Fast-forward 240 million years, and the evolutionary landscape had transformed dramatically.
Vertebrates had conquered the land, ecosystems had diversified, and Earth’s surface teemed with reptilian innovators, amphibians the size of crocodiles, and the early ancestors of mammals. Among these emerging terrestrial titans strode the Gorgonopsians, or “Gorgons”—ferocious sabre-toothed therapsids that dominated the Middle to Late Permian, from about 265 to 252 million years ago.
These were no sluggish proto-reptiles. Gorgons were highly specialized predators, boasting elongated canine teeth worthy of any future saber-toothed cat, powerful jaws, and sleek, muscular bodies built for pursuit. Their anatomy blended the primitive and the prophetic: reptile-like postures paired with early mammalian traits such as differentiated teeth and strong jaw musculature.
Their clawed limbs, keen forward-facing eyes, and cutting-edge predatory adaptations placed them firmly at the top of the Permian food chain. In a world long before dinosaurs, they were the undisputed apex hunters.
My own fascination with these remarkable creatures was ignited by Gorgons, Peter Ward’s wonderfully wry and insightful dive into the ancient landscapes of South Africa. Ward’s vivid tales of fieldwork in the blistering, bone-dry vastness of the Karoo Basin—ancestral home of the Gorgons—captured both the hardships and the sheer exhilaration of unearthing deep time.
His descriptions of sunburn and scientific revelations in that arid world made me laugh more than once. It is a highly enjoyable read.
The Great Karoo itself is a geological and paleontological marvel. This enormous, semi-arid expanse formed within a vast inland basin roughly 320 million years ago, at a time when the part of Gondwana destined to become Africa lay draped across the South Pole.
Layer upon layer of sedimentary rock accumulated as glaciers advanced and retreated, rivers meandered, lakes dried, and ecosystems rose and fell. Today, those layers read like a grand evolutionary chronicle, preserving a world populated by beaked herbivores, hulking amphibians, and the charismatic, toothy Gorgonopsians.
This was a pivotal chapter in Earth’s history—just before the catastrophic Permian-Triassic extinction swept away nearly 90% of life. Yet in the twilight of the Permian, before that great dying, the Karoo thrived with innovation and ecological complexity. It was a world where the early steps toward warm-bloodedness were being taken, where synapsids (our own deep ancestors) were experimenting with new forms, and where the Gorgons reigned supreme.
