lougheed Hwy, Harrison Mills. From there, it will take about an hour to get to the site. Look for signs for the Chehalis River Fish Hatchery to get you started. 

Drive 30 km up Forestry Road #1, and stop just past Hale Creek at 49.5° N, 121.9° W (paleo-coordinates 42.5° N, 63.4° W) on the west side of Harrison Lake. You will see Long Island to your right. 

The first of the yummy fossil exposures are just north of Hale Creek on the west side of the road. Keep in mind that this is an active logging road, so watch your kids and pets carefully. Everyone should be wearing something bright so they can be easily spotted.

How to Spot the Fossils

The fossils here are easily collected—look in the bedrock and in the loose material that gathers in the ditches. Specimens will show up as either dark grey, grey-brown or black. Look for the large, dark-grey boulders the size of smart cars packed with Buchia. 

And while you are at it, be on the lookout for anything that looks like bone. This site is also ripe for marine reptiles—think plesiosaur, mosasaur and elasmosaur. As a citizen scientist and budding palaeontologist, you might just find something new!

What to Know Before You Go

Fill your gas tank and pack a tasty lunch. As with all trips into British Columbia's wild places, dress for the weather. You will need hiking boots, rain gear, gloves, eye protection, and a good geologic hammer and rock (cold) chisel. 

Wear bright clothing and keep your head covered. Slides are common, and you may start a few if you hike the cliffs. If you are with a group, those collecting below may want to consider hardhats in case of rockfall — chunks of rock the size of your fist up to the size of a grapefruit. They pack a punch. 

Bring a colourful towel or something to put your keepers on. Once you set rock down, it can be hard to find again given the terrain. I take the extra precaution of spraying the ends of my hammers and chisels with yellow fluorescent paint, as I have lost too many in the field. You will also want to bring a camera for the blocks of Buchia that are too big to carry home. 

Identifying Your Treasures

When you have finished for the day, compare your treasures to see which ones you would like to keep. In British Columbia, you are a steward of the fossil, which means they belong to the province, but you can keep them safe. You are not allowed to sell or ship them outside British Columbia without a permit. 

Once you get home, wash and identify your finds. Harrison Lake does not have a large variety of fossil fauna, so this should not be difficult. If your find is coiled and round, it is an ammonite. If it is long and straight, it is a belemnite. And if it looks like a wee fat baby oyster, it is Buchia. This is not always true, but mostly true.

What about collecting fossils in all seasons?. Everyone has a preference. I prefer not to collect in the snow, but I have done it. While sunny days are lovely, it can also be easier to see the specimens when the rock is wet. So, do we do this in the rain? Heck, yeah! 

In torrential rain? 

Yes — once you are hooked, but for your casual friends or the kiddos, the answer is likely no. Choose your battles. They may come with you, but a cold day getting soaked is no fun. 

In time, you will find your inner fossil geek — probably with your first find. And that's just the tip of the iceberg. First, it will be you, then your kids, your friends and then your neighbour. Once you start, it is easy to get hooked. Fossil addiction is real, and the only cure is to get out there and do it some more. You've got this!

References and further information:

A. J. Arthur, P. L. Smith, J. W. H. Monger and H. W. Tipper. 1993. Mesozoic stratigraphy and Jurassic palaeontology west of Harrison Lake, southwestern British Columbia. Geological Survey of Canada Bulletin 441:1-62

R. W. Imlay. 1953. Callovian (Jurassic) ammonites from the United States and Alaska Part 2. The Alaska Peninsula and Cook Inlet regions. United States Geological Survey Professional Paper 249-B:41-108

An overview of the tectonic history of the southern Coast Mountains, British Columbia; Monger, J W H; in, Field trips to Harrison Lake and Vancouver Island, British Columbia; Haggart, J W (ed.); Smith, P L (ed.). Canadian Paleontology Conference, Field Trip Guidebook 16, 2011 p. 1-11 (ESS Cont.# 20110248).

JELLYFISH: GAGISAMA

These festive lovelies are jellyfish. Jellyfish are found all over the world, from surface waters to our deepest seas — and they are old. They are some of the oldest animals in the fossil record.

Sea jellies and jellyfish are the common names for the medusa-phase or adult phase of certain gelatinous members of the subphylum Medusozoa, a major part of the phylum Cnidaria — more closely related to anemones and corals.

Jellyfish are not fish at all. Jellyfish evolved millions of years before true fish. 

The oldest conulariid scyphozoans — picture an ice-cream cone with fourfold symmetry — appeared between 635 and 577 million years ago in the Neoproterozoic of the Lantian Formation a 150-meter-thick sequence of rocks deposited in southern China. 

Others are found in the youngest Ediacaran rocks of the Tamengo Formation of Brazil, c. 505 mya, through to the Triassic. Cubozoans and hydrozoans appeared in the Cambrian of the Marjum Formation in Utah, USA, c. 540 mya. Like other soft-bodied organisms, ctenophores (comb jellies), sea jellies and jellyfish only produce fossils only under exceptional taphonomic conditions — think rare.

I have seen all sorts of their brethren growing up on the west coast of Canada. I have seen them in tide pools, washed up on the beach and swam amongst thousands of Moon Jellyfish while scuba diving in the Salish Sea. Their movement in the water is marvellous.  

In the Kwak̓wala language of the Kwakiutl or Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, jellyfish are known as ǥaǥisama.

The watercolour ǥaǥisama you see here is a bit of fancy. While I chose blue, purple and pink for these lovelies, they also come in bright yellow, orange and relatively clear — and are often luminescent.

Jellyfish such as comb jellies produce bright flashes to startle a predator, others such as siphonophores can produce a chain of light or release thousands of glowing particles into the water as a mimic of small plankton to confuse the predator.

For most jellyfish bioluminescence is used for defence against predators — and about half of all jellyfish are bioluminescent. Some produce a glowing sticky slime that clings to predators making them vulnerable to other predators. Some jellyfish can release their tentacles as glowing decoys. So you see that there are many strategies for using bioluminescence by jellyfish.

All bioluminescence comes from energy released from a chemical reaction. This is very different from other sources of light, such as from the sun or a light bulb, where the energy comes from heat. In a luminescent reaction, two types of chemicals, called luciferin and luciferase, combine together. The luciferase acts as an enzyme, allowing the luciferin to release energy as it is oxidized. The colour of the light depends on the chemical structures of the chemicals. 

There are more than a dozen known chemical luminescent systems, indicating that bioluminescence evolved independently in different groups of organisms. One type of luciferin is called coelenterazine, found in jellyfish, shrimp, and fish. Dinoflagellates and krill share another class of unique luciferins, while ostracods (firefleas) and some fish have a completely different luciferin. The occurrence of identical luciferins for different types of organisms suggests a dietary source for some groups. Organisms such as bacteria and fireflies have unique luminescent chemistries. In many other groups, the chemistry is still unknown

Some of the most amazing deep-sea jellyfish are the comb jellies, which can get as large as a basketball, and are in some cases so fragile that they are almost impossible to collect intact.

Also spectacular are the siphonophores, some of which can reach several meters in length. Siphonophores deploy many tentacles like a gill net casting for small fish.

HETEROPTERA: SNEAKOPTERA

A sweet little water bug from the suborder Heteroptera (Latreille, 1810). He looks more like a cartoon character that any other specimen I've seen. 

This fun fellow is in the collections of Tim Dingman. The deeply awesome Jim Barkley gets credit for this charming photo. The cartoon effect comes from this guy missing his abdomen. He hails from Eocene deposits of the Green River Formation of Western Colorado.

The Green River Formation is an Eocene geologic formation that records a 12 million year history of sedimentation in a group of intermountain lakes in three basins along the present-day Green River in Colorado, Wyoming, and Utah. It is one of the most important outcrops we have for insight into life in the Eocene. It gives a window into what our world looked like about 50 million years ago. 

The first documented records of invertebrate fossils from what is now called the Green River Formation are in the journals of early missionaries and explorers such as S. A. Parker, 1840, and J. C. Fremont, 1845. Geologist Dr. John Evans collected the first fossil fish, described as Culpea humilis — later renamed Knightia eocaena — from the Green River beds in 1856.

Edward Drinker Cope collected extensively from the area and produced several publications on the fossil fish from 1870 onwards. Ferdinand Vandeveer Hayden, geologist-in-charge of the United States Geological and Geographical Survey of the Territories, the forerunner of the United States Geological Survey,  first used the name "Green River Shales" for the fossil sites in 1869.

Millions of fish fossils have been collected from the area, commercial collectors operating from legal quarries on state and private land have been responsible for the majority of Green River vertebrate fossils in public and private collections all over the world.

ICE AGE PROBOSCIDEANS: WOOLLY MAMMOTHS

This disarticulated fellow is Mammutus primigenius a Woolly Mammoth from the Pleistocene of Siberia, Russia. 

He's now housed in the Museo Nacional De Ciencias Naturales in Madrid, Spain in a display that shows thoughtful comparisons between the proboscideans. They have a wonderful display of mammoth teeth, the diagnostic flat enamel plates and the equally distinct pointy cusped molars of the mastodons.

He was a true elephant, unlike his less robust cousins, the mastodons. Mammoths were bigger — both in girth and height — weighing in at a max of 13 tonnes. 

They are closely related to Asian elephants and were about the size of the African elephants you see roaming the grasslands of Africa today.

If you stood beside him and reached way up, you might be able to touch his tusks but likely not reach up to his mouth or even his eyes. 

He would have had a shaggy coat of light or dark coloured hair with long outer hair strands covering a dense thick undercoat. His oil glands would have worked overtime to secrete oils, giving him natural — and I'm guessing stinky — waterproofing.

Some of the hair strands we have recovered are more than a meter in length. These behemoth proboscideans boasted long, curved tusks, little ears, short tails and grazed on leaves, shrubs and grasses that would have been work to get at as much of the northern hemisphere was covered in ice and snow during his reign. It is often the teeth of mammoths like those you see in the photo here that we see displayed. 

Their molar teeth were large and have always struck me as looking like ink plates from a printing press. If they are allowed to dry out in collection, they fall apart into discreet plates that can be mistaken for mineralized or calcified rock and not the bits and pieces of mammoth molars that they indeed are. Their large surface area was perfect for grinding down the low nutrient, but for the most part, plentiful grasses that sustained them.

Woolly Mammoth Tusk, Wrangel Island

How did they use their tusks? Likely for displays of strength, protecting their delicate trunks, digging up ground vegetation and in dry riverbeds, digging holes to get at the precious life-giving water. It's a genius design, really. A bit like having a plough on the front of your skull. In the photo here you can see a tusk washed clean in a creek bed on Wrangel Island.

Their size offered protection against other predators once the mammoth was full grown. Sadly for the juveniles, they offered tasty prey to big cats like Homotherium who roamed those ancient grasslands alongside them.

They roamed widely in the Pliocene to Holocene, roaming much of Africa, Europe, Asia and North America. We see them first some 150,000 years ago from remains in Russia then expanding out from Spain to Alaska. They enjoyed a very long lifespan of 60-80 — up to 20 years longer than a mastodon and longer than modern elephants. They enjoyed the prime position as the Apex predator of the megafauna, then declined — partially because of the environment and food resources and partially because of their co-existence with humans. In places where the fossil record shows a preference for hunting smaller prey, humans and megafauna do better together. We see this in places like the Indian Subcontinent where primates and rodents made the menu more often than the large megafauna who roamed there. We also see this in present-day Africa, where the last of the large and lovely megafauna show remarkable resilience in the face of human co-existence.  

The woolly mammoths from the Ukrainian-Russian plains died out 15,000 years ago. This population was followed by woolly mammoths from St. Paul Island in Alaska who died out 5,600 years ago — and quite surprisingly, at least to me, the last mammoth died just 4,000 years ago in the frosty ice on the small island of Wrangel in the Arctic Ocean — their final days spent scratching out a dwindling existence of genetic mutations, howling winds, rain-darkened hills and subsistence on tough grasses grown in thin soil. 

Further reading: Laura Arppe, Juha A. Karhu, Sergey Vartanyan, Dorothée G. Drucker, Heli Etu-Sihvola, Hervé Bocherens. Thriving or surviving? The isotopic record of the Wrangel Island woolly mammoth population. Quaternary Science Reviews, 2019; 222: 105884 DOI: 10.1016/j.quascirev.2019.105884

ALCIDS AUKS: PUFFLING AND DUTIFUL PARENTING

Puffins are any of three small species of alcids or auks in the bird genus Fratercula with a brightly coloured beak during the breeding season.

Their sexy orange beaks shift from a dull grey to bright orange when it is time to attract a mate. While not strictly monogamous, most Puffins choose the same mate year upon year producing adorable chicks or pufflings (awe) from their mating efforts.

Female Puffins produce one single white egg which the parents take turns to incubate over a course of about six weeks. Their dutiful parents share the honour of feeding the wee pufflings five to eight times a day until the chick is ready to fly. Towards the end of July, the fledgeling Puffins begin to venture from the safety of their parents and dry land. Once they take to the seas, mom and dad are released from duty and the newest members of the colony are left to hunt and survive on their own.

These are pelagic seabirds that feed primarily by diving in the water. They breed in large colonies on coastal cliffs or offshore islands, nesting in crevices among rocks or in burrows in the soil. Two species, the tufted puffin and horned puffin are found in the North Pacific Ocean, while the Atlantic puffin is found in the North Atlantic Ocean. This lovely fellow, with his distinctive colouring, is an Atlantic Puffin or "Sea Parrot" from Skomer Island near Pembrokeshire in the southwest of Wales. Wales is bordered by Camarthenshire to the east and Ceredigion to the northeast with the sea bordering everything else. It is a fine place to do some birding if it's seabirds you're after.

These Atlantic Puffins are one of the most famous of all the seabirds and form the largest colony in Southern Britain. They live about 25 years making a living in our cold seas dining on herring, hake and sand eels. Some have been known to live to almost 40 years of age. They are good little swimmers as you might expect, but surprisingly they are great flyers, too! They are hindered by short wings, which makes flight challenging but still possible with effort. Once they get some speed on board, they can fly up to 88 km an hour.

The oldest alcid fossil is Hydrotherikornis from Oregon dating to the Late Eocene while fossils of Aethia and Uria go back to the Late Miocene. Molecular clocks have been used to suggest an origin in the Pacific in the Paleocene. Fossils from North Carolina were originally thought to have been of two Fratercula species but were later reassigned to one Fratercula, the tufted puffin, and a Cerorhinca species. Another extinct species, Dow's puffin, Fratercula dowi,  was found on the Channel Islands of California until the Late Pleistocene or early Holocene.

The Fraterculini are thought to have originated in the Pacific primarily because of their greater diversity in the region. There is only one extant species in the Atlantic, compared to two in the Pacific. The Fraterculini fossil record in the Pacific extends at least as far back as the middle Miocene, with three fossil species of Cerorhinca, and material tentatively referred to that genus, in the middle Miocene to late Pliocene of southern California and northern Mexico.

Although there no records from the Miocene in the Atlantic, a re-examination of the North Carolina material indicated that the diversity of puffins in the early Pliocene was as great in the Atlantic as it is in the Pacific today. This diversity was achieved through influxes of puffins from the Pacific; the later loss of species was due to major oceanographic changes in the late Pliocene due to closure of the Panamanian Seaway and the onset of severe glacial cycles in the North Atlantic.

AMMONITE OF THE RHÔNE

An exquisite specimen of the delicately ridged ammonite, Porpoceras verticosum, from Middle Toarcian outcrops adjacent the Rhône in southeastern France.

Porpoceras (Buchman, 1911) is a genus of ammonite that lived during the early and middle Toarcian stage of the Early Jurassic. We see members of this genus from the uppermost part of Serpentinum Zone to Variabilis Subzone. These beauties are found in Europe, Asia, North America and South America.

Ammonites belonging to this genus have evolute shells, with compressed to depressed whorl section. Flanks were slightly convex and venter has been low. The whorl section is sub-rectangular. 

The rib is pronounced and somewhat fibulate on the inner whorls — just wee nodes here — and tuberculate to spined on the ventrolateral shoulder. It differs from Peronoceras by not having a compressed whorl section and regular nodes or fibulation. Catacoeloceras is also similar, but it has regular ventrolateral tubercules and is missing the classic nodes or fibulation of his cousins.

This specimen hails from southern France near the Rhône, one of the major rivers of Europe. It has twice the average water level of the Loire and is fed by the Rhône Glacier in the Swiss Alps at the far eastern end of the Swiss canton of Valais then passes through Lake Geneva before running through southeastern France. This 10 cm specimen was prepared by the supremely talented José Juárez Ruiz

OH, MEDUSA: JELLYFISH. A HALF BILLION YEARS IN THE MAKING

Mesmerizing, delicate and seemingly impossible — this lovely luminescent denizen of the sea has been living in our oceans for more than half a billion years.

Jellyfish are found all over the world, from surface waters to our deepest seas — and they are old. They are some of the oldest animals in the fossil record.

Jellyfish are not fish at all. These gossamer wonders evolved millions of years before true fish.

Jellyfish and sea jellies are the informal common names given to the medusa-phase or adult phase of certain gelatinous members of the subphylum Medusozoa, a major part of the phylum Cnidaria — more closely related to anemones and corals.

The oldest conulariid scyphozoans appeared between 635 and 577 mya in the Neoproterozoic of the Lantian Formation in China. Others are found in the youngest Ediacaran rocks of the Tamengo Formation of Brazil, c. 505 mya, through to the Triassic. Cubozoans and hydrozoans appeared in the Cambrian of the Marjum Formation in Utah, USA, c. 540 mya.

I have seen all sorts of their brethren growing up on the west coast of Canada in tide pools, washed up on the beach and swam amongst thousands of Moon Jellyfish while scuba diving in the Salish Sea. Their pulsating movements are marvellous.  

In the Kwak̓wala language of the Kwakiutl or Kwakwaka'wakw, speakers of Kwak'wala, of the Pacific Northwest, jellyfish are known as ǥaǥisama.

The dreamy blue and purple ǥaǥisama you see here is but one of a large variety of colours and designs. Jellyfish come in bright yellow, orange, clear with pink spots and are often luminescent.

HAIDA GWAII: HAADALA GWAII-AI

A wreck with tales to tell at Naikoon, Haida Gwaii. The islands have gone by many names. To the people who call the islands home, Haida Gwaii means “island of the people,” it is a shortened version of an earlier name, Haadala Gwaii-ai, or “taken out of concealment.” 

Back at the time of Nangkilslas, it was called Didakwaa Gwaii, or “shoreward country.” 

By any name, the islands are a place of rugged beauty and spirit and enjoy a special place in both the natural and supernatural world. The enormous difference between high and low tide in Haida Gwaii – up to twenty three vertical feet – means that twice a day, vast swathes of shellfish are unveiled, free for the taking. 

An ancient Haida saying is still often heard today, “When the tide is out, the table is set.” Archaeological evidence shows that by about five thousand years ago, gathering shellfish replaced hunting and fishing as their primary food source. The shellfish meat was skewered on sticks, smoked and stored for use in winter or for travel.

Steeped in mist and mythology, the islands of Haida Gwaii abound in local lore that surrounds their beginnings. Today, the Hecate Strait is a tempestuous 40-mile wide channel that separates the mist-shrouded archipelago of Haida Gwaii from the BC mainland. Haida oral tradition tells of a time when the strait was mostly dry, dotted here and there with lakes. During the last ice age, glaciers locked up so much water that the sea level was hundreds of feet lower than it is today. Soil samples from the sea floor contain wood, pollen, and other terrestrial plant materials that tell of a tundra-like environment.

The islands of Haida Gwaii are at the western edge of the continental shelf and form part of Wrangellia, an exotic terrane of former island arcs, which also includes Vancouver Island, parts of western mainland British Columbia and southern Alaska. 

Brewericeras hulenense (Anderson, 1938)

While we’ll see that there are two competing schools of thought on Wrangellia’s more recent history, both sides agree that many of the rocks, and the fossils they contain, were laid down somewhere near the equator. 

They had a long, arduous journey, first being pushed by advancing plates, then being uplifted, intruded, folded, and finally thrust up again. It’s reminiscent of how pastry is balled up, kneaded over and over, finally rolled out, then the process is repeated again.

This violent history applies to most of the rock that makes up the Insular Belt, the outermost edge of the Cordillera. Once in their present location, the rocks that make up the mountains and valleys of this island group were glaciated and eroded to their present form. Despite this tumultuous past, the islands have arguably the best-preserved and most fossil-rich rocks in the Canadian Cordillera, dating from very recent to more than 200 million years old. 

The fossils found in the Triassic rock of Wrangellia are equatorial or low latitude life forms quite different from those found today on the Continent at the latitude of Haida Gwaii. This suggests those rocks were in the equatorial region during the Late Triassic, just over 200 million years ago. 

The Lower Jurassic ammonite faunas found at Haida Gwaii are very similar to those found in the Eastern Pacific around South America and in the Mediterranean. The strata exposed at Maple Island, Haida Gwaii are stratigraphically higher than the majority of Albian localities in Skidegate Inlet. The macrofossil fauna belonged to the Upper part of the Sandstone Member of the Haida formation.

The western end of the island contains numerous well-preserved inoceramids such as Birostrina concentrica and a few rare ammonites of Desmoceras bearskinese. The eastern shores are home to unusual ammonite fauna in the finer grained sandstones. Here we find the fossils as extremely hard concretions while others were loose in the shale. Species include Anagaudryceras sacya and Tetragonites subtimotheanus. A large whorl section of the rare Ammonoceratites crenucostatus has also been found here. The ammonites, Desmoceras; Brewericeras hulenense; Cleoniceras perezianum, Douvelliceras spiniferum are all found in Lower Cretaceous, Middle Albian, Haida Formation deposits.

CTENOPHORES: COMB JELLIES

Cannibalistic Comb Jellies

This festive lantern looking lovely belongs to a group of invertebrates known as comb jellies.

Comb jellies are named for their unique plates of giant fused cilia, or combs, which run in eight rows up and down the length of their bodies. They are armed with sticky cells or colloblasts, that do not sting but display wonderful bioluminescent colouring as they move through the sea.

Ctenophores or comb jellies are one of the phylogenetically most important and controversial metazoan groups. They are not jellyfish and are not closely related, though they do share some characteristics with the gelatinous members of the subphylum Medusozoa. 

Comb jellies are not picky eaters. Their tastes range to what is at hand, including cannibalizing other comb jellies. They will feast on their kin along with tasty plankton, zooplankton, crustaceans and wee fish.

Interest in their fossil record has been catalysed by spectacularly preserved soft-bodied specimens from Cambrian Lagerstätten of the 518-million-years-old Chengjiang Biota, the 505-million-years-old Burgess Shale and other Burgess Shale-like deposits. 

We find them in the Late Devonian Escuminac Formation at Miguasha National Park, Quebec, Canada — a UNESCO world heritage site famous for its abundance of well-preserved vertebrate fossils including most major evolutionary groups of Devonian lower vertebrates from jawless fish to stem-tetrapods.

Based on morphological similarities of this Canadian fossil with stem-ctenophore fossils from the Cambrian Lagerstätte of the Chinese locality Chengjiang, they have been assessed for their affinity to stem-group ctenophores (dinomischids, Siphusauctum, scleroctenophorans) and early crown-group ctenophores. Modern ctenophores and many fossil forms lack mineralized hard parts, which renders the rare fossils that have been extracted from several Lagerstätten quite remarkable. 

Like the soft bodies of jellyfish and the polyps of hydrozoans and anthozoans, the probability for such soft bodies (or body regions) to become fossilized is extremely low. In spite of this low preservation potential, remains of stem-ctenophores have become known from several Cambrian and younger conservation deposits, and with even older candidate ctenophores in the Ediacaran. 

While Cambrian Lagerstätten have yielded several genera, ctenophore remains are much rarer in the Devonian; in particular, two studies, describing material from the German Hunsrück Slate. 

Bioluminescent Comb Jellies

This Early Devonian material, however, appears to belong to crown ctenophores morphologically similar to living forms such as Pleurobrachia, unlike the stem Cambrian taxa and the new Devonian stem taxon described here.

The most basal stem ctenophores are the dinomischids: sessile benthic petaloid invertebrates, many of which are equipped with a stalk. This group first was described from the Middle Cambrian Burgess Shale. Based on the genus Dinomischus, these early stalked forms were commonly called ‘dinomischids’. 

Zhao et al. shared that dinomischids "form a grade in the lower part of the ctenophore stem group” and include taxa such as Xianguangia, Daihua, and Dinomischus that have hexaradiate-based symmetry (e.g., sixfold, 18-fold). 

Some later, skeletonised stem-ctenophores were termed ‘Scleroctenophora’; ‘scleroctenophorans’ have a shorter stalk, lack the ‘petals’ and have no bracts and might be monophyletic. 

To date, all known dinomischids and scleroctenophorans are Cambrian. Remarkably, analysis of the material described here suggests it is a very late-surviving member of this part of the ctenophore tree, occurring in strata over a hundred million years younger with no intervening known record, thus making it a Lazarus taxon with an extensive ghost lineage. 

Palaeozoic sediments yield a growing number of fossil invertebrates with radial symmetries, some being quite enigmatic with body plans differing radically from those of extant organisms.

The morphological similarities to Cambrian forms and the mix of characters regarding overall shape and symmetries render this discovery important. The aims of this study are to describe the only known specimen of this Devonian ctenophore, discuss its phylogenetic and systematic position, and the impact of fossil data for ctenophore affinities, and assess its palaeoecological role.