BLUE LIAS ICHTHYOSAUR

This well-preserved partial ichthyosaur was found in the Blue Lias shales by Lewis Winchester-Ellis in 2018. The vertebrae you see here are from the tail section of this marine reptile.

The find includes stomach contents which tell us a little about how this particular fellow liked to dine.

As with most of his brethren, he enjoyed fish and cephalopods. Lewis found fishbone and squid tentacle hooklets in his belly. Oh yes, these ancient cephies had grasping hooklets on their tentacles. I'm picturing them wiggling all ominously. The hooklets were the only hard parts of the animal preserved in this case as the softer parts of this ancient calamari were fully or partially digested before this ichthyosaur met his end.

Ichthyosaurus was an extinct marine reptile first described from fossil fragments found in 1699 in Wales. Shortly thereafter, fossil vertebrae were published in 1708 from the Lower Jurassic and the first member of the order Ichthyosauria to be discovered.

To give that a bit of historical significance, this was the age of James Stuart, Jacobite hopeful to the British throne. While scientific journals of the day were publishing the first vertebrae ichthyosaur finds, he was avoiding the French fleet in the Firth of Forth off Scotland. This wasn’t Bonnie Prince Charlie, this was his Dad. Yes, that far back.

The first complete skeleton was discovered in the early 19th century by Mary Anning and her brother Joseph along the Dorset Jurassic Coast. Joseph had mistakenly, but quite reasonably, taken the find for an ancient crocodile. Mary excavated the specimen a year later and it was this and others that she found that would supply the research base others would soon publish on.

Mary's find was described by a British surgeon, Sir Everard Home, an elected Fellow of the Royal Society, in 1814. The specimen is now on display at the Natural History Museum in London bearing the name Temnodontosaurus platyodon, or “cutting-tooth lizard.”

Ichthyosaurus communis

In 1821, William Conybeare and Henry De La Beche, a friend of Mary's, published a paper describing three new species of unknown marine reptiles based on the Anning's finds.

Rev. William Buckland would go on to describe two small ichthyosaurs from the Lias of Lyme Regis, Ichthyosaurus communis and Ichthyosaurus intermedius, in 1837.

Remarkable, you'll recall that he was a theologian, geologist, palaeontologist AND Dean of Westminster. It was Buckland who published the first full account of a dinosaur in 1824, coining the name, "Megalosaurus."

The Age of Dinosaurs and Era of the Mighty Marine Reptile had begun.

Ichthyosaurs have been collected in the Blue Lias near Lyme Regis and the Black Ven Marls. More recently, specimens have been collected from the higher succession near Seatown. Paddy Howe, Lyme Regis Museum geologist, found a rather nice Ichthyosaurus breviceps skull a few years back. A landslip in 2008 unveiled some ribs poking out of the Church cliffs and a bit of digging revealed the ninth fossil skull ever found of a breviceps, with teeth and paddles to boot.

Specimens have since been found in Europe in Belgium, England, Germany, Switzerland and in Indonesia. Many tremendously well-preserved specimens come from the limestone quarries in Holzmaden, southern Germany.

Ichthyosaurs ranged from quite small, just a foot or two, to well over twenty-six metres in length and resembled both modern fish and dolphins.

Dean Lomax and Sven Sachs, both active (and delightful) vertebrate paleontologists, have described a colossal beast, Shonisaurus sikanniensis from the Upper Triassic (Norian) Pardonet Formation of northeastern British Columbia, Canada, measuring 3-3.5 meters in length. The specimen is now on display in the Royal Tyrrell Museum of Palaeontology in Alberta, Canada. It was this discovery that tipped the balance in the vote, making it British Columbia's Official Fossil. Ichthyosaurs have been found at other sites in British Columbia, on Vancouver Island and the Queen Charlotte Islands (Haida Gwaii) but Shoni tipped the ballot.

The first specimens of Shonisaurus were found in the 1990s by Peter Langham at Doniford Bay on the Somerset coast of England.

Dr. Betsy Nicholls, Rolex Laureate Vertebrate Palaeontologist from the Royal Tyrrell Museum, excavated the type specimen of Shonisaurus sikanniensis over three field sessions in one of the most ambitious fossil excavations ever ventured. Her efforts from 1999 through 2001, both in the field and lobbying back at home, paid off. Betsy published on this new species in 2004, the culmination of her life’s work and her last paper as we lost her to cancer in autumn of that year.

Roy Chapman Andrews, AMNH 1928 Expedition to the Gobi Desert

Charmingly, Betsy had a mail correspondence with Roy Chapman Andrews, former director of the American Museum of Natural History, going back to the late 1950s as she explored her potential career in palaeontology. Do you recall the AMNH’s sexy paleo photos of expeditions to the Gobi Desert in southern Mongolia in China in the early 20th century? I've posted a picture here to jog your memory. Roy Chapman Andrews was the lead on that trip. The man was dead sexy. His photos are what fueled the flames of my own interest in paleo.

We've found at least 37 specimens of Shonisaurus in Triassic outcrops of the Luning Formation in the Shoshone Mountains of Nevada, USA. The finds go back to the 1920s. The specimens that may it to publication were collected by M. Wheat and C. L. Camp in the 1950s.  The aptly named Shonisaurus popularis became the Nevada State Fossil in 1984. Our Shoni got around. Isolated remains have been found in a section of sandstone in Belluno, in the Eastern Dolomites, Veneto region of northeastern Italy. The specimens were published by Vecchia et al. in 2002.

For a time, Shonisaurus was the largest ichthyosaurus known.

Move over, Shoni, as a new marine reptile find competes with the Green Anaconda (Eunectes murinus) and the Blue Whale (Balaenoptera musculus) for size at a whopping twenty-six (26) metres.

The find is the prize of fossil collector turned co-author, Paul de la Salle, who (you guessed it) found it in the lower part of the intertidal area that outcrops strata from the latest Triassic Westbury Mudstone Formation of Lilstock on the Somerset coast. He contacted Dean Lomax and Judy Massare who became co-authors on the paper.

The find and conclusions from their paper put "dinosaur" bones from the historic Westbury Mudstone Formation of Aust Cliff, Gloucestershire, UK site into full reinterpretation.

And remember that ichthyosaur the good Reverend Buckland described back in 1837, the Ichthyosaurus communis? Dean Lomax was the first to describe a wee baby. A wee baby ichthyosaur! Awe. I know, right? He and paleontologist Nigel Larkin published this adorable first in the journal of Historical Biology in 2017.

They had teamed up previously on another first back in 2014 when they completed the reconstruction of an entire large marine reptile skull and mandible in 3-D, then graciously making it available to fellow researchers and the public. The skull and braincase in question were from an Early Jurassic, and relatively rare, Protoichthyosaurus prostaxalis. The specimen had been unearthed in Warwickshire back in the 1950s. Unlike most ichthyosaur finds of this age, it was not compressed and allowed the team to look at a 3-D specimen through the lens of computerized tomography (CT) scanning.

Another superb 3-D ichthyosaur skull was found near Lyme Regis by fossil hunter-turned-entrepreneur-local David Sole and prepped by the late David Costain. I'm rather hoping it went into a museum collection as it would be wonderful to see the specimen studied, imaged, scanned and 3-D printed for all to share. Here's hoping.

Ichthyosaurus somersetensis Credit: Dean R Lomax

Lomax and Sven Sachs also published on an embryo from one of the largest ichthyosaurs known, a new species named Ichthyosaurus somersetensis.

Their paper in the ACTA Palaeontologica Polonica from 2017, describes the third embryo known for Ichthyosaurus and the first to be positively identified to species level. The specimen was collected from Lower Jurassic strata (lower Hettangian, Blue Lias Formation) of Doniford Bay, Somerset, UK and is housed in the collection of the Niedersächsisches Landesmuseum (Lower Saxony State Museum) in Hannover, Germany.

We've learned a lot about them in the time we've been studying them. We now have thousands of specimens, some whole, some as bits and pieces. Many specimens that have been collected are only just now being studied and the tools we are using to study them are getting better and better.

Link to Lomax Paper: https://journals.plos.org/plosone/article…

Link to Nathan's Paper: https://www.tandfonline.com/…/10.1080/03115518.2018.1523462…

Nicholls Paper: E. L. Nicholls and M. Manabe. 2004. Giant ichthyosaurs of the Triassic - a new species of Shonisaurus from the Pardonet Formation (Norian: Late Triassic) of British Columbia. Journal of Vertebrate Paleontology 24(4):838-849 [M. Carrano/H. Street]

PLESIOSAURS OF THE YORKSHIRE COAST

These two lovely Plesiosaur vertebrae were found by Liam Langley on fossil field trips to the Yorkshire Coast on the east coast of England.

Plesiosaurus was a large, carnivorous air-breathing marine reptile with strong jaws and sharp teeth that moved through the water with four flippers. We'd originally thought that this might not be the most aerodynamic design but it was clearly effective as they used the extra set to create a wee vortex that aided in their propulsion. In terms of mechanical design, they have a little something in common with dragonflies.

We've recreated plesiosaur movements and discovered that they were able to optimize propulsion to make use of their own wake. As their front flippers paddled in big circular movements, the propelled water created little whirlpools under their bellies.

The back flippers would then paddle between these whirlpools pushing the plesiosaur forward to maximal effect. They were very successful hunters, outcompeting ichthyosaurs who thrived in the Triassic but were replaced in the Jurassic and Cretaceous by these new aquatic beasties. Our ancient seas teemed with these predatory marine reptiles with their long necks and barrel-shaped bodies. Plesiosaurs were smaller than their pliosaur cousins, weighing in at about 450 kg or 1,000 lbs and reaching about 4.5 metres or 15 feet in length. For a modern comparison, they were roughly twice as long as a standard horse or about as long as a good size hippo.

Plesiosaurs first appeared in the latest Triassic, during the Rhaetian. They thrived in the Jurassic and vanished at the end of the Cretaceous in time with the K-Pg extinction event along with a host of other species. They are one of the marine reptiles that we associate with the infamous Mary Anning, a paleo darling of the early 19th century who found her first fossil specimen in the winter of 1823. These two vertebrae grace the home of the talented Mr. Langley. Anning's plesiosaur can be viewed in London's Natural History Museum.

SHONISAURUS OF NEVADA

The beauties you see here are ichthyosaurs. The largest of their lineage is the genus Shonisaurus who ruled our ancient seas 217 million years ago.

At least 37 incomplete fossil specimens of the marine reptile have been found in hard limestone deposits of the Luning Formation, in far northwestern Nye County of Nevada. This formation dates to the late Carnian age of the late Triassic period when present-day Nevada and parts of the western United States were covered by an ancient ocean.

The first researcher to recognize the Nevada fossil specimens as ichthyosaurs was Siemon W. Muller of Stanford University. He had the work of Sir Richard Owen and others to build on. That being said, there are very few contenders for a species that boasts vertebrae over a foot wide and weighing in at almost 10 kg or 21 lbs. Muller contacted the University of California Museum of Paleontology at Berkeley. Surface collecting by locals continued at the site but no major excavation was planned.

Sir Richard Owen, the British biologist, comparative anatomist and paleontologist, coined the name ichthyopterygia, or "fish flippers," one hundred and fourteen years earlier, but that wee bit of scientific knowledge hadn't made its way west to the general population. The finds at Luning were still, "marine monsters."

Owen, too, was building on research going back to 1699, the very first recorded fossil fragments found of these beasties in Wales. Shortly thereafter, fossil vertebrae were published in 1708 from the Lower Jurassic.

The first complete skeleton was discovered in the early 19th century by Mary Anning and her brother Joseph along the Dorset Jurassic Coast. Mary's find was described by a British surgeon, Sir Everard Home, an elected Fellow of the Royal Society, in 1814. The specimen is now on display at the Natural History Museum in London bearing the name Temnodontosaurus platyodon, or “cutting-tooth lizard.”

In 1821, William Conybeare and Henry De La Beche, a friend of Mary's, published a paper describing three new species of unknown marine reptiles based on the Anning's finds. The Rev. William Buckland would go on to describe two small ichthyosaurs from the Lias of Lyme Regis, Ichthyosaurus communis and Ichthyosaurus intermedius. All of this early work was instrumental in aiding the researchers who would join the project at Luning.

Owen is considered to have been an outstanding naturalist with a remarkable gift for interpreting fossils. Contrary to common belief, advanced study does help with identifying fossils, but what is truly needed is a keen eye. The finds at Luning were blessed to be seen by an enthusiastic local with just that right kind of keen eye.

Almost a quarter of a century after Muller's initial reports, Dr. Charles L. Camp from UCMP received correspondence further detailing the finds from a lovely Mrs. Margaret Wheat of Fallon. She wrote to Camp in September of 1928 to say that she'd been giving the quarry section a bit of a sweep, as you do, and had uncovered a nice aligned section of vertebrae with her broom. The following year, Dr. Charles L. Camp went out to survey the finds and began working on the specimens, his first field season of many, in 1954.

Back in the 1950s, these large marine reptiles were rumoured to be "marine monsters," as the concept of an ichthyosaur was not well understood by the local townsfolk. Excitement soon hit West Union Canyon as the quarry began to reveal the sheer size of these mighty beasts. Four of the specimens were fully excavated. Most of the ichthyosaur bones were left in situ, partially because the work was tremendously difficult, and partially to allow others to see how the specimens were laid down over 200 million years ago.

Camp continued to work with Wheat at the site and brought on Sam Welles and a host of students to help with excavations. The team understood the need for protection at the site. They canvassed the Nevada Legislature to establish the Ichthyosaur Paleontological State Monument. You can see one of the Park Rangers above giving a tour within the lovely Fossil Hut building they built on the site to protect the fossils.

In 1957, the site was incorporated into the State Park System and Berlin-Ichthyosaur State Park was born. The park Twenty years later, in 1977, the population of Nevada weighed in and the Legislature designated Shonisaurus popularis as the State Fossil of Nevada. Visitors are welcome to collect fossils from the exposures of the Upper Triassic (Early Norian, Kerri Zone) of the Luning Formation, West Union Canyon, just outside Berlin-Ichthyosaur State Park.

Address: State route 844, Austin, NV 89310, United States. Area: 4.58 km². Open 24 hours;
Elevation: 6,975 ft (2,126 m); Tel: +1 775-964-2440; http://parks.nv.gov/parks/berlin-ichthyosaur

CENOMANIAN-TURONIAN IMPACT

Ichthyosaur and Plesiosaur by Edouard Riou, 1863

During the early Triassic period, ichthyosaurs evolved from a group of unidentified land reptiles that returned to the sea. They were particularly abundant in the later Triassic and early Jurassic periods before being replaced as a premier aquatic predator by another marine reptilian group, the Plesiosauria, in the later Jurassic and Cretaceous periods.

In the Late Cretaceous, ichthyosaurs were hard hit by the Cenomanian-Turonian anoxic event. As the deepest benthos layers of the seas became anoxic, poisoned by hydrogen sulphide, deep water marine life died off. This caused a cascade that wreaked havoc all the way up the food chain. At the end of that chain were our mighty predaceous marine reptiles.

Bounty turned to scarcity and a race for survival began. The ichthyosaurs lost that race as the last of their lineage became extinct. It may have been their conservative evolution as a genus when faced with a need for adaptation to the world in which they found themselves and/or being outcompeted by early mosasaurs.